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<document id="D3205AC4893B2C7B2F5AF1F60BBC1417" ID-CLB-Dataset="32777" ID-DOI="10.11646/zootaxa.4282.3.2" ID-GBIF-Dataset="13966ea2-f556-4d72-917e-a1b7dd85d0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="827888" ID-ZooBank="797FDD34-9129-4EC4-8A8F-1B7DEBA77E06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500107109175" checkinUser="plazi" docAuthor="Kluge, Nikita J. &amp; Novikova, Eugenia A." docDate="2017" docId="8E2587C2FFAD682C9ED4FBEF4676FDC7" docLanguage="en" docName="zootaxa.4282.3.2.pdf" docOrigin="Zootaxa 4282 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anafroptilum orthostylus Kluge &amp; Novikova, 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="463" masterDocId="721CFFBAFFA568269E43FFC84520FF98" masterDocTitle="Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region" masterLastPageNumber="472" masterPageNumber="453" pageNumber="461" updateTime="1698464953366" updateUser="ExternalLinkService">
<mods:mods id="1FC074A92503D91AA4701376D07B305A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="31383DF1DBA41D34456819AC59358CBC">Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region</mods:title>
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<mods:name id="21297B0098C2754E2050FEBF7E66AEF8" type="personal">
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<mods:namePart id="201C9F1FE26F8188A73BA1BF4DE606F5">Kluge, Nikita J.</mods:namePart>
</mods:name>
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<mods:role id="AFE3C3550644407E5079F8D9B8263B31">
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<mods:namePart id="BF4A80DE1A9BFF0EB7B51D62310CDFE3">Novikova, Eugenia A.</mods:namePart>
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<mods:typeOfResource id="EC50DCDD4A31F5431FF8B7AFA3B3B076">text</mods:typeOfResource>
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<mods:title id="475795B99D8BCAF3C45E2DE1EF39A4FC">Zootaxa</mods:title>
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<mods:part id="6570BE2DCBCAD0A1306688A2383B46CC">
<mods:date id="8EED53093BEB558BF64C4A7876FA0610">2017</mods:date>
<mods:detail id="E69F2D841E28EBF46C24BCDF06596A0B" type="volume">
<mods:number id="D5CDFE308C4930906254DA3532E4E636">4282</mods:number>
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<mods:number id="0658C01479453AA470AB52C67BA56DE1">3</mods:number>
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<mods:classification id="8688A95EF29CC8E2DB977DA1C3503543">journal article</mods:classification>
<mods:identifier id="167EB655AFA74D90A170A94E03329932" type="CLB-Dataset">32777</mods:identifier>
<mods:identifier id="209791EAF940F5E0C70486AA584AA79E" type="DOI">10.11646/zootaxa.4282.3.2</mods:identifier>
<mods:identifier id="099AA7F974EA1A4D7F9B49FFC7DB9DEB" type="GBIF-Dataset">13966ea2-f556-4d72-917e-a1b7dd85d0ec</mods:identifier>
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<treatment id="8E2587C2FFAD682C9ED4FBEF4676FDC7" ID-DOI="http://doi.org/10.5281/zenodo.6028651" ID-GBIF-Taxon="132548382" ID-Zenodo-Dep="6028651" LSID="urn:lsid:plazi:treatment:8E2587C2FFAD682C9ED4FBEF4676FDC7" httpUri="http://treatment.plazi.org/id/8E2587C2FFAD682C9ED4FBEF4676FDC7" lastPageId="10" lastPageNumber="463" pageId="8" pageNumber="461">
<subSubSection id="4E96655FFFAD682E9ED4FBEF4439FBFB" pageId="8" pageNumber="461" type="nomenclature">
<paragraph id="063336D4FFAD682E9ED4FBEF4731FBD9" blockId="8.[151,529,1063,1123]" box="[151,529,1063,1089]" pageId="8" pageNumber="461">
<heading id="5D7B81B8FFAD682E9ED4FBEF4731FBD9" bold="true" box="[151,529,1063,1089]" fontSize="11" level="1" pageId="8" pageNumber="461" reason="1">
<emphasis id="34F8EAC6FFAD682E9ED4FBEF4731FBD9" bold="true" box="[151,529,1063,1089]" pageId="8" pageNumber="461">
<taxonomicName id="C18C4D57FFAD682E9ED4FBEF44E9FBD9" box="[151,457,1063,1089]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFAD682E9ED4FBEF44E9FBD9" bold="true" box="[151,457,1063,1089]" italics="true" pageId="8" pageNumber="461">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFAD682E9F8CFBEF4731FBD9" box="[463,529,1063,1089]" pageId="8" pageNumber="461" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="063336D4FFAD682E9ED4FB824439FBFB" blockId="8.[151,529,1063,1123]" box="[151,281,1098,1123]" pageId="8" pageNumber="461">
(
<figureCitation id="9EB72A51FFAD682E9EDCFB824430FBFB" box="[159,272,1098,1123]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 18" captionStart-2="FIGURES 19 – 27" captionStart-3="FIGURES 28 – 36" captionStart-4="FIGURES 37 – 43" captionStart-5="FIGURES 44 – 51" captionStart-6="FIGURES 52 – 53" captionStartId-0="2.[151,264,1929,1952]" captionStartId-1="3.[151,264,1927,1950]" captionStartId-2="4.[151,264,1722,1745]" captionStartId-3="5.[151,264,1949,1972]" captionStartId-4="6.[151,264,1794,1817]" captionStartId-5="7.[151,264,1927,1950]" captionStartId-6="8.[151,264,960,983]" captionTargetBox-0="[166,1430,195,1902]" captionTargetBox-1="[151,1436,193,1906]" captionTargetBox-2="[151,1436,193,1696]" captionTargetBox-3="[220,1301,599,1910]" captionTargetBox-4="[201,1383,200,1772]" captionTargetBox-5="[151,1436,193,1906]" captionTargetBox-6="[151,1436,425,911]" captionTargetId-0="figure@2.[166,1430,195,1903]" captionTargetId-1="figure@3.[151,1436,193,1906]" captionTargetId-2="figure@4.[151,1436,193,1701]" captionTargetId-3="figure@5.[216,1351,589,1927]" captionTargetId-4="figure@6.[201,1395,193,1781]" captionTargetId-5="figure@7.[151,1436,193,1906]" captionTargetId-6="figure@8.[151,1436,425,911]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." captionText-1="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." captionText-2="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." captionText-3="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." captionText-4="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." captionText-5="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." captionText-6="FIGURES 52 – 53. Anafroptilum orthostylus sp. n., eggs." httpUri-0="https://zenodo.org/record/827890/files/figure.png" httpUri-1="https://zenodo.org/record/827892/files/figure.png" httpUri-2="https://zenodo.org/record/827894/files/figure.png" httpUri-3="https://zenodo.org/record/827896/files/figure.png" httpUri-4="https://zenodo.org/record/827898/files/figure.png" httpUri-5="https://zenodo.org/record/827900/files/figure.png" httpUri-6="https://zenodo.org/record/827902/files/figure.png" pageId="8" pageNumber="461">Figs 1–53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAD682E9ED4FB5A4100FB33" box="[151,1056,1170,1195]" pageId="8" pageNumber="461" type="etymology">
<paragraph id="063336D4FFAD682E9ED4FB5A4100FB33" blockId="8.[151,1437,1170,2023]" box="[151,1056,1170,1195]" pageId="8" pageNumber="461">
<emphasis id="34F8EAC6FFAD682E9ED4FB5A443EFB33" bold="true" box="[151,286,1170,1195]" pageId="8" pageNumber="461">Etymology.</emphasis>
Referring to straight shape of male imaginal gonostyli (
<figureCitation id="9EB72A51FFAD682E9DD2FB5A46C8FB33" box="[913,1000,1170,1195]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="8" pageNumber="461">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFAD682E9DB7FB5A4134FB33" box="[1012,1044,1170,1195]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="8" pageNumber="461">50</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAD682E9E84FB7E464AF897" pageId="8" pageNumber="461" type="materials_examined">
<paragraph id="063336D4FFAD682E9E84FB7E4662FA8F" blockId="8.[151,1437,1170,2023]" pageId="8" pageNumber="461">
<materialsCitation id="B6E43C89FFAD682E9E84FB7E47C4FB6B" ID-GBIF-Occurrence="1572324439" collectingDate="2015-02-06" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" location="Erawan Falls" pageId="8" pageNumber="461" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi Province" typeStatus="holotype">
<emphasis id="34F8EAC6FFAD682E9E84FB7E4495FB57" bold="true" box="[199,437,1206,1231]" pageId="8" pageNumber="461">Material examined.</emphasis>
<typeStatus id="D9378876FFAD682E9F82FB7E470EFB57" box="[449,558,1206,1231]" pageId="8" pageNumber="461" type="holotype">Holotype</typeStatus>
: L-S-I
<specimenCount id="108AFD5DFFAD682E9C3FFB7F47AFFB57" box="[636,655,1207,1231]" pageId="8" pageNumber="461" type="male">♂</specimenCount>
{specimen [X] (13) 2015},
<collectingCountry id="7E9B7644FFAD682E9DA7FB7F4158FB57" box="[996,1144,1207,1231]" name="Thailand" pageId="8" pageNumber="461">THAILAND</collectingCountry>
,
<collectingRegion id="C448F836FFAD682E9ACBFB7E40B8FB57" box="[1160,1432,1206,1231]" country="Thailand" name="Kanchanaburi" pageId="8" pageNumber="461">Kanchanaburi Province</collectingRegion>
,
<location id="0353600FFFAD682E9ED4FB13440CFB6B" LSID="urn:lsid:plazi:treatment:8E2587C2FFAD682C9ED4FBEF4676FDC7:0353600FFFAD682E9ED4FB13440CFB6B" box="[151,300,1242,1267]" country="Thailand" name="Erawan Falls" pageId="8" pageNumber="461" stateProvince="Kanchanaburi Province">Erawan Falls</location>
,
<collectingDate id="6276E9FCFFAD682E9F74FB124481FB6B" box="[311,417,1242,1267]" pageId="8" pageNumber="461" value="2015-02-06">
<date id="72321014FFAD682E9F74FB124481FB6B" box="[311,417,1242,1267]" pageId="8" pageNumber="461" value="2015-02-06">6.II.2015</date>
</collectingDate>
, coll.
<collectorName id="AB795302FFAD682E9FA0FB134769FB6B" box="[483,585,1242,1267]" pageId="8" pageNumber="461">N. Kluge</collectorName>
&amp;
<collectorName id="AB795302FFAD682E9C2EFB1347C4FB6B" box="[621,740,1242,1267]" pageId="8" pageNumber="461">L. Sheyko</collectorName>
</materialsCitation>
.
<materialsCitation id="B6E43C89FFAD682E9CADFB134468FA8F" ID-GBIF-Occurrence="1572324441" collectingDate="2015-02-02" collectingDateMax="2015-02-04" collectingDateMin="2015-02-02" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" location="Erawan Falls" pageId="8" pageNumber="461" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi Province" typeStatus="paratype">
<typeStatus id="D9378876FFAD682E9CADFB13467FFB6B" box="[750,863,1243,1267]" pageId="8" pageNumber="461" type="paratype">Paratypes</typeStatus>
: the same locality,
<collectingDate id="6276E9FCFFAD682E9A71FB124197FB6B" box="[1074,1207,1242,1267]" pageId="8" pageNumber="461" value="2015-02-02" valueMax="2015-02-04" valueMin="2015-02-02">
<date id="72321014FFAD682E9A71FB124197FB6B" box="[1074,1207,1242,1267]" pageId="8" pageNumber="461" value="2015-02-02" valueMax="2015-02-04" valueMin="2015-02-02">2–4.II.2015</date>
</collectingDate>
, coll.
<collectorName id="AB795302FFAD682E9AB9FB134040FB6B" box="[1274,1376,1242,1267]" pageId="8" pageNumber="461">N. Kluge</collectorName>
&amp;
<collectorName id="AB795302FFAD682E9BC7FB1345CFFA8F" pageId="8" pageNumber="461">L. Sheyko</collectorName>
: 2 S-I
<specimenCount id="108AFD5DFFAD682E9F77FB374468FA8F" box="[308,328,1279,1303]" pageId="8" pageNumber="461" type="male">♂</specimenCount>
</materialsCitation>
,
<date id="72321014FFAD682E9F17FB3744A1FA8E" box="[340,385,1279,1303]" pageId="8" pageNumber="461" value="1924-01">24 I</date>
♂, 1 L/S♂, 1 L♂, 5 S-I♀,
<date id="72321014FFAD682E9CE4FB3747E9FA8E" box="[679,713,1279,1303]" pageId="8" pageNumber="461">1 I</date>
♀, 2 L/S♀.
</paragraph>
<paragraph id="063336D4FFAD682E9E84FAEA464AF897" blockId="8.[151,1437,1170,2023]" pageId="8" pageNumber="461">
<emphasis id="34F8EAC6FFAD682E9E84FAEA4798FAA3" bold="true" box="[199,696,1314,1339]" pageId="8" pageNumber="461">
Descriptions.
<emphasis id="34F8EAC6FFAD682E9F29FAEB4495FAA2" bold="true" box="[362,437,1315,1338]" italics="true" pageId="8" pageNumber="461">Larva.</emphasis>
Cuticular coloration.
</emphasis>
Head with colorless and very light brownish areas. Pronotum and mesonotum light brownish with darker brown markings; protoptera light with darker stripes corresponding to longitudinal veins (
<figureCitation id="9EB72A51FFAD682E9F33FAA24495FA1B" box="[368,437,1386,1411]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Fig. 4</figureCitation>
). Metanotum, thoracic pleura and sterna brown, membranes colorless (
<figureCitation id="9EB72A51FFAD682E9A93FAA24016FA1B" box="[1232,1334,1386,1411]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Figs 1–3</figureCitation>
). Legs at most part colorless, with brown band at distal part of femur (
<figureCitation id="9EB72A51FFAD682E9D7FFA46465CFA3F" box="[828,892,1422,1447]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Fig. 1</figureCitation>
), sometimes also with brown marking at base of femur and at apex of tibia. Abdomen with variable brown and colorless areas; in all 5 examined individuals, tergum II with contrasting dark brown V-shaped middle spot (
<figureCitation id="9EB72A51FFAD682E9CB7FA1E4656FA77" box="[756,886,1494,1519]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Figs 2, 5–8</figureCitation>
); terga III–V either nearly entirely colorless (
<figureCitation id="9EB72A51FFAD682E9B2CFA1E4587F98B" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Fig. 6</figureCitation>
), or with more or less extensive brown maculation; tergum VI darkest, sometimes much darker than others (
<figureCitation id="9EB72A51FFAD682E9B29FA3245E8F9AF" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Figs 2, 6</figureCitation>
). Abdominal sterna with large brown median markings, sometimes darker than terga (
<figureCitation id="9EB72A51FFAD682E9A89F9D64033F9AF" box="[1226,1299,1566,1591]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Fig. 3</figureCitation>
). Among 5 examined larval individuals, all
<specimenCount id="108AFD5DFFAD682E9C46F98B4771F9C3" box="[517,593,1602,1627]" pageId="8" pageNumber="461" type="male">3 male</specimenCount>
larvae have abdominal tergum and sternum VI contrastingly much darker than others (
<figureCitation id="9EB72A51FFAD682E9F62F9AE449BF9E7" box="[289,443,1638,1663]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Figs 1–3, 5, 6</figureCitation>
) and both female larvae have abdominal coloration less contrasting (
<figureCitation id="9EB72A51FFAD682E9AFCF9AE4003F9E7" box="[1215,1315,1638,1663]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Figs 7, 8</figureCitation>
); possibly, sex-specific character. Cerci and paracercus at most part colorless or light ocher, each with two contrasting narrow brown bands: one at middle and another at distal part; darkening on posterior margin of each 4th segment poorly expressed (
<figureCitation id="9EB72A51FFAD682E9F57F91A44B6F973" box="[276,406,1746,1771]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="8" pageNumber="461">Figs 1, 3, 4</figureCitation>
). On light portions of cerci and paracercus swimming setae also light; on brown bands each swimming seta brown proximally, colorless distally (
<figureCitation id="9EB72A51FFAD682E9CABF93E467DF897" box="[744,861,1782,1807]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="8" pageNumber="461">Figs 9–10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAD682C9E84F8D24676FDC7" lastPageId="10" lastPageNumber="463" pageId="8" pageNumber="461" type="description">
<paragraph id="063336D4FFAD682E9E84F8D247B3F8AB" blockId="8.[151,1437,1170,2023]" box="[199,659,1818,1843]" pageId="8" pageNumber="461">
<emphasis id="34F8EAC6FFAD682E9E84F8D244C5F8AB" bold="true" box="[199,485,1818,1843]" pageId="8" pageNumber="461">Hypodermal coloration.</emphasis>
Not expressed.
</paragraph>
<paragraph id="063336D4FFAD682F9E84F8F6460AFD80" blockId="8.[151,1437,1170,2023]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="462" pageId="8" pageNumber="461">
<emphasis id="34F8EAC6FFAD682E9E84F8F6441FF8CF" bold="true" box="[199,319,1854,1879]" pageId="8" pageNumber="461">Structure.</emphasis>
Frons between antennae with flat elevation; frontal suture relatively long and obtuse-angled (
<figureCitation id="9EB72A51FFAD682E9B2CF8F64597F8E3" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="8" pageNumber="461">Fig. 37</figureCitation>
). Labrum truncate, nearly rectangular; dorsal surface with irregularly situated setae [without constant pair of submedian setae and/or pair of latero-distal setal rows –see
<taxonomicName id="C18C4D57FFAD682E9D7CF84E46F3F806" box="[831,979,1926,1950]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAD682E9D7CF84E46F3F806" box="[831,979,1926,1950]" italics="true" pageId="8" pageNumber="461">Anafroptilum</emphasis>
</taxonomicName>
(9)] (
<figureCitation id="9EB72A51FFAD682E9A55F84E414EF807" box="[1046,1134,1926,1951]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="8" pageNumber="461">Figs 12</figureCitation>
,
<figureCitation id="9EB72A51FFAD682E9A38F84E41BAF807" box="[1147,1178,1926,1951]" captionStart="FIGURES 28 – 36" captionStartId="5.[151,264,1949,1972]" captionTargetBox="[220,1301,599,1910]" captionTargetId="figure@5.[216,1351,589,1927]" captionTargetPageId="5" captionText="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="8" pageNumber="461">35</figureCitation>
). Both mandibles with incisor and kinetodontium deeply separated [see
<taxonomicName id="C18C4D57FFAD682E9CADF86246A2F85A" box="[750,898,1962,1986]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAD682E9CADF86246A2F85A" box="[750,898,1962,1986]" italics="true" pageId="8" pageNumber="461">Anafroptilum</emphasis>
</taxonomicName>
(10)]; planes of incisor, kinetodontium and prostheca perpendicular to plane of mandible (
<figureCitation id="9EB72A51FFAD682E9CE7F8064606F87F" box="[676,806,1998,2023]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="8" pageNumber="461">Figs 13–14</figureCitation>
,
<figureCitation id="9EB72A51FFAD682E9D72F807465CF87F" box="[817,892,1999,2023]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="8" pageNumber="461">38–39</figureCitation>
). On left mandible incisor with 4 thick denticles and slender ventral denticle; kinetodontium with 3 thick denticles and serration on dorsal margin; prostheca bifurcate (
<figureCitation id="9EB72A51FFAC682F9F50FF744450FF4C" box="[275,368,188,213]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="9" pageNumber="462">Fig. 38</figureCitation>
). On right mandible incisor with 3 thick apical denticles and slender ventral denticle; kinetodontium with 2 thick apical denticles and thin denticles on both margins; prostheca asymmetrically pointed (
<figureCitation id="9EB72A51FFAC682F9EDCFECC45D3FE84" box="[159,243,260,285]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="9" pageNumber="462">Fig. 39</figureCitation>
). Molar processes of left mandible longer than distal molar projection; each process terminated by elongate apical lamella (
<figureCitation id="9EB72A51FFAC682F9F02FEEF44B4FED8" box="[321,404,295,320]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 15</figureCitation>
: al). Maxilla with 3 canines, 3 dentisetae, 2–4 setae proximad of dentisetae and 9–11 setae of inner ventral row; all canines, dentisetae and setae slender, pointed and bent at the same direction (
<figureCitation id="9EB72A51FFAC682F9B47FE844079FEFC" box="[1284,1369,332,357]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 17</figureCitation>
; as in
<figureCitation id="9EB72A51FFAC682F9ED4FEA745CBFE10" box="[151,235,367,392]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="9" pageNumber="462">Fig. 77</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAC682F9F74FEB844ECFE10" box="[311,460,368,392]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9F74FEB844ECFE10" box="[311,460,368,392]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(12)]. Maxillary palp 2-segmented (
<figureCitation id="9EB72A51FFAC682F9D3AFEA746F0FE10" box="[889,976,367,392]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 17</figureCitation>
). Labium with glossae and paraglossae narrowed near bases, widened at midlength; setae of median row of glossa relatively short, shifted to dorsal side of glossa; setae of dorso-lateral row of glossa relatively short; setae of dorso-median row of paraglossa located far from median margin (
<figureCitation id="9EB72A51FFAC682F9FCDFE1444C3FE6D" box="[398,483,476,501]" captionStart="FIGURES 28 – 36" captionStartId="5.[151,264,1949,1972]" captionTargetBox="[220,1301,599,1910]" captionTargetId="figure@5.[216,1351,589,1927]" captionTargetPageId="5" captionText="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="9" pageNumber="462">Fig. 36</figureCitation>
). Labial palp with 2nd segment bearing weak muscle, 3rd segment widened apically and truncate (
<figureCitation id="9EB72A51FFAC682F9F73FE3744A3FD80" box="[304,387,511,536]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 16</figureCitation>
; as in
<figureCitation id="9EB72A51FFAC682F9F89FE37473AFD80" box="[458,538,511,536]" captionStart="FIGURE 78" captionStartId="15.[151,250,1478,1500]" captionTargetBox="[190,1386,217,1416]" captionTargetId="figure@15.[170,1417,193,1457]" captionTargetPageId="15" captionText="FIGURE 78. Anafroptilum odontostylus sp. n., labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827912/files/figure.png" pageId="9" pageNumber="462">Fig. 78</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAC682F9C1DFDC847D2FD80" box="[606,754,512,536]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9C1DFDC847D2FD80" box="[606,754,512,536]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(2)].
</paragraph>
<paragraph id="063336D4FFAC682F9E84FDEC41F3FC8D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="462">
Hind protoptera completely absent. Legs as in A.
<taxonomicName id="C18C4D57FFAC682F9D60FDEC4684FDA4" box="[803,932,548,572]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="species" species="kazlauskasi">
<emphasis id="34F8EAC6FFAC682F9D60FDEC4684FDA4" box="[803,932,548,572]" italics="true" pageId="9" pageNumber="462">kazlauskasi</emphasis>
</taxonomicName>
(
<bibRefCitation id="621D4B25FFAC682F9DFAFDEC41C1FDA4" author="Kluge" box="[953,1249,548,573]" pageId="9" pageNumber="462" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2012">
Kluge 2012:
<figureCitation id="9EB72A51FFAC682F9A1BFDEC41C1FDA4" box="[1112,1249,548,573]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="9" pageNumber="462">Figs 38–40</figureCitation>
</bibRefCitation>
): femora nearly parallel-sided, fore femur widest at middle, middle and hind femora widest at apex; apex of each femur finely serrate (as in
<figureCitation id="9EB72A51FFAC682F9F6DFDA444A2FD1D" box="[302,386,620,645]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="9" pageNumber="462">Fig. 59</figureCitation>
); each femur without stout setae on outer margin and apex; only inner margin with few small stout setae [see
<taxonomicName id="C18C4D57FFAC682F9F0BFD5844FCFD30" box="[328,476,656,680]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9F0BFD5844FCFD30" box="[328,476,656,680]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(14)]; fore tibia without patella-tibial suture, middle and hind tibia with patella-tibial suture [see
<taxonomicName id="C18C4D57FFAC682F9F5BFD7C448CFD54" box="[280,428,692,716]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9F5BFD7C448CFD54" box="[280,428,692,716]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(1)]; each tibia with short ark of fine setae at outer side near base; outer side of tibia and tarsus without stout setae, inner side of tibia and tarsus with sparse spine-like setae. Claw with two equal regular rows of numerous slender denticles, reaching 1/2 of claw length (
<figureCitation id="9EB72A51FFAC682F9D30FD3446E4FC8C" box="[883,964,764,789]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 18</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAC682F9A4BFD3441BCFC8C" box="[1032,1180,764,788]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9A4BFD3441BCFC8C" box="[1032,1180,764,788]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(5)].
</paragraph>
<paragraph id="063336D4FFAC682F9E84FCD741E4FA58" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="462">
Abdominal terga and sterna with small denticles and narrow scales; scales sparse on anterior segments and dense on II–X terga and III–IX sterna (
<figureCitation id="9EB72A51FFAC682F9C15FC8C478BFCC4" box="[598,683,836,861]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Fig. 19</figureCitation>
); sterna with irregularly situated simple long setae (as in
<figureCitation id="9EB72A51FFAC682F9B78FC8C40AFFCC5" box="[1339,1423,836,861]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="9" pageNumber="462">Fig. 67</figureCitation>
). Posterior margin of abdominal tergum I without denticles; posterior margins of terga II–VI with small blunt denticles (
<figureCitation id="9EB72A51FFAC682F9F4AFC4444A7FC3C" box="[265,391,908,933]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Fig. 19–21</figureCitation>
); posterior margins of terga VII–X with longer and pointed denticles (
<figureCitation id="9EB72A51FFAC682F9AD0FC444036FC3C" box="[1171,1302,908,933]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Figs 22–23</figureCitation>
); tergum IX without denticles behind pair of submedian pores (
<figureCitation id="9EB72A51FFAC682F9CB0FC67466BFC50" box="[755,843,943,968]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Fig. 23</figureCitation>
); tergum X as in
<taxonomicName id="C18C4D57FFAC682F9A65FC7841F7FC50" box="[1062,1239,944,968]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFAC682F9A65FC7841F7FC50" box="[1062,1239,944,968]" italics="true" pageId="9" pageNumber="462">A. odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFAC682F9AA7FC674009FC50" box="[1252,1321,943,968]" pageId="9" pageNumber="462" rank="species">
<emphasis id="34F8EAC6FFAC682F9AA7FC674009FC50" bold="true" box="[1252,1321,943,968]" pageId="9" pageNumber="462">sp. n.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="9EB72A51FFAC682F9B7FFC6740AFFC50" box="[1340,1423,943,968]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="9" pageNumber="462">Fig 58</figureCitation>
). Posterior margin of abdominal sterna I–III without denticles; posterior margin of sterna IV–VIII with very small blunt and cleft denticles (
<figureCitation id="9EB72A51FFAC682F9FF4FC3F4717FB88" box="[439,567,1015,1040]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Fig. 24–25</figureCitation>
); posterior margin of sternum IX of female larva with similar blunt and cleft denticles (
<figureCitation id="9EB72A51FFAC682F9F57FBD44452FBAD" box="[276,370,1052,1077]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Fig. 26</figureCitation>
); posterior margin of sternum IX of mature male larva with very short and shallow protogonostyli, with few very small denticles (
<figureCitation id="9EB72A51FFAC682F9CE2FBF747D4FBC0" box="[673,756,1087,1112]" captionStart="FIGURES 19 – 27" captionStartId="4.[151,264,1722,1745]" captionTargetBox="[151,1436,193,1696]" captionTargetId="figure@4.[151,1436,193,1701]" captionTargetPageId="4" captionText="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="9" pageNumber="462">Fig. 27</figureCitation>
,
<figureCitation id="9EB72A51FFAC682F9CBCFB88463FFBC0" box="[767,799,1088,1112]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="9" pageNumber="462">43</figureCitation>
). Paraprocts with pointed denticles on apex; most part of median side smooth (
<figureCitation id="9EB72A51FFAC682F9FC7FBAC44EDFBE4" box="[388,461,1124,1149]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Figs 9</figureCitation>
,
<figureCitation id="9EB72A51FFAC682F9F9BFBAC44D8FBE4" box="[472,504,1124,1148]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="9" pageNumber="462">40</figureCitation>
). Lateral denticles absent [see
<taxonomicName id="C18C4D57FFAC682F9D0EFBAC46C2FBE4" box="[845,994,1124,1148]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9D0EFBAC46C2FBE4" box="[845,994,1124,1148]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(3)]. Each tergalius I–II with costal rib, without anal rib; each tergalius III–VII with long costal and anal marginal ribs, without denticles, distal margin without rib; tergalius I very narrow, bent backward; tergalius II slightly wider; tergalii III–VI nearly symmetric, widest at distal part; tergalius VII longest, asymmetrically bent backward; each tergalius with contrasting dark brown middle macula formed by pigment granules on main trachea (
<figureCitation id="9EB72A51FFAC682F9DD9FB3C413FFA94" box="[922,1055,1268,1293]" captionStart="FIGURES 28 – 36" captionStartId="5.[151,264,1949,1972]" captionTargetBox="[220,1301,599,1910]" captionTargetId="figure@5.[216,1351,589,1927]" captionTargetPageId="5" captionText="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="9" pageNumber="462">Figs 28–34</figureCitation>
). Cerci with small denticles on all segments; one simple or composed larger spine can be present on lateral side of cercus near tip, distad of distal brown band (
<figureCitation id="9EB72A51FFAC682F9F6AFAF44488FACC" box="[297,424,1340,1365]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Figs 10, 11</figureCitation>
). Among 5 examined individuals, one has such spine on left and right cercus (
<figureCitation id="9EB72A51FFAC682F9B4DFAF4407FFACC" box="[1294,1375,1340,1365]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 11</figureCitation>
), one (
<typeStatus id="D9378876FFAC682F9EDDFA974424FAE0" box="[158,260,1375,1400]" pageId="9" pageNumber="462" type="holotype">holotype</typeStatus>
) has spine on right cercus and vestigial spine on left cercus (
<figureCitation id="9EB72A51FFAC682F9DEDFA974103FAE0" box="[942,1059,1375,1400]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Figs 9–10</figureCitation>
), one has spine on left cercus and no spine on right cercus, one has no spines on both cerci (
<figureCitation id="9EB72A51FFAC682F9D64FA4C46ABFA04" box="[807,907,1412,1437]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1929,1952]" captionTargetBox="[166,1430,195,1902]" captionTargetId="figure@2.[166,1430,195,1903]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="9" pageNumber="462">Figs 1–3</figureCitation>
), and one has no spine on right cercus and left cercus missing. Outer side of cercus with additional row of sparse fine colorless setae (
<figureCitation id="9EB72A51FFAC682F9A27FA6F4196FA58" box="[1124,1206,1447,1472]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="9" pageNumber="462">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="063336D4FFAC682F9E84FA034624F9ED" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="462">
<emphasis id="34F8EAC6FFAC682F9E84FA034767FA7D" bold="true" box="[199,583,1483,1509]" pageId="9" pageNumber="462">
<emphasis id="34F8EAC6FFAC682F9E84FA03441CFA7C" bold="true" box="[199,316,1483,1508]" italics="true" pageId="9" pageNumber="462">Subimago</emphasis>
. Cuticular coloration.
</emphasis>
Head colorless. Pronotum and prosternum light brown. Mesonotum nearly colorless with brown microtrichia, certain sutures light brown (
<figureCitation id="9EB72A51FFAC682F9D2EFA2746E4F990" box="[877,964,1519,1544]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="9" pageNumber="462">Fig. 46</figureCitation>
). Thoracic pleura and sterna at most part nearly colorless with brown microtrichia, certain sclerites light brow (
<figureCitation id="9EB72A51FFAC682F9DF6F9DC412BF9B5" box="[949,1035,1556,1581]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="9" pageNumber="462">Fig. 45</figureCitation>
). Legs nearly colorless with brown microtrichia. Abdominal terga and sterna very light brownish with brown microtrichia; terga with brownish sublateral markings. Cerci colorless with brown spines.
</paragraph>
<paragraph id="063336D4FFAC682F9E84F9B746DFF89C" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="462">
<emphasis id="34F8EAC6FFAC682F9E84F9B74408F900" bold="true" box="[199,296,1663,1688]" pageId="9" pageNumber="462">Texture.</emphasis>
On fore legs of male and female all tarsomeres covered by pointed microlepides. On middle and hind legs of both sexes proximal half of 1st tarsomere (corresponding to primary 1st tarsomere) covered by microtrichiae (as tibia), distal half of 1st tarsomere and other three tarsomeres (corresponding to 2nd–5th primary tarsomeres) covered by pointed microlepides (
<figureCitation id="9EB72A51FFAC682F9CDDF92447CFF89C" box="[670,751,1772,1797]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="9" pageNumber="462">Fig. 49</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAC682F9D70F92446E8F89C" box="[819,968,1772,1796]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAC682F9D70F92446E8F89C" box="[819,968,1772,1796]" italics="true" pageId="9" pageNumber="462">Anafroptilum</emphasis>
</taxonomicName>
(7)].
</paragraph>
<paragraph id="063336D4FFAC682C9E84F8D84792FF4D" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,608]" lastPageId="10" lastPageNumber="463" pageId="9" pageNumber="462">
<emphasis id="34F8EAC6FFAC682F9E84F8D84479F8B0" bold="true" box="[199,345,1807,1832]" italics="true" pageId="9" pageNumber="462">Imago, male</emphasis>
(
<figureCitation id="9EB72A51FFAC682F9F28F8C744E1F8B0" box="[363,449,1807,1832]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="9" pageNumber="462">Fig. 44</figureCitation>
). Turbinate eyes with stem dull reddish, grayish distally; facetted surface dull reddish. Thorax brown with membranous areas white because of fat located under cuticle. Legs pale ocher, femur with white spot (formed by internal fat) at middle and near apex. In
<typeStatus id="D9378876FFAC682F9DD1F89F46D7F8E8" box="[914,1015,1879,1904]" pageId="9" pageNumber="462" type="holotype">holotype</typeStatus>
, proportion of femur / tibia / tarsal segments of fore leg 70: 90: 3: 40: 27: 15: 13; middle leg 60: 52: 19: 6: 3: 12; hind leg 60: 47: 19: 6: 3: 12. Tarsus of middle and hind leg with one apical spine on 2nd tarsomere (primary 3rd tarsomere). Fore wing with membrane and veins colorless; pterostigma with 2–4 simple oblique veins. Hind wings absent. Abdominal segments I–VI colorless and translucent, with pair of dark brown stripes on lateral tracheal trunks; VII–X terga brown, sterna lighter. Cerci unicolor light ocher.
</paragraph>
<paragraph id="063336D4FFAF682C9E84FF1744B0FE10" blockId="10.[151,1437,151,608]" pageId="10" pageNumber="463">
Unistyligers and gonostyli colorless (
<figureCitation id="9EB72A51FFAF682C9C2DFF1747E4FF60" box="[622,708,223,248]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="10" pageNumber="463">Fig. 50</figureCitation>
). Unistyligers basally separated, with styligeral muscle attached between them (
<figureCitation id="9EB72A51FFAF682C9F0EFECC4485FE84" box="[333,421,260,285]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Fig. 41</figureCitation>
). Finger-like projection between unistyligers (
<figureCitation id="9EB72A51FFAF682C9D87FECC4100FE84" box="[964,1056,260,285]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFAF682C9A72FECC4171FE85" box="[1073,1105,260,285]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="10" pageNumber="463">50</figureCitation>
). 1st segment of gonostylus without projection; 2nd segment nearly straight; 3rd segment relatively small (
<figureCitation id="9EB72A51FFAF682C9A42FEEF4177FED8" box="[1025,1111,295,320]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFAF682C9A20FEEF41A0FED8" box="[1123,1152,295,320]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="10" pageNumber="463">50</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAF682C9A81FEE04076FED8" box="[1218,1366,296,320]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9A81FEE04076FED8" box="[1218,1366,296,320]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(15)]. Penis light brownish; external part of penis widely rounded; apical margin of penis with unusual spines bent laterally (
<figureCitation id="9EB72A51FFAF682C9F42FEA74478FE10" box="[257,344,367,392]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFAF682C9F27FEA744A3FE10" box="[356,387,367,392]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="10" pageNumber="463">51</figureCitation>
).
</paragraph>
<paragraph id="063336D4FFAF682C9E84FE5C4483FD8F" blockId="10.[151,1437,151,608]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9E84FE5C444FFE34" bold="true" box="[199,367,403,428]" italics="true" pageId="10" pageNumber="463">Imago, female</emphasis>
(
<figureCitation id="9EB72A51FFAF682C9F3DFE5C44DEFE34" box="[382,510,404,429]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="10" pageNumber="463">Fig. 47–48</figureCitation>
). Head dorsally ocher with variable light brown markings; eyes widely separated. Thorax dorsally light brown with ocher, ventrally light ocher. Tarsus of fore leg with one apical spine on 3rd tarsomere; middle and hind legs as in male. Abdomen ocher with variable brown markings on terga, with lateral brown stripes on sterna.
</paragraph>
<paragraph id="063336D4FFAF682C9E84FDEC41ABFDA5" blockId="10.[151,1437,151,608]" box="[199,1163,548,573]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9E84FDEC4421FDA3" bold="true" box="[199,257,548,571]" italics="true" pageId="10" pageNumber="463">Eggs</emphasis>
(
<figureCitation id="9EB72A51FFAF682C9F4CFDEC44B3FDA5" box="[271,403,548,573]" captionStart="FIGURES 52 – 53" captionStartId="8.[151,264,960,983]" captionTargetBox="[151,1436,425,911]" captionTargetId="figure@8.[151,1436,425,911]" captionTargetPageId="8" captionText="FIGURES 52 – 53. Anafroptilum orthostylus sp. n., eggs." httpUri="https://zenodo.org/record/827902/files/figure.png" pageId="10" pageNumber="463">Figs 52–53</figureCitation>
). About 0.12 mm length. Elongate-oval, with regular net-like relief.
</paragraph>
<paragraph id="063336D4FFAF682C9E84FD8F4676FDC7" blockId="10.[151,1437,151,608]" box="[199,854,583,608]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9E84FD8F446EFDF8" bold="true" box="[199,334,583,608]" pageId="10" pageNumber="463">Dimension.</emphasis>
Fore wing length of male and female 3.5 mm.
</paragraph>
</subSubSection>
</treatment>
</document>
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