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<mods:title id="33D3580F881661EF176286E1E87EB6D0">Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region</mods:title>
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<mods:namePart id="FC9EF161135D1D0D068027FDDA957518">Kluge, Nikita J.</mods:namePart>
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<mods:namePart id="0B963A6D4CCE8D39C40BE51CB0614107">Novikova, Eugenia A.</mods:namePart>
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<mods:date id="51B4639215ACD1E14525DD5F5F1552DE">2017</mods:date>
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<treatment id="8E2587C2FFA4682E9ED4FDA44181FEFD" ID-DOI="http://doi.org/10.5281/zenodo.6028649" ID-GBIF-Taxon="132548380" ID-Zenodo-Dep="6028649" LSID="urn:lsid:plazi:treatment:8E2587C2FFA4682E9ED4FDA44181FEFD" httpUri="http://treatment.plazi.org/id/8E2587C2FFA4682E9ED4FDA44181FEFD" lastPageId="8" lastPageNumber="461" pageId="1" pageNumber="454">
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Genus
<taxonomicName id="C18C4D57FFA468279EADFDA44707FD1E" ID-CoL="62BNC" authority="Kluge 2012" authorityName="Kluge" authorityYear="2012" box="[238,551,620,646]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279EADFDA444B3FD1E" bold="true" box="[238,403,620,646]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
<bibRefCitation id="621D4B25FFA468279FD9FDA44707FD1E" author="Kluge" box="[410,551,620,646]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2012">Kluge 2012</bibRefCitation>
</taxonomicName>
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<paragraph id="063336D4FFA468279ED4FD464439FD3F" blockId="1.[151,551,620,679]" box="[151,281,654,679]" pageId="1" pageNumber="454">
(
<figureCitation id="9EB72A51FFA468279EDCFD464430FD3F" box="[159,272,654,679]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 18" captionStart-10="FIGURES 69 – 77" captionStart-11="FIGURE 78" captionStart-12="FIGURES 79 – 81" captionStart-13="FIGURES 82 – 85" captionStart-14="FIGURES 86 – 87" captionStart-2="FIGURES 19 – 27" captionStart-3="FIGURES 28 – 36" captionStart-4="FIGURES 37 – 43" captionStart-5="FIGURES 44 – 51" captionStart-6="FIGURES 52 – 53" captionStart-7="FIGURES 54 – 59" captionStart-8="FIGURES 60 – 64" captionStart-9="FIGURES 65 – 68" captionStartId-0="2.[151,264,1929,1952]" captionStartId-1="3.[151,264,1927,1950]" captionStartId-10="14.[151,264,1940,1963]" captionStartId-11="15.[151,250,1478,1500]" captionStartId-12="16.[151,264,1498,1521]" captionStartId-13="17.[151,264,1964,1987]" captionStartId-14="18.[151,264,905,928]" captionStartId-2="4.[151,264,1722,1745]" captionStartId-3="5.[151,264,1949,1972]" captionStartId-4="6.[151,264,1794,1817]" captionStartId-5="7.[151,264,1927,1950]" captionStartId-6="8.[151,264,960,983]" captionStartId-7="11.[151,264,1943,1966]" captionStartId-8="12.[151,264,1198,1221]" captionStartId-9="13.[151,264,1895,1918]" captionTargetBox-0="[166,1430,195,1902]" captionTargetBox-1="[151,1436,193,1906]" captionTargetBox-10="[200,1394,209,1894]" captionTargetBox-11="[190,1386,217,1416]" captionTargetBox-12="[176,1371,195,1448]" captionTargetBox-13="[152,1435,769,1943]" captionTargetBox-14="[151,1436,195,884]" captionTargetBox-2="[151,1436,193,1696]" captionTargetBox-3="[220,1301,599,1910]" captionTargetBox-4="[201,1383,200,1772]" captionTargetBox-5="[151,1436,193,1906]" captionTargetBox-6="[151,1436,425,911]" captionTargetBox-7="[170,1414,193,1927]" captionTargetBox-8="[151,1436,193,1177]" captionTargetBox-9="[151,1436,193,1874]" captionTargetId-0="figure@2.[166,1430,195,1903]" captionTargetId-1="figure@3.[151,1436,193,1906]" captionTargetId-10="figure@14.[170,1417,193,1919]" captionTargetId-11="figure@15.[170,1417,193,1457]" captionTargetId-12="figure@16.[173,1410,185,1455]" captionTargetId-13="figure@17.[151,1436,769,1943]" captionTargetId-14="figure@18.[151,1436,195,884]" captionTargetId-2="figure@4.[151,1436,193,1701]" captionTargetId-3="figure@5.[216,1351,589,1927]" captionTargetId-4="figure@6.[201,1395,193,1781]" captionTargetId-5="figure@7.[151,1436,193,1906]" captionTargetId-6="figure@8.[151,1436,425,911]" captionTargetId-7="figure@11.[170,1414,193,1927]" captionTargetId-8="figure@12.[151,1436,193,1177]" captionTargetId-9="figure@13.[151,1436,193,1874]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="14" captionTargetPageId-11="15" captionTargetPageId-12="16" captionTargetPageId-13="17" captionTargetPageId-14="18" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURES 1 – 8. Anafroptilum orthostylus sp. n., larvae. 1 – 3, male larva: lateral, dorsal and ventral view; 4 – 8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5 – 8, abdominal terga (left) and sterna (right): 5 – 6, males; 7 – 8, females (5, holotype). Abbreviations: I – X, abdominal segments I – X." captionText-1="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." captionText-10="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." captionText-11="FIGURE 78. Anafroptilum odontostylus sp. n., labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." captionText-12="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." captionText-13="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." captionText-14="FIGURES 86 – 87. Anafroptilum spp., median portion of posterior margin of abdominal tergum IX of larva. 86, Anafroptilum kazlauslasi (holotype); 87, Anafroptilum bifurcatum (USA, Nebraska, Cherry Co., R + 61, Snake River, 6. VI. 1995, coll. B. Kondratieff)." captionText-2="FIGURES 19 – 27. Anafroptilum orthostylus sp. n., eXuviae of larval abdomen. 19, median part of tergum IV; 20 – 23, posterior margin of terga V, VI, VII and IX; 24 – 26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20 – 22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV – sIX, sterna V – IX; tIV – tIX, terga IV – IX." captionText-3="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." captionText-4="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." captionText-5="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." captionText-6="FIGURES 52 – 53. Anafroptilum orthostylus sp. n., eggs." captionText-7="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." captionText-8="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." captionText-9="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri-0="https://zenodo.org/record/827890/files/figure.png" httpUri-1="https://zenodo.org/record/827892/files/figure.png" httpUri-10="https://zenodo.org/record/827910/files/figure.png" httpUri-11="https://zenodo.org/record/827912/files/figure.png" httpUri-12="https://zenodo.org/record/827914/files/figure.png" httpUri-13="https://zenodo.org/record/827916/files/figure.png" httpUri-14="https://zenodo.org/record/827918/files/figure.png" httpUri-2="https://zenodo.org/record/827894/files/figure.png" httpUri-3="https://zenodo.org/record/827896/files/figure.png" httpUri-4="https://zenodo.org/record/827898/files/figure.png" httpUri-5="https://zenodo.org/record/827900/files/figure.png" httpUri-6="https://zenodo.org/record/827902/files/figure.png" httpUri-7="https://zenodo.org/record/827904/files/figure.png" httpUri-8="https://zenodo.org/record/827906/files/figure.png" httpUri-9="https://zenodo.org/record/827908/files/figure.png" pageId="1" pageNumber="454">Figs 1–87</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFA468279ED4FD1E44F5FD74" box="[151,469,726,748]" pageId="1" pageNumber="454" type="reference_group">
<paragraph id="063336D4FFA468279ED4FD1E44F5FD74" blockId="1.[151,683,726,779]" box="[151,469,726,748]" pageId="1" pageNumber="454">
<treatmentCitationGroup id="269C11FAFFA468279ED4FD1E44F5FD74" box="[151,469,726,748]" pageId="1" pageNumber="454">
<taxonomicName id="C18C4D57FFA468279ED4FD1E44F0FD74" ID-CoL="62BNC" authority="Kluge 2012: 372" authorityName="Kluge" authorityPageNumber="372" authorityYear="2012" box="[151,464,726,748]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279ED4FD1E443DFD74" box="[151,285,726,748]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
<treatmentCitation id="872D10C5FFA468279F60FD1E44F0FD74" author="Kluge" box="[291,464,726,748]" page="372" pageId="1" pageNumber="454" year="2012">
<bibRefCitation id="621D4B25FFA468279F60FD1E44BEFD74" author="Kluge" box="[291,414,726,748]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2012">Kluge 2012</bibRefCitation>
: 372
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFA468279ED4FD3E40A8FC00" pageId="1" pageNumber="454" type="materials_examined">
<paragraph id="063336D4FFA468279ED4FD3E478BFC93" blockId="1.[151,683,726,779]" box="[151,683,757,779]" pageId="1" pageNumber="454">
Type species:
<taxonomicName id="C18C4D57FFA468279F65FD3D4787FC93" authority="Kluge 1983" authorityName="Kluge" authorityYear="1983" box="[294,679,757,779]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="kazlauskasi">
<emphasis id="34F8EAC6FFA468279F65FD3D4707FC93" box="[294,551,757,779]" italics="true" pageId="1" pageNumber="454">Centroptilum kazlauskasi</emphasis>
<bibRefCitation id="621D4B25FFA468279C6EFD3D4787FC93" author="Kluge" box="[557,679,757,779]" pageId="1" pageNumber="454" refString="Kluge, N. J. (1983) [New and little known mayflies of the family Baetidae (Ephemeroptera) from Primorye Territiry]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 61 (1), 65 - 79. [in Russian, English translation: Entomoljgical Review, 62 (1), 53 - 68.]" type="journal article" year="1983">Kluge 1983</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="063336D4FFA468279ED4FCFF40A8FC00" blockId="1.[151,1437,823,2037]" pageId="1" pageNumber="454">
<emphasis id="34F8EAC6FFA468279ED4FCFF4660FCC8" bold="true" box="[151,832,823,848]" pageId="1" pageNumber="454">
Systematic position and characteristics of
<taxonomicName id="C18C4D57FFA468279CDFFCFF4619FCC8" box="[668,825,823,848]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279CDFFCFF4619FCC8" bold="true" box="[668,825,823,848]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
.
</emphasis>
Systematic position of
<taxonomicName id="C18C4D57FFA468279A16FCF041C9FCC8" box="[1109,1257,824,848]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279A16FCF041C9FCC8" box="[1109,1257,824,848]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
is vague: from one hand, it shares some peculiar characters common with Cloeonini
<emphasis id="34F8EAC6FFA468279D87FC954122FCEC" box="[964,1026,861,884]" italics="true" pageId="1" pageNumber="454">sensu</emphasis>
<bibRefCitation id="621D4B25FFA468279A4EFC944185FCED" author="Kluge" box="[1037,1189,860,885]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2016 b) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. http: // dX. doi. org / 10.11646 / ZootaXa. 4107.4.2" type="journal article" year="2016" yearSuffix="b">Kluge 2016b</bibRefCitation>
; from other hand, its patella-tibial suture is characteristic for the taxon Protopatellata, while Cloeonini do not belong to Protopatellata.
</paragraph>
</subSubSection>
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<paragraph id="063336D4FFA468279E84FC6C44EFFB67" blockId="1.[151,1437,823,2037]" pageId="1" pageNumber="454">
<emphasis id="34F8EAC6FFA468279E84FC6C46BAFC25" bold="true" box="[199,922,931,957]" pageId="1" pageNumber="454">
Character of
<taxonomicName id="C18C4D57FFA468279F28FC6B4728FC24" box="[363,520,931,956]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279F28FC6B4728FC24" bold="true" box="[363,520,931,956]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
common with Protopatellata. (1)
</emphasis>
In larva, subimago and imago of both sexes patella-tibial suture is absent on fore legs and present on middle and hind legs (
<bibRefCitation id="621D4B25FFA468279A77FC0F4073FC78" author="Kluge" box="[1076,1363,967,992]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2012">
Kluge 2012:
<figureCitation id="9EB72A51FFA468279A8EFC0F4073FC78" box="[1229,1363,967,992]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="1" pageNumber="454">Figs 13, 14</figureCitation>
</bibRefCitation>
). This character is plesiomorphic, being the same as in the outgroup, i.e. in the most non-baetid mayflies and in the primitive baetid taxa,
<taxonomicName id="C18C4D57FFA468279FCCFBD847F3FBB0" authority="Penniket 1962" authorityName="Penniket" authorityYear="1962" box="[399,723,1039,1064]" class="Insecta" family="Siphlaenigmatidae" genus="Siphlaenigma" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279FCCFBD84709FBB0" box="[399,553,1040,1064]" italics="true" pageId="1" pageNumber="454">Siphlaenigma</emphasis>
<bibRefCitation id="621D4B25FFA468279C72FBC747F3FBB0" author="Penniket" box="[561,723,1039,1064]" pageId="1" pageNumber="454" refString="Penniket, J. G. (1962) Notes on New Zealand Ephemeroptera. III. A new family, genus and species. Records of the Canterbury Museum, 7 (5), 389 - 398." type="journal article" year="1962">Penniket 1962</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="C18C4D57FFA468279D48FBD841F2FBB0" authority="Kluge 1997 (Kluge 1997)" authorityName="Kluge 1997 (Kluge" authorityYear="1997" box="[779,1234,1039,1064]" class="Insecta" family="Baetidae" genus="Palaeocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279D48FBD84685FBB0" box="[779,933,1040,1064]" italics="true" pageId="1" pageNumber="454">Palaeocloeon</emphasis>
<bibRefCitation id="621D4B25FFA468279DEFFBC74111FBB0" author="Kluge" box="[940,1073,1039,1064]" pageId="1" pageNumber="454" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535. [Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne]" type="journal volume" year="1997">Kluge 1997</bibRefCitation>
(
<bibRefCitation id="621D4B25FFA468279A03FBC741EAFBB0" author="Kluge" box="[1088,1226,1039,1064]" pageId="1" pageNumber="454" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535. [Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne]" type="journal volume" year="1997">Kluge 1997</bibRefCitation>
)
</taxonomicName>
. Among
<taxonomicName id="C18C4D57FFA468279B79FBC740BCFBB0" box="[1338,1436,1039,1064]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
in the narrow sense, i.e. Turbanoculata
<bibRefCitation id="621D4B25FFA468279C1AFBFC47C2FBD4" author="Kluge" box="[601,738,1076,1101]" pageId="1" pageNumber="454" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535. [Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne]" type="journal volume" year="1997">Kluge 1997</bibRefCitation>
, this character is present in the taxon Protopatellata
<bibRefCitation id="621D4B25FFA468279B7AFBFC4468FBE8" author="Kluge" pageId="1" pageNumber="454" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taXon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
(= Afroptilinae
<emphasis id="34F8EAC6FFA468279C4DFB91476CFBE8" box="[526,588,1113,1136]" italics="true" pageId="1" pageNumber="454">sensu</emphasis>
<bibRefCitation id="621D4B25FFA468279C1BFB9F47C5FBE8" author="Kluge" box="[600,741,1111,1136]" pageId="1" pageNumber="454" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535. [Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne]" type="journal volume" year="1997">Kluge 1997</bibRefCitation>
). In contrast to this, the taxon Anteropatellata
<bibRefCitation id="621D4B25FFA468279B57FB9F40BCFBE8" author="Kluge" box="[1300,1436,1111,1136]" pageId="1" pageNumber="454" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535. [Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne]" type="journal volume" year="1997">Kluge 1997</bibRefCitation>
(comprising overwhelming majority of
<taxonomicName id="C18C4D57FFA468279C1EFBB447E1FB0D" box="[605,705,1148,1173]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
) is characterized by presence of patella-tibial suture on all legs other than fore legs of male imago and subimago. Exceptions are made only by baetids with highly modified leg structure; unlike them,
<taxonomicName id="C18C4D57FFA468279FEBFB0C471CFB44" box="[424,572,1220,1244]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279FEBFB0C471CFB44" box="[424,572,1220,1244]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
has leg structure non-modified, similar to that of Cloeonini
<emphasis id="34F8EAC6FFA468279B4EFB0D406BFB44" box="[1293,1355,1221,1244]" italics="true" pageId="1" pageNumber="454">sensu</emphasis>
<bibRefCitation id="621D4B25FFA468279B14FB0C45FDFA98" author="Kluge" pageId="1" pageNumber="454" refString="Kluge, N. J. (2016 b) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. http: // dX. doi. org / 10.11646 / ZootaXa. 4107.4.2" type="journal article" year="2016" yearSuffix="b">Kluge 2016b</bibRefCitation>
and many other taxa.
</paragraph>
<paragraph id="063336D4FFA468279E84FAC441D3F94D" blockId="1.[151,1437,823,2037]" pageId="1" pageNumber="454">
<emphasis id="34F8EAC6FFA468279E84FAC4466FFABD" bold="true" box="[199,847,1291,1317]" pageId="1" pageNumber="454">
Characters of
<taxonomicName id="C18C4D57FFA468279F39FAC34737FABC" box="[378,535,1291,1316]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279F39FAC34737FABC" bold="true" box="[378,535,1291,1316]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
common with Cloeonini.
</emphasis>
The taxon Cloeonini
<emphasis id="34F8EAC6FFA468279A17FAC541B2FABC" box="[1108,1170,1293,1316]" italics="true" pageId="1" pageNumber="454">sensu</emphasis>
<bibRefCitation id="621D4B25FFA468279ADDFAC44014FABD" author="Kluge" box="[1182,1332,1292,1317]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2016 b) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. http: // dX. doi. org / 10.11646 / ZootaXa. 4107.4.2" type="journal article" year="2016" yearSuffix="b">Kluge 2016b</bibRefCitation>
includes genera
<taxonomicName id="C18C4D57FFA468279EABFAF844E3FAD0" authority="Leach 1815" authorityName="Leach" authorityYear="1815" box="[232,451,1327,1352]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279EABFAF84419FAD0" box="[232,313,1328,1352]" italics="true" pageId="1" pageNumber="454">Cloeon</emphasis>
<bibRefCitation id="621D4B25FFA468279F03FAE744E3FAD0" author="Leach" box="[320,451,1327,1352]" pageId="1" pageNumber="454" refString="Leach, W. E. (1815) Entomology. Brewster's Edinburg Encyclopedia, 1 st Edition, 9 (1), 57 - 172." type="journal article" year="1815">Leach 1815</bibRefCitation>
</taxonomicName>
(in narrowest sense),
<taxonomicName id="C18C4D57FFA468279CFAFAF846C9FAD0" authority="Bengtsson 1915" authorityName="Bengtsson" authorityYear="1915" box="[697,1001,1327,1352]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279CFAFAF8460DFAD0" box="[697,813,1328,1352]" italics="true" pageId="1" pageNumber="454">Procloeon</emphasis>
<bibRefCitation id="621D4B25FFA468279D70FAF846C9FAD0" author="Bengtsson" box="[819,1001,1327,1352]" pageId="1" pageNumber="454" refString="Bengtsson, S. (1915) Eine Namensanderung. Entomologisk Tidskrift, 36 (1), 34." type="journal article" year="1915">Bengtsson 1915</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C18C4D57FFA468279DB7FAF840BCFAD0" authority="Kluge &amp; Novikova 1992" authorityName="Kluge &amp; Novikova" authorityYear="1992" box="[1012,1436,1327,1352]" class="Insecta" family="Baetidae" genus="Similicloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279DB7FAF841A0FAD0" box="[1012,1152,1328,1352]" italics="true" pageId="1" pageNumber="454">Similicloeon</emphasis>
<bibRefCitation id="621D4B25FFA468279AC4FAE740BCFAD0" author="Kluge" box="[1159,1436,1327,1352]" pageId="1" pageNumber="454" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83 (in Russian). English translation: Entomological Review, 71 (9), 29 - 54." type="journal article" year="1992">Kluge &amp; Novikova 1992</bibRefCitation>
</taxonomicName>
and related taxa (
<bibRefCitation id="621D4B25FFA468279F22FA9C44DCFAF5" author="Kluge" box="[353,508,1364,1389]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2016 b) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. http: // dX. doi. org / 10.11646 / ZootaXa. 4107.4.2" type="journal article" year="2016" yearSuffix="b">Kluge 2016b</bibRefCitation>
). In contrast to
<taxonomicName id="C18C4D57FFA468279CF5FA9C466AFAF4" box="[694,842,1364,1388]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279CF5FA9C466AFAF4" box="[694,842,1364,1388]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
, all representatives of Cloeonini have patella-tibial suture equally developed on all legs of larva and female, so Cloeonini undoubtedly belong to Anteropatellata (see above). Besides evident symplesiomorphies,
<taxonomicName id="C18C4D57FFA468279CD7FA544608FA2C" box="[660,808,1436,1460]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279CD7FA544608FA2C" box="[660,808,1436,1460]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
and Cloeonini share the following common characters:
<emphasis id="34F8EAC6FFA468279ED4FA774598FA40" bold="true" box="[151,184,1471,1496]" pageId="1" pageNumber="454">(2)</emphasis>
Larval labial palp with 3rd segment truncate and 2nd segment non-projected (
<figureCitation id="9EB72A51FFA468279A77FA7741ACFA40" box="[1076,1164,1471,1496]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="1" pageNumber="454">Figs 16</figureCitation>
,
<figureCitation id="9EB72A51FFA468279ADAFA084199FA40" box="[1177,1209,1472,1496]" captionStart="FIGURE 78" captionStartId="15.[151,250,1478,1500]" captionTargetBox="[190,1386,217,1416]" captionTargetId="figure@15.[170,1417,193,1457]" captionTargetPageId="15" captionText="FIGURE 78. Anafroptilum odontostylus sp. n., labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827912/files/figure.png" pageId="1" pageNumber="454">78</figureCitation>
). The same in some other taxa.
<emphasis id="34F8EAC6FFA468279F54FA2C4418FA65" bold="true" box="[279,312,1508,1533]" pageId="1" pageNumber="454">(3)</emphasis>
Larval abdominal segments VIII and IX with spines forming one longitudinal row on each lateral margin; sometimes such spines are present also on some of previous abdominal segments. This character is constant for Cloeonini and not found in any taxa other than Cloeonini and
<taxonomicName id="C18C4D57FFA468279DBDF9E441B2F9DC" box="[1022,1170,1580,1604]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279DBDF9E441B2F9DC" box="[1022,1170,1580,1604]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
. Among
<taxonomicName id="C18C4D57FFA468279B42F9E440B5F9DC" box="[1281,1429,1580,1604]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279B42F9E440B5F9DC" box="[1281,1429,1580,1604]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
, these spines are either constantly present (e.g. in
<taxonomicName id="C18C4D57FFA468279CFBF9984677F9F0" box="[696,855,1616,1640]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="kazlauskasi">
<emphasis id="34F8EAC6FFA468279CFBF9984677F9F0" box="[696,855,1616,1640]" italics="true" pageId="1" pageNumber="454">A. kazlauskasi</emphasis>
</taxonomicName>
,
<taxonomicName id="C18C4D57FFA468279D26F99846DEF9F0" box="[869,1022,1616,1640]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="bifurcatum">
<emphasis id="34F8EAC6FFA468279D26F99846DEF9F0" box="[869,1022,1616,1640]" italics="true" pageId="1" pageNumber="454">A. bifurcatum</emphasis>
</taxonomicName>
,
<taxonomicName id="C18C4D57FFA468279A48F99841ADF9FF" box="[1035,1165,1616,1639]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="34F8EAC6FFA468279A48F99841ADF9FF" box="[1035,1165,1616,1639]" italics="true" pageId="1" pageNumber="454">A. victoriae</emphasis>
</taxonomicName>
), or occasionally absent (e.g., in
<taxonomicName id="C18C4D57FFA468279EB1F9BC4477F914" box="[242,343,1652,1676]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="album">
<emphasis id="34F8EAC6FFA468279EB1F9BC4477F914" box="[242,343,1652,1676]" italics="true" pageId="1" pageNumber="454">A. album</emphasis>
</taxonomicName>
,
<taxonomicName id="C18C4D57FFA468279F27F9BC4732F914" box="[356,530,1652,1676]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="conturbatum">
<emphasis id="34F8EAC6FFA468279F27F9BC4732F914" box="[356,530,1652,1676]" italics="true" pageId="1" pageNumber="454">A. conturbatum</emphasis>
</taxonomicName>
) (
<bibRefCitation id="621D4B25FFA468279C6AF9BC4672F914" author="Lowen" box="[553,850,1652,1677]" pageId="1" pageNumber="454" refString="Lowen, R. G. &amp; Flannagan, J. F. (1991) Four Manitoba species of Centroptilum Eaton (Ephemeroptera: Baetidae) with remarks on the genus. In: Alba-Tercedor, J. &amp; SancheZ-Ortega, A. (Eds.), Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 189 - 205. [Proc. 6 th Int. Congf. Ephemeroptera &amp; 10 th Int. Symp. Plecoptera, 24 - 30 July 1989, Granada, Spain]" type="journal volume" year="1991">Lowen &amp; Flannagan 1991</bibRefCitation>
). In the two new species from
<collectingCountry id="7E9B7644FFA468279AE6F9BC402BF915" box="[1189,1291,1652,1677]" name="Thailand" pageId="1" pageNumber="454">Thailand</collectingCountry>
, these spines are either poorly developed, or absent (see below).
<emphasis id="34F8EAC6FFA468279C92F95F47D1F928" bold="true" box="[721,753,1687,1712]" pageId="1" pageNumber="454">(4)</emphasis>
Male imaginal gonovectes are fused with penial bridge (
<figureCitation id="9EB72A51FFA468279B29F95F4595F94C" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="1" pageNumber="454">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFA468279E82F97445C1F94C" box="[193,225,1724,1748]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="1" pageNumber="454">79</figureCitation>
). Among Protopatellata, the same in the Afrotropical taxon Rhithrocloeoninae (
<bibRefCitation id="621D4B25FFA468279A1EF97441C7F94D" author="Kluge" box="[1117,1255,1724,1749]" pageId="1" pageNumber="454" refString="Kluge, N. J. (2015) First description of winged stages of Thraulobaetodes Elouard &amp; HideuX 1991 and reclassification of Rhithrocloeoninae (Ephemeroptera, Baetidae). Zootaxa, 3949 (4), 491 - 514. http: // dX. doi. org / 10.11646 / ZootaXa. 3949.4.2" type="journal article" year="2015">Kluge 2015</bibRefCitation>
).
</paragraph>
<paragraph id="063336D4FFA468279E84F9174180F834" blockId="1.[151,1437,823,2037]" pageId="1" pageNumber="454">
<emphasis id="34F8EAC6FFA468279E84F9174069F960" bold="true" box="[199,1353,1759,1784]" pageId="1" pageNumber="454">
Symplesiomorphies of
<taxonomicName id="C18C4D57FFA468279F9FF9174631F960" authority="Cloeoninae" authorityName="Cloeoninae" box="[476,785,1759,1784]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279F9FF9174759F960" bold="true" box="[476,633,1759,1784]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
, Cloeoninae
</taxonomicName>
, some Protopatellata and some other taxa. (5)
</emphasis>
Larval claw with 2 equal rows of denticles (in contrast to Baetungulata and some other taxa).
<emphasis id="34F8EAC6FFA468279A3AF8CC41B9F885" bold="true" box="[1145,1177,1796,1821]" pageId="1" pageNumber="454">(6)</emphasis>
Fore wing with single marginal intercalaries (in contrast to Baetovectata).
<emphasis id="34F8EAC6FFA468279C9AF8EF47DAF8D8" bold="true" box="[729,762,1831,1856]" pageId="1" pageNumber="454">(7)</emphasis>
In subimago of both sexes either all segments of all tarsi are covered with pointed microlepides, or only proximal part of tarsus is covered with microtrichiae (
<figureCitation id="9EB72A51FFA468279AB5F884406DF8FC" box="[1270,1357,1868,1893]" captionStart="FIGURES 44 – 51" captionStartId="7.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 44 – 51. Anafroptilum orthostylus sp. n. 44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47 – 48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45 – 46, 50 – 51, holotype)." httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="1" pageNumber="454">Fig. 49</figureCitation>
); blunt microlepides are absent on all tarsomeres (in contrast to Baetungulata).
<emphasis id="34F8EAC6FFA468279DA4F8A74128F810" bold="true" box="[999,1032,1903,1928]" pageId="1" pageNumber="454">(8)</emphasis>
Subimaginal gonostyli developing under larval cuticle are folded in «
<taxonomicName id="C18C4D57FFA468279C5BF85C4749F834" box="[536,617,1940,1964]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279C5BF85C4749F834" box="[536,617,1940,1964]" italics="true" pageId="1" pageNumber="454">Cloeon</emphasis>
</taxonomicName>
-
<typeStatus id="D9378876FFA468279C32F85D4784F835" box="[625,676,1941,1965]" pageId="1" pageNumber="454">type</typeStatus>
» pose (
<figureCitation id="9EB72A51FFA468279CB8F85C466CF834" box="[763,844,1940,1965]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="1" pageNumber="454">Fig. 43</figureCitation>
) (in contrast to Baetovectata).
</paragraph>
<paragraph id="063336D4FFA468279E84F87040BCF86D" blockId="1.[151,1437,823,2037]" pageId="1" pageNumber="454">
<emphasis id="34F8EAC6FFA468279E84F8704699F849" bold="true" box="[199,953,1975,2001]" pageId="1" pageNumber="454">
Characters of
<taxonomicName id="C18C4D57FFA468279F34F87F4734F848" box="[375,532,1975,2000]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA468279F34F87F4734F848" bold="true" box="[375,532,1975,2000]" italics="true" pageId="1" pageNumber="454">Anafroptilum</emphasis>
</taxonomicName>
of unclear phylogenetic status. (9)
</emphasis>
Dorsal surface of labrum with irregularly situated setae, without constant pair of submedian setae and without regular latero-distal setal rows (
<figureCitation id="9EB72A51FFA468279AB9F8144075F86D" box="[1274,1365,2012,2037]" captionStart="FIGURES 28 – 36" captionStartId="5.[151,264,1949,1972]" captionTargetBox="[220,1301,599,1910]" captionTargetId="figure@5.[216,1351,589,1927]" captionTargetPageId="5" captionText="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="1" pageNumber="454">Figs 35</figureCitation>
,
<figureCitation id="9EB72A51FFA468279B33F81440B0F86D" box="[1392,1424,2012,2037]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="1" pageNumber="454">76</figureCitation>
).
</paragraph>
<caption id="52F3665CFFA768249ED4F84147DBF845" httpUri="https://zenodo.org/record/827890/files/figure.png" pageId="2" pageNumber="455" targetBox="[166,1430,195,1902]" targetPageId="2">
<paragraph id="063336D4FFA768249ED4F84147DBF845" blockId="2.[151,1435,1929,2013]" pageId="2" pageNumber="455">
<emphasis id="34F8EAC6FFA768249ED4F841441BF806" bold="true" box="[151,315,1929,1952]" pageId="2" pageNumber="455">FIGURES 1–8.</emphasis>
<taxonomicName id="C18C4D57FFA768249F01F8414761F807" box="[322,577,1929,1951]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="455" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA768249F01F8414761F807" box="[322,577,1929,1951]" italics="true" pageId="2" pageNumber="455">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA768249C04F84247A1F807" box="[583,641,1930,1951]" pageId="2" pageNumber="455" rank="species">
<emphasis id="34F8EAC6FFA768249C04F84247A1F807" bold="true" box="[583,641,1930,1951]" pageId="2" pageNumber="455">sp. n.</emphasis>
</taxonomicNameLabel>
, larvae. 1–3, male larva: lateral, dorsal and ventral view; 4–8, eXuviae of last instar larvae: 4, left half of pronotum and mesonotum; 5–8, abdominal terga (left) and sterna (right): 5–6, males; 7–8, females (5, holotype). Abbreviations: I–X, abdominal segments I–X.
</paragraph>
</caption>
<caption id="52F3665CFFA668259ED4F84F41C8F843" httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="3" pageNumber="456" targetBox="[151,1436,193,1906]" targetPageId="3">
<paragraph id="063336D4FFA668259ED4F84F41C8F843" blockId="3.[151,1436,1927,2011]" pageId="3" pageNumber="456">
<emphasis id="34F8EAC6FFA668259ED4F84F446AF805" bold="true" box="[151,330,1927,1950]" pageId="3" pageNumber="456">FIGURES 9–18.</emphasis>
<taxonomicName id="C18C4D57FFA668259F10F8404773F806" box="[339,595,1928,1950]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA668259F10F8404773F806" box="[339,595,1928,1950]" italics="true" pageId="3" pageNumber="456">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA668259C1FF84047B7F805" box="[604,663,1928,1949]" pageId="3" pageNumber="456" rank="species">
<emphasis id="34F8EAC6FFA668259C1FF84047B7F805" bold="true" box="[604,663,1928,1949]" pageId="3" pageNumber="456">sp. n.</emphasis>
</taxonomicNameLabel>
, larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13–15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9–10 and 12–15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection.
</paragraph>
</caption>
<caption id="52F3665CFFA168229ED4F97247CFF8B5" httpUri="https://zenodo.org/record/827894/files/figure.png" pageId="4" pageNumber="457" targetBox="[151,1436,193,1696]" targetPageId="4">
<paragraph id="063336D4FFA168229ED4F97247CFF8B5" blockId="4.[151,1436,1722,1837]" pageId="4" pageNumber="457">
<emphasis id="34F8EAC6FFA168229ED4F9724474F948" bold="true" box="[151,340,1722,1745]" pageId="4" pageNumber="457">FIGURES 19–27.</emphasis>
<taxonomicName id="C18C4D57FFA168229F19F9734778F949" box="[346,600,1723,1745]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA168229F19F9734778F949" box="[346,600,1723,1745]" italics="true" pageId="4" pageNumber="457">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA168229C1DF97347B7F948" box="[606,663,1723,1744]" pageId="4" pageNumber="457" rank="species">
<emphasis id="34F8EAC6FFA168229C1DF97347B7F948" bold="true" box="[606,663,1723,1744]" pageId="4" pageNumber="457">sp. n.</emphasis>
</taxonomicNameLabel>
, eXuviae of larval abdomen. 19, median part of tergum IV; 20–23, posterior margin of terga V, VI, VII and IX; 24–26, posterior margin of sterna V, VIII and sternum IX of female; 27, sternum IX of male with protogonostyli (20–22, 27, holotype). Arrow shows paired submedian pore near posterior margin of tergum IX; abbreviations: sV–sIX, sterna V–IX; tIV–tIX, terga IV–IX.
</paragraph>
</caption>
<paragraph id="063336D4FFA168229ED4F89140AFF80E" blockId="4.[151,1436,1881,2015]" pageId="4" pageNumber="457">
The same in some Cloeonini and in
<taxonomicName id="C18C4D57FFA168229C7EF89247F1F8EA" box="[573,721,1882,1906]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA168229C7EF89247F1F8EA" box="[573,721,1882,1906]" italics="true" pageId="4" pageNumber="457">Centroptilum</emphasis>
</taxonomicName>
.
<emphasis id="34F8EAC6FFA168229CAFF891463BF8EA" bold="true" box="[748,795,1881,1906]" pageId="4" pageNumber="457">(10)</emphasis>
Both mandibles with incisor and kinetodontium deeply separated (
<figureCitation id="9EB72A51FFA168229F4CF8B544B1F80E" box="[271,401,1917,1942]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="4" pageNumber="457">Figs 13–14</figureCitation>
,
<figureCitation id="9EB72A51FFA168229FDFF8B644C5F80E" box="[412,485,1918,1942]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="4" pageNumber="457">38–39</figureCitation>
,
<figureCitation id="9EB72A51FFA168229FB3F8B5471AF80E" box="[496,570,1917,1942]" captionStart="FIGURES 60 – 64" captionStartId="12.[151,264,1198,1221]" captionTargetBox="[151,1436,193,1177]" captionTargetId="figure@12.[151,1436,193,1177]" captionTargetPageId="12" captionText="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="4" pageNumber="457">61–62</figureCitation>
). The same in
<taxonomicName id="C18C4D57FFA168229C9EF8B64651F80E" box="[733,881,1918,1942]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA168229C9EF8B64651F80E" box="[733,881,1918,1942]" italics="true" pageId="4" pageNumber="457">Centroptilum</emphasis>
</taxonomicName>
, selected species of
<taxonomicName id="C18C4D57FFA168229A14F8B641EBF80E" box="[1111,1227,1918,1942]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA168229A14F8B641EBF80E" box="[1111,1227,1918,1942]" italics="true" pageId="4" pageNumber="457">Procloeon</emphasis>
</taxonomicName>
and some others.
</paragraph>
<paragraph id="063336D4FFA168239E84F86945C6FD80" blockId="4.[151,1436,1881,2015]" lastBlockId="5.[151,1437,151,536]" lastPageId="5" lastPageNumber="458" pageId="4" pageNumber="457">
<emphasis id="34F8EAC6FFA168229E84F86945D3F822" bold="true" box="[199,243,1953,1978]" pageId="4" pageNumber="457">(11)</emphasis>
Both mandibles with dense setae between prostheca and mola (
<figureCitation id="9EB72A51FFA168229DA8F8694151F822" box="[1003,1137,1953,1978]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="4" pageNumber="457">Figs 13–14</figureCitation>
,
<figureCitation id="9EB72A51FFA168229AC2F86941ECF822" box="[1153,1228,1953,1978]" captionStart="FIGURES 60 – 64" captionStartId="12.[151,264,1198,1221]" captionTargetBox="[151,1436,193,1177]" captionTargetId="figure@12.[151,1436,193,1177]" captionTargetPageId="12" captionText="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="4" pageNumber="457">61–63</figureCitation>
). The same in all Cloeonini,
<taxonomicName id="C18C4D57FFA168229F59F80E448FF846" box="[282,431,1990,2014]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA168229F59F80E448FF846" box="[282,431,1990,2014]" italics="true" pageId="4" pageNumber="457">Centroptilum</emphasis>
</taxonomicName>
and some others.
<emphasis id="34F8EAC6FFA168229CD0F80D47E1F846" bold="true" box="[659,705,1989,2014]" pageId="4" pageNumber="457">(12)</emphasis>
Maxilla of the «
<taxonomicName id="C18C4D57FFA168229DD5F80E46C7F846" box="[918,999,1990,2014]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA168229DD5F80E46C7F846" box="[918,999,1990,2014]" italics="true" pageId="4" pageNumber="457">Cloeon</emphasis>
</taxonomicName>
-
<typeStatus id="D9378876FFA168229DACF80F4104F847" box="[1007,1060,1991,2015]" pageId="4" pageNumber="457">type</typeStatus>
», i.e. with all 3 canines and 3 dentisetae slender, pointed and bent at the same direction (
<figureCitation id="9EB72A51FFA068239D68FF5F46A2FF28" box="[811,898,151,176]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1927,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@3.[151,1436,193,1906]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Anafroptilum orthostylus sp. n., larvae. 9, tenth abdominal segment and caudalii; 10, enlarged portion of cerci; 11, portion of cercus of another specimen; 12, labrum; 13 – 15, left and right mandibles; 16, labium; 17, maXilla; 18, claw (9 – 10 and 12 – 15, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827892/files/figure.png" pageId="5" pageNumber="458">Figs 17</figureCitation>
,
<figureCitation id="9EB72A51FFA068239DCCFF50468FFF28" box="[911,943,152,176]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="5" pageNumber="458">77</figureCitation>
). The same in many other taxa.
<emphasis id="34F8EAC6FFA068239B5AFF5F4068FF28" bold="true" box="[1305,1352,151,176]" pageId="5" pageNumber="458">(13)</emphasis>
Glossa and paraglossa of the «
<taxonomicName id="C18C4D57FFA068239FD8FF7444CCFF4C" box="[411,492,188,212]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA068239FD8FF7444CCFF4C" box="[411,492,188,212]" italics="true" pageId="5" pageNumber="458">Cloeon</emphasis>
</taxonomicName>
-
<typeStatus id="D9378876FFA068239FB0FF754708FF4D" box="[499,552,189,213]" pageId="5" pageNumber="458">type</typeStatus>
», i.e. subequal in length and width, pointed, paraglossa with no more than one regular row on outer-apical margin (
<figureCitation id="9EB72A51FFA068239C6DFF1747A5FF60" box="[558,645,223,248]" captionStart="FIGURES 28 – 36" captionStartId="5.[151,264,1949,1972]" captionTargetBox="[220,1301,599,1910]" captionTargetId="figure@5.[216,1351,589,1927]" captionTargetPageId="5" captionText="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="5" pageNumber="458">Figs 36</figureCitation>
,
<figureCitation id="9EB72A51FFA068239CD2FF284791FF60" box="[657,689,224,248]" captionStart="FIGURE 78" captionStartId="15.[151,250,1478,1500]" captionTargetBox="[190,1386,217,1416]" captionTargetId="figure@15.[170,1417,193,1457]" captionTargetPageId="15" captionText="FIGURE 78. Anafroptilum odontostylus sp. n., labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827912/files/figure.png" pageId="5" pageNumber="458">78</figureCitation>
). The same in many other taxa.
<emphasis id="34F8EAC6FFA068239A5BFF174166FF60" bold="true" box="[1048,1094,223,248]" pageId="5" pageNumber="458">(14)</emphasis>
Larval femora without row of stout setae on outer margin and without stout outer-apical setae; outer-apical margin serrate, with sparse thin hairs (
<figureCitation id="9EB72A51FFA068239EDCFEEF45D5FED8" box="[159,245,295,320]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="5" pageNumber="458">Fig. 59</figureCitation>
). The same in
<taxonomicName id="C18C4D57FFA068239FE3FEE04715FED8" box="[416,565,296,320]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA068239FE3FEE04715FED8" box="[416,565,296,320]" italics="true" pageId="5" pageNumber="458">Centroptilum</emphasis>
</taxonomicName>
and selected species of
<taxonomicName id="C18C4D57FFA068239D0FFEE046E0FED8" box="[844,960,296,320]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA068239D0FFEE046E0FED8" box="[844,960,296,320]" italics="true" pageId="5" pageNumber="458">Procloeon</emphasis>
</taxonomicName>
.
<emphasis id="34F8EAC6FFA068239D93FEEF46DFFED8" bold="true" box="[976,1023,295,320]" pageId="5" pageNumber="458">(15)</emphasis>
Male imaginal gonostylus with 2nd segment thickened apically, 3rd segment petiolate (
<figureCitation id="9EB72A51FFA068239CADFE844669FEFC" box="[750,841,332,357]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="5" pageNumber="458">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFA068239D19FE84465AFEFC" box="[858,890,332,356]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="5" pageNumber="458">79</figureCitation>
). The same in
<taxonomicName id="C18C4D57FFA068239A6DFE8441E2FEFC" box="[1070,1218,332,356]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA068239A6DFE8441E2FEFC" box="[1070,1218,332,356]" italics="true" pageId="5" pageNumber="458">Centroptilum</emphasis>
</taxonomicName>
and
<taxonomicName id="C18C4D57FFA068239B41FE8440BCFEFC" box="[1282,1436,332,356]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA068239B41FE8440BCFEFC" box="[1282,1436,332,356]" italics="true" pageId="5" pageNumber="458">Cheleocloeon</emphasis>
</taxonomicName>
(
<bibRefCitation id="621D4B25FFA068239EDDFEA74766FE10" author="Kluge" box="[158,582,367,392]" pageId="5" pageNumber="458" refString="Kluge, N. J. (2016 a) Redescription of the genus Cheleocloeon Wuillot &amp; Gillies 1993 (Ephemeroptera: Baetidae) with descriptions of three new species from Zambia and Uganda. Zootaxa, 4067 (2), 135 - 167. http: // dX. doi. org / 10.11646 / ZootaXa. 4067.2.2" type="journal article" year="2016" yearSuffix="a">Kluge 2016a: Figs 98, 106, 108, 110</bibRefCitation>
). In Cloeonini, second segment of gonostylus is also widened apically, and third segment is also petiolate, but smaller. In other respects external genitals of
<taxonomicName id="C18C4D57FFA068239A65FE5C419AFE34" box="[1062,1210,404,428]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA068239A65FE5C419AFE34" box="[1062,1210,404,428]" italics="true" pageId="5" pageNumber="458">Anafroptilum</emphasis>
</taxonomicName>
are highly variable: 1st segment of gonostylus either with characteristic projection on inner side (
<figureCitation id="9EB72A51FFA068239A42FE7F4176FE48" box="[1025,1110,439,464]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="5" pageNumber="458">Fig. 79</figureCitation>
), or without it (
<figureCitation id="9EB72A51FFA068239B4BFE7F407CFE48" box="[1288,1372,439,464]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="5" pageNumber="458">Fig. 41</figureCitation>
); 2nd segment either arched (
<figureCitation id="9EB72A51FFA068239FDDFE1444D2FE6C" box="[414,498,476,501]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="5" pageNumber="458">Fig. 79</figureCitation>
), or straight (
<figureCitation id="9EB72A51FFA068239CC9FE1447FEFE6C" box="[650,734,476,501]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="5" pageNumber="458">Fig. 41</figureCitation>
); median finger-like projection either present (
<figureCitation id="9EB72A51FFA068239AAFFE144064FE6C" box="[1260,1348,476,501]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="5" pageNumber="458">Figs 41</figureCitation>
,
<figureCitation id="9EB72A51FFA068239B12FE144050FE6C" box="[1361,1392,476,500]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="5" pageNumber="458">79</figureCitation>
), or absent.
</paragraph>
<caption id="52F3665CFFA068239ED4F85546CBF84B" httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="5" pageNumber="458" targetBox="[220,1301,599,1910]" targetPageId="5">
<paragraph id="063336D4FFA068239ED4F85546CBF84B" blockId="5.[151,1436,1949,2003]" pageId="5" pageNumber="458">
<emphasis id="34F8EAC6FFA068239ED4F8554475F82B" bold="true" box="[151,341,1949,1972]" pageId="5" pageNumber="458">FIGURES 28–36.</emphasis>
<taxonomicName id="C18C4D57FFA068239F1FF856477BF82C" box="[348,603,1950,1972]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA068239F1FF856477BF82C" box="[348,603,1950,1972]" italics="true" pageId="5" pageNumber="458">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA068239C20F85747BDF82C" box="[611,669,1951,1972]" pageId="5" pageNumber="458" rank="species">
<emphasis id="34F8EAC6FFA068239C20F85747BDF82C" bold="true" box="[611,669,1951,1972]" pageId="5" pageNumber="458">sp. n.</emphasis>
</taxonomicNameLabel>
, larvae. 28–34, tergalii I–VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa.
</paragraph>
</caption>
<caption id="52F3665CFFA368209ED4F8CA41CEF868" httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="6" pageNumber="459" targetBox="[201,1383,200,1772]" targetPageId="6">
<paragraph id="063336D4FFA368209ED4F8CA41CEF868" blockId="6.[151,1436,1794,2032]" pageId="6" pageNumber="459">
<emphasis id="34F8EAC6FFA368209ED4F8CA4474F880" bold="true" box="[151,340,1794,1817]" pageId="6" pageNumber="459">FIGURES 37–43.</emphasis>
<taxonomicName id="C18C4D57FFA368209F1AF8CB4776F881" box="[345,598,1795,1817]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="459" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA368209F1AF8CB4776F881" box="[345,598,1795,1817]" italics="true" pageId="6" pageNumber="459">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA368209C1FF8CB47B4F880" box="[604,660,1795,1816]" pageId="6" pageNumber="459" rank="species">
<emphasis id="34F8EAC6FFA368209C1FF8CB47B4F880" bold="true" box="[604,660,1795,1816]" pageId="6" pageNumber="459">sp. n.</emphasis>
</taxonomicNameLabel>
37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38–39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs1, gs2, gs3, segments of gonostylus; in, incisor; kd, kinetodontoum; m.IX-X, areas of anterior attachment of median paraproctal muscles (i.e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger.
</paragraph>
</caption>
<caption id="52F3665CFFA268219ED4F84F47BAF843" httpUri="https://zenodo.org/record/827900/files/figure.png" pageId="7" pageNumber="460" targetBox="[151,1436,193,1906]" targetPageId="7">
<paragraph id="063336D4FFA268219ED4F84F47BAF843" blockId="7.[151,1436,1927,2011]" pageId="7" pageNumber="460">
<emphasis id="34F8EAC6FFA268219ED4F84F4474F805" bold="true" box="[151,340,1927,1950]" pageId="7" pageNumber="460">FIGURES 44–51.</emphasis>
<taxonomicName id="C18C4D57FFA268219F1AF8404776F806" box="[345,598,1928,1950]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="orthostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA268219F1AF8404776F806" box="[345,598,1928,1950]" italics="true" pageId="7" pageNumber="460">Anafroptilum orthostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA268219C1FF84047B4F805" box="[604,660,1928,1949]" pageId="7" pageNumber="460" rank="species">
<emphasis id="34F8EAC6FFA268219C1FF84047B4F805" bold="true" box="[604,660,1928,1949]" pageId="7" pageNumber="460">sp. n.</emphasis>
</taxonomicNameLabel>
44, male imago; 45, eXuviae of subimaginal pleuron; 46, left half of eXuviae of subimaginal mesonotum; 47–48, female imago; 49, first tarsomere of middle leg of subimago; 50, genitals of male imago; 51, penis of male imago (45–46, 50–51, holotype).
</paragraph>
</caption>
<paragraph id="063336D4FFAD682E9E84FF5F4127FED8" blockId="8.[151,1436,151,357]" pageId="8" pageNumber="461">
<emphasis id="34F8EAC6FFAD682E9E84FF5F479CFF28" bold="true" box="[199,700,151,176]" pageId="8" pageNumber="461">
Variable characters of
<taxonomicName id="C18C4D57FFAD682E9F9CFF5F475CFF28" box="[479,636,151,176]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAD682E9F9CFF5F475CFF28" bold="true" box="[479,636,151,176]" italics="true" pageId="8" pageNumber="461">Anafroptilum</emphasis>
</taxonomicName>
. (16)
</emphasis>
Hind wings either present, or absent (in
<taxonomicName id="C18C4D57FFAD682E9ADAFF5041DEFF37" box="[1177,1278,152,175]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="34F8EAC6FFAD682E9ADAFF5041DEFF37" box="[1177,1278,152,175]" italics="true" pageId="8" pageNumber="461">A. minor</emphasis>
</taxonomicName>
and two new species described below). If present, hind wing is either of the «
<taxonomicName id="C18C4D57FFAD682E9DDAFF7546FFFF4C" box="[921,991,189,212]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAD682E9DDAFF7546FFFF4C" box="[921,991,189,212]" italics="true" pageId="8" pageNumber="461">Baetis</emphasis>
</taxonomicName>
-type», i.e. relatively wide, with three longitudinal veins and short costal projection (e.g., in
<taxonomicName id="C18C4D57FFAD682E9D45FF2846BBFF60" box="[774,923,224,248]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="species" species="bifircatum">
<emphasis id="34F8EAC6FFAD682E9D45FF2846BBFF60" box="[774,923,224,248]" italics="true" pageId="8" pageNumber="461">A. bifircatum</emphasis>
</taxonomicName>
), or of the «
<taxonomicName id="C18C4D57FFAD682E9A73FF2841E4FF60" box="[1072,1220,224,248]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAD682E9A73FF2841E4FF60" box="[1072,1220,224,248]" italics="true" pageId="8" pageNumber="461">Centroptilum</emphasis>
</taxonomicName>
-type», i.e. narrow, with two longitudinal veins and hooked costal projection (e.g., in
<taxonomicName id="C18C4D57FFAD682E9DA6FECC41B0FE84" box="[997,1168,260,284]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="461" phylum="Arthropoda" rank="species" species="kazlauskasi">
<emphasis id="34F8EAC6FFAD682E9DA6FECC41B0FE84" box="[997,1168,260,284]" italics="true" pageId="8" pageNumber="461">A. kazlauskasi</emphasis>
</taxonomicName>
).
<emphasis id="34F8EAC6FFAD682E9AF0FECC41C2FE85" bold="true" box="[1203,1250,260,285]" pageId="8" pageNumber="461">(17)</emphasis>
Tergalii either asymmetrically widened (
<figureCitation id="9EB72A51FFAD682E9FFBFEEF471BFED8" box="[440,571,295,320]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="8" pageNumber="461">Figs 69–75</figureCitation>
), or narrow and symmetric (
<figureCitation id="9EB72A51FFAD682E9D34FEEF46DBFED8" box="[887,1019,295,320]" captionStart="FIGURES 28 – 36" captionStartId="5.[151,264,1949,1972]" captionTargetBox="[220,1301,599,1910]" captionTargetId="figure@5.[216,1351,589,1927]" captionTargetPageId="5" captionText="FIGURES 28 – 36. Anafroptilum orthostylus sp. n., larvae. 28 – 34, tergalii I – VII; 35, labrum and clypeus; 36, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827896/files/figure.png" pageId="8" pageNumber="461">Figs 28–34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAD682E9E84FE844181FEFD" box="[199,1185,332,357]" pageId="8" pageNumber="461" type="distribution">
<paragraph id="063336D4FFAD682E9E84FE844181FEFD" blockId="8.[151,1436,151,357]" box="[199,1185,332,357]" pageId="8" pageNumber="461">
<emphasis id="34F8EAC6FFAD682E9E84FE844440FEFD" bold="true" box="[199,352,332,357]" pageId="8" pageNumber="461">Distribution.</emphasis>
Amphipacific: Asia (East Palaearctic and Oriental Regions) and Nearctic.
</paragraph>
</subSubSection>
</treatment>
</document>
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