120 lines
12 KiB
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120 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.81737" ID-GBIF-Dataset="d148b8fa-4f4c-4069-80e5-f04f98561bfe" ID-Pensoft-Pub="2747-8106-96-51" ID-Pensoft-UUID="FCEF2A912BC258BBB022681E86486B06" ID-ZooBank="CD8C70BA025345408EFCD430A12F6B35" LSID="urn:lsid:zoobank.org:act:6CF0B3D2-345A-4225-BC1E-769F8444169F" ModsDocID="2747-8106-96-51" checkinTime="1649795137967" checkinUser="pensoft" docAuthor="Schwarzhans, Werner & Keupp, Helmut" docDate="2022" docId="D082390C309159C1AABF15137BEE6AF4" docLanguage="en" docName="Zitteliana 96: 51-67" docOrigin="Zitteliana 96" docPubDate="2022-04-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.81737" docTitle="Archaeotolithus doppelsteini Schwarzhans & Keupp 2022, sp. nov." docType="treatment" docUuid="6CF0B3D2-345A-4225-BC1E-769F8444169F" docUuidSource="ZooBank" docVersion="2" id="FCEF2A912BC258BBB022681E86486B06" lastPageNumber="51" masterDocId="FCEF2A912BC258BBB022681E86486B06" masterDocTitle="Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany" masterLastPageNumber="67" masterPageNumber="51" pageId="0" pageNumber="51" updateTime="1649795760618" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schwarzhans, Werner</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4842-7989</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark & Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">wwschwarz@aol.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Keupp, Helmut</mods:namePart>
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<mods:affiliation>Freie Universitaet Berlin, Institut fuer Geologische Wissenschaften, Fachrichtung Palaeontologie, Malteser Strasse 74 - 100, Haus D, 12249 Berlin, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zitteliana</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>67</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zitteliana.96.81737</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.81737</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2747-8106-96-51</mods:identifier>
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<mods:identifier type="ZooBank">CD8C70BA025345408EFCD430A12F6B35</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FCEF2A912BC258BBB022681E86486B06</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:6CF0B3D2-345A-4225-BC1E-769F8444169F" httpUri="http://treatment.plazi.org/id/D082390C309159C1AABF15137BEE6AF4" lastPageNumber="51" pageId="0" pageNumber="51">
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<subSubSection pageId="0" pageNumber="51" type="nomenclature">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName LSID="http://zoobank.org/6CF0B3D2-345A-4225-BC1E-769F8444169F" authority="Schwarzhans & Keupp, 2022" authorityName="Schwarzhans & Keupp" authorityYear="2022" genus="Archaeotolithus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Archaeotolithus doppelsteini" order="Ophidiiformes" pageId="0" pageNumber="51" rank="species" species="doppelsteini" status="sp. nov.">Archaeotolithus doppelsteini</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="description">
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<paragraph pageId="0" pageNumber="51">
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<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. Otoliths of Archaeotolithus. fig. 1: Achaeotolithus bornholmiensis (Malling & Gronwall, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon. figs 2 - 4: Archaeotolithus doppelsteini sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon; 2: holotype SNSB-BSPG 2022 IV 8; 3 - 4: paratypes SNSB-BSPG 2022 IV 9. figs 5 - 7: Archaeotolithus trigonalis Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12." figureDoi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51">Plate 4, figs 2-4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="reference_group">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName authority="Schwarzhans & Keupp, 2022" authorityName="Schwarzhans & Keupp" authorityYear="2022" genus="Lapillus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lapillus unbekannter" pageId="0" pageNumber="51" rank="species" species="unbekannter" status="sp. nov.">Lapillus unbekannter</taxonomicName>
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2021b
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<taxonomicName authority="Schwarzhans & Keupp, 2022" authorityName="Schwarzhans & Keupp" authorityYear="2022" genus="Lapillus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lapillus unbekannter" pageId="0" pageNumber="51" rank="species" species="unbekannter" status="sp. nov.">Lapillus unbekannter</taxonomicName>
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Zuordnung - Keupp: fig. 9.3 D.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="etymology">
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<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="51">Named after Bernd Doppelstein (Berlin), who has intensely collected from the Buttenheim clay pit and provided the largest specimen of this species.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="holotype">
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<paragraph pageId="0" pageNumber="51">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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SNSB-BSPG 2022 IV 8 (Plate
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<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. Otoliths of Archaeotolithus. fig. 1: Achaeotolithus bornholmiensis (Malling & Gronwall, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon. figs 2 - 4: Archaeotolithus doppelsteini sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon; 2: holotype SNSB-BSPG 2022 IV 8; 3 - 4: paratypes SNSB-BSPG 2022 IV 9. figs 5 - 7: Archaeotolithus trigonalis Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12." figureDoi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51">4</figureCitation>
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, fig. 2), Clay pit near Buttenheim, upper Pliensbachian,
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<taxonomicName genus="Hawkerense" lsidName="Hawkerense" pageId="0" pageNumber="51" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="51">Hawkerense</emphasis>
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</taxonomicName>
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subzone, EPH horizon.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="paratypes">
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<paragraph pageId="0" pageNumber="51">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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4 specimens same data as holotype (SNSB-BSPG 2022 IV 9, Plate
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<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. Otoliths of Archaeotolithus. fig. 1: Achaeotolithus bornholmiensis (Malling & Gronwall, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon. figs 2 - 4: Archaeotolithus doppelsteini sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon; 2: holotype SNSB-BSPG 2022 IV 8; 3 - 4: paratypes SNSB-BSPG 2022 IV 9. figs 5 - 7: Archaeotolithus trigonalis Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12." figureDoi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51">4</figureCitation>
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, figs 3-4).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="diagnosis">
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<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="51">OL:OH = 0.9-0.93; OH:OT = 2.8-3.6. Ventral rim deeply curved, middorsal angle with small process. Outer face with complex pattern of subvertical furrows.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="description">
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<paragraph pageId="0" pageNumber="51">Description.</paragraph>
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<paragraph pageId="0" pageNumber="51">Very high-bodied otoliths up to 3.25 mm in length (holotype 2.2 mm) with rounded ventral rim and rounded postventral angle. Middorsal angle with small projection. Anterior and posterior rims smooth, ventral rim regularly and coarsely crenulated.</paragraph>
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<paragraph pageId="0" pageNumber="51">Inner face moderately convex with distinct supramedian positioned sulcus. Dorsal margin of sulcus relatively well-defined, ventral margin gradational and indistinct. Sulcus anteriorly open and ventrally widened, posteriorly curved slightly downward and terminating close to rounded postventral angle. Distinction in ostium and cauda indistinct or with broad shallow ventral bend at junction; OsL:CaL notably variable, ranging from 0.7 to 1.2. Outer face flat, with set of opposing subvertical furrows on pre- and postventral fields, diminishing in intensity with size.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="discussion">
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<paragraph pageId="0" pageNumber="51">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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The relatively small holotype is well preserved whereas the large figured paratype (Plate
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<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. Otoliths of Archaeotolithus. fig. 1: Achaeotolithus bornholmiensis (Malling & Gronwall, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon. figs 2 - 4: Archaeotolithus doppelsteini sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon; 2: holotype SNSB-BSPG 2022 IV 8; 3 - 4: paratypes SNSB-BSPG 2022 IV 9. figs 5 - 7: Archaeotolithus trigonalis Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12." figureDoi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51">4</figureCitation>
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, fig. 4) has severe incrustations across much of the sulcus that obliterate part of its morphology.
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<taxonomicName authorityName="Schwarzhans & Keupp" authorityYear="2022" genus="Archaeotolithus" kingdom="Animalia" lsidName="Archaeotolithus doppelsteini" order="Ophidiiformes" pageId="0" pageNumber="51" rank="species" species="doppelsteini">
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<emphasis italics="true" pageId="0" pageNumber="51">Archaeotolithus doppelsteini</emphasis>
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</taxonomicName>
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is readily distinguished from its congeners by its deep ventral rim, rounded postventral angle, and process of the middorsal angle. It differs from
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<taxonomicName genus="A." lsidName="A. trigonalis" pageId="0" pageNumber="51" rank="species" species="trigonalis">
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<emphasis italics="true" pageId="0" pageNumber="51">A. trigonalis</emphasis>
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</taxonomicName>
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in its higher body shape, expressed in the lower OL:OH ratio (0.9-0.93 vs. 1.05-1.12) and the different proportions of the ratio OsL:CaL (0.7-1.2 vs. 0.3-0.4). Furthermore, the ornamentation of the outer face differs from that observed in its congeners.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |