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<document id="AF1FC16C269BF4235526E9AF1DB376FF" ID-DOI="10.5281/zenodo.197810" ID-GBIF-Dataset="3fed616e-ea51-496b-b522-145b87775b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="197810" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460279155734" checkinUser="plazi" docAuthor="Rahmadi, Cahyo, Harvey, Mark S. &amp; Kojima, Jun-Ichi" docDate="2010" docId="AD1687F3FF8EFF960CEB4F702F74FB96" docLanguage="en" docName="zt02612p021.pdf" docOrigin="Zootaxa 2612" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sarax sarawakensis Thorell 1888" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="512FFF8BFF87FF9C0C7C4D7D2E59FFA4" masterDocTitle="Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island" masterLastPageNumber="21" masterPageNumber="1" pageNumber="10" updateTime="1698249151004" updateUser="plazi">
<mods:mods id="F4DAEEABF2BF070795FCF3076DBD34AD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9B85ADA7CB0CDC6EBF89657DD0835FD1">Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island</mods:title>
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<mods:name id="4C9715E3A862B0A9D4A30AE532170834" type="personal">
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<mods:namePart id="FB7D42A105E32621F8DAEB8E48FD4122">Rahmadi, Cahyo</mods:namePart>
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<mods:namePart id="0208CB1093643413BA348A88935CB2E8">Harvey, Mark S.</mods:namePart>
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<mods:namePart id="88F208CF96558D9731D0FA1817B40555">Kojima, Jun-Ichi</mods:namePart>
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<mods:title id="DA0AA2FDC1870F7C928565A7E725D64A">Zootaxa</mods:title>
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<mods:part id="9F0490D4D86D1AD47877A3C5C15DB89D">
<mods:date id="489EFA7F3196332F0162EF5E07E54D26">2010</mods:date>
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<mods:number id="723C94156181C56FAE86698B7622DED4">2612</mods:number>
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<mods:identifier id="3489BC15CA160EC44985A1E805C08171" type="DOI">10.5281/zenodo.197810</mods:identifier>
<mods:identifier id="A6D13D62C2DD9A26D3E6E76DF3D7CDF4" type="GBIF-Dataset">3fed616e-ea51-496b-b522-145b87775b5a</mods:identifier>
<mods:identifier id="3CA9083DB757F96EFBA97D169EDD4BDC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="90CC3115B76BEDE3E153D67CEDF4B2C6" type="Zenodo-Dep">197810</mods:identifier>
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<treatment id="AD1687F3FF8EFF960CEB4F702F74FB96" ID-DOI="http://doi.org/10.5281/zenodo.6207613" ID-GBIF-Taxon="119396508" ID-Zenodo-Dep="6207613" LSID="urn:lsid:plazi:treatment:AD1687F3FF8EFF960CEB4F702F74FB96" httpUri="http://treatment.plazi.org/id/AD1687F3FF8EFF960CEB4F702F74FB96" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="6DA5656EFF8EFF950CEB4F702F6DFDEE" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="250036E5FF8EFF950CEB4F702C66FD83" blockId="9.[151,575,525,586]" box="[151,575,525,551]" pageId="9" pageNumber="10">
<heading id="7E488189FF8EFF950CEB4F702C66FD83" bold="true" box="[151,575,525,551]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="E2BF4D66FF8EFF950CEB4F702C66FD83" ID-CoL="79MF7" authority="Thorell 1888" authorityName="Thorell" authorityYear="1888" box="[151,575,525,551]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8EFF950CEB4F702C66FD83" bold="true" box="[151,575,525,551]" pageId="9" pageNumber="10">
<emphasis id="17CBEAF7FF8EFF950CEB4F702FDEFD83" bold="true" box="[151,391,525,551]" italics="true" pageId="9" pageNumber="10">Sarax sarawakensis</emphasis>
(
<bibRefCitation id="412E4B14FF8EFF950DEA4F702C6EFD83" author="Thorell" box="[406,567,525,551]" pageId="9" pageNumber="10" refString="Thorell, T. (1888) Pedipalpi e Scorpioni dell'Arcipelago Malese conservati nel Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova, 26, 327 - 428." type="journal article" year="1888">Thorell 1888</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="250036E5FF8EFF950CEB4F4D2F6DFDEE" blockId="9.[151,575,525,586]" box="[151,308,560,586]" pageId="9" pageNumber="10">
(
<figureCitation id="BD842A60FF8EFF950CE34F4D2F74FDEE" box="[159,301,560,586]" captionStart="FIGURES 16 20" captionStartId="9.[151,269,1902,1926]" captionTargetBox="[174,1406,1066,1809]" captionTargetId="figure@9.[151,1436,1029,1881]" captionTargetPageId="9" captionText="FIGURES 16 20. Sarax sarawakensis (Thorell 1888), female holotype from Borneo unless stated otherwise: 16. Frontal carapace and right pedipalp; 17. External view of right chelicerae; 18. Pedipalpal tarsus, internal view; 19. Tibial and tarsal segments of leg I showing three most distal tibial segments, and five most proximal tarsal segments; 20. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV, (MZB. Ambl. 149)." httpUri="https://zenodo.org/record/197814/files/figure.png" pageId="9" pageNumber="10">Figs. 1620</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8EFF950CEB4F072AC8FD15" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="250036E5FF8EFF950CEB4F072C15FD35" blockId="9.[151,1169,634,689]" box="[151,588,634,657]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="05AF11CBFF8EFF950CEB4F072C15FD35" box="[151,588,634,657]" pageId="9" pageNumber="10">
<taxonomicName id="E2BF4D66FF8EFF950CEB4F072C11FD35" authority="Thorell 1888: 354" authorityName="Thorell" authorityPageNumber="354" authorityYear="1888" box="[151,584,634,657]" class="Arachnida" family="Charontidae" genus="Charon" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8EFF950CEB4F072F27FD35" box="[151,382,634,657]" italics="true" pageId="9" pageNumber="10">Charon sarawakensis</emphasis>
<treatmentCitation id="A41E10F4FF8EFF950DF84F072C11FD35" author="Thorell" box="[388,584,634,657]" page="354" pageId="9" pageNumber="10" year="1888">
<bibRefCitation id="412E4B14FF8EFF950DF84F072C4BFD35" author="Thorell" box="[388,530,634,657]" pageId="9" pageNumber="10" refString="Thorell, T. (1888) Pedipalpi e Scorpioni dell'Arcipelago Malese conservati nel Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova, 26, 327 - 428." type="journal article" year="1888">Thorell 1888</bibRefCitation>
: 354
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="250036E5FF8EFF950CEB4FE72AC8FD15" blockId="9.[151,1169,634,689]" box="[151,1169,666,689]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="05AF11CBFF8EFF950CEB4FE72AC8FD15" box="[151,1169,666,689]" pageId="9" pageNumber="10">
<taxonomicName id="E2BF4D66FF8EFF950CEB4FE72F8EFD15" ID-CoL="79MF7" authority="Thorell" authorityName="Thorell" box="[151,471,666,689]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8EFF950CEB4FE72F35FD15" box="[151,364,666,689]" italics="true" pageId="9" pageNumber="10">Sarax sarawakensis</emphasis>
(Thorell)
</taxonomicName>
:
<treatmentCitation id="A41E10F4FF8EFF950D9C4FE72CD7FD15" author="Simon" box="[480,654,666,689]" page="48" pageId="9" pageNumber="10" year="1892">
<bibRefCitation id="412E4B14FF8EFF950D9C4FE72C3FFD15" author="Simon" box="[480,614,666,689]" pageId="9" pageNumber="10" refString="Simon, E. (1892) Arachnides. In: A. Raffray, Bolivar, I. and Simon, E. (Eds.) Etude sur les Arthropodes cavernicoles de ile Luzon, Voyage de M. E. Simon aux iles Philippines. Mars et avril 1890. Annales de la Societe Entomologique de France, 61, 35 - 52." type="journal article" year="1892">Simon 1892</bibRefCitation>
: 48
</treatmentCitation>
;
<treatmentCitation id="A41E10F4FF8EFF950EE54FE72D19FD15" author="Harvey" box="[665,832,666,689]" page="8" pageId="9" pageNumber="10" year="2003">
<bibRefCitation id="412E4B14FF8EFF950EE54FE72D7EFD15" author="Harvey" box="[665,807,666,689]" pageId="9" pageNumber="10" refString="Harvey, M. S. (2003) Catalogue of the smaller arachnid orders of the world: Amblypygi, Uropygi, Schizomida, Palpigradi, Ricinulei and Solifugae. CSIRO Publishing, Melbourne." type="book" year="2003">Harvey 2003</bibRefCitation>
: 8
</treatmentCitation>
; Rahmadi and Kojima in press.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8EFF950CEB4FA22C56FC2B" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="250036E5FF8EFF950CEB4FA22C56FC2B" blockId="9.[151,1438,735,986]" pageId="9" pageNumber="10">
<emphasis id="17CBEAF7FF8EFF950CEB4FA22FD6FD5D" bold="true" box="[151,399,735,761]" pageId="9" pageNumber="10">Material examined:</emphasis>
Female
<typeStatus id="FA048847FF8EFF950D894F9D2C05FD5E" box="[501,604,736,762]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(
<collectionCode id="43AEAE20FF8EFF950E114F9D2C9FFD5E" box="[621,710,736,762]" httpUri="http://grbio.org/cool/q7tf-sdrm" name="Museo Civico di Storia Naturale, Grosseto" pageId="9" pageNumber="10">MCSG</collectionCode>
),
<collectingCountry id="5DA87675FF8EFF950EA74F9D2D21FD5E" box="[731,888,736,762]" name="Malaysia" pageId="9" pageNumber="10">MALAYSIA</collectingCountry>
:
<emphasis id="17CBEAF7FF8EFF950FF84F9D2DADFD5D" box="[900,1012,736,761]" italics="true" pageId="9" pageNumber="10">
<collectingCountry id="5DA87675FF8EFF950FF84F9D2DB6FD5D" box="[900,1007,736,761]" name="Malaysia" pageId="9" pageNumber="10">Sarawak</collectingCountry>
:
</emphasis>
(labeled “
<taxonomicName id="E2BF4D66FF8EFF95080C4F9D2BC5FD5E" authority="Thor." authorityName="Thor." box="[1136,1436,736,762]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="saravakensis">
<emphasis id="17CBEAF7FF8EFF95080C4F9D2B0CFD5D" box="[1136,1365,736,761]" italics="true" pageId="9" pageNumber="10">Sarax saravakensis</emphasis>
Thor.
</taxonomicName>
<collectingCountry id="5DA87675FF8EFF950CEB4E782F51FCBB" box="[151,264,773,799]" name="Malaysia" pageId="9" pageNumber="10">Sarawak</collectingCountry>
, Viag Doria and Beccari”).
<specimenCount id="33B9FD6CFF8EFF950E0E4E782CBBFCBB" box="[626,738,773,799]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
(
<collectionCode id="43AEAE20FF8EFF950E894E782D39FCBB" LSID="urn:lsid:biocol.org:col:34988" box="[757,864,773,799]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
.Am.17),
<collectingCountry id="5DA87675FF8EFF950FAE4E782ADBFCBB" box="[978,1154,773,799]" name="Indonesia" pageId="9" pageNumber="10">INDONESIA</collectingCountry>
:
<emphasis id="17CBEAF7FF8EFF9508F34E782A92FCBA" box="[1167,1227,773,798]" italics="true" pageId="9" pageNumber="10">Java</emphasis>
: labeled “
<taxonomicName id="E2BF4D66FF8EFF95092F4E782C43FCE0" authority="Thor., Kraepelin" authorityName="Thor., Kraepelin" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="saravakensis">
<emphasis id="17CBEAF7FF8EFF95092F4E782F65FCE7" italics="true" pageId="9" pageNumber="10">Sarax saravakensis</emphasis>
Thor., Kraepelin
</taxonomicName>
det. 1900, Java, Raffray 1005.77, 1877”.
<specimenCount id="33B9FD6CFF8EFF9508474E572AF2FCE0" box="[1083,1195,810,836]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
(
<collectionCode id="43AEAE20FF8EFF9508C14E572B5EFCE0" box="[1213,1287,810,836]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="10">MZB</collectionCode>
.Ambl.149),
<collectingCountry id="5DA87675FF8EFF950CEB4E2D2F62FCCE" box="[151,315,848,874]" name="Indonesia" pageId="9" pageNumber="10">INDONESIA</collectingCountry>
:
<emphasis id="17CBEAF7FF8EFF950D3A4E2D2FE6FCCD" box="[326,447,848,873]" italics="true" pageId="9" pageNumber="10">Lampung:</emphasis>
Kalianda, Raja Basa, GPS coordinates
<geoCoordinate id="408B5022FF8EFF950FEF4E2D2A42FCCE" box="[915,1051,848,874]" direction="south" orientation="latitude" pageId="9" pageNumber="10" precision="1" value="-5.7816">5°46.896´S</geoCoordinate>
,
<geoCoordinate id="408B5022FF8EFF9508544E2D2A97FCCE" box="[1064,1230,848,874]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="1" value="105.62812">105°37.687´E</geoCoordinate>
, altitude
<quantity id="E2479B00FF8EFF9509474E2D2BC1FCCE" box="[1339,1432,848,874]" metricMagnitude="3" metricUnit="m" metricValue="1.255" pageId="9" pageNumber="10" unit="m" value="1255.0">1255 m</quantity>
. asl.,
<date id="51011025FF8EFF950CB14E082FD2FC2B" box="[205,395,885,911]" pageId="9" pageNumber="10" value="2004-08-15">15 August 2004</date>
, A. Riedel.
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8EFF950CB94EE72DF7FC7E" pageId="9" pageNumber="10" type="distribution">
<paragraph id="250036E5FF8EFF950CB94EE72DF7FC7E" blockId="9.[151,1438,735,986]" pageId="9" pageNumber="10">
<emphasis id="17CBEAF7FF8EFF950CB94EE72F31FC10" bold="true" box="[197,360,922,948]" pageId="9" pageNumber="10">Distribution:</emphasis>
<taxonomicName id="E2BF4D66FF8EFF950D134EE72C00FC17" box="[367,601,922,947]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8EFF950D134EE72C00FC17" box="[367,601,922,947]" italics="true" pageId="9" pageNumber="10">Sarax sarawakensis</emphasis>
</taxonomicName>
is known from Andaman Island, Peninsular
<collectingCountry id="5DA87675FF8EFF9508184EE72A96FC10" box="[1124,1231,922,948]" name="Malaysia" pageId="9" pageNumber="10">Malaysia</collectingCountry>
(Selangor), Java, Sumatra [new record] (
<collectingCountry id="5DA87675FF8EFF950DDA4EBD2C47FC7E" box="[422,542,960,986]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
), and Borneo (
<collectingCountry id="5DA87675FF8EFF950EB54EBD2D68FC7E" box="[713,817,960,986]" name="Malaysia" pageId="9" pageNumber="10">Sarawak</collectingCountry>
) (
<figureCitation id="BD842A60FF8EFF950F354EBD2DF8FC7E" box="[841,929,960,986]" captionStart="FIGURES 41 42" captionStartId="16.[151,272,1727,1751]" captionTargetBox="[200,1385,221,1687]" captionTargetId="figure@16.[191,1395,194,1706]" captionTargetPageId="16" captionText="FIGURES 41 42. Map showing the distribution of Sarax species: 41. The distribution in Southeast Asia, except for Sarax species from India and New Guinea. 42. A detailed map showing the distribution in the Sangkulirang Karst, East Kalimantan." httpUri="https://zenodo.org/record/197818/files/figure.png" pageId="9" pageNumber="10">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="71C0666DFF8EFF950CEB4A132AB2F84B" httpUri="https://zenodo.org/record/197814/files/figure.png" pageId="9" pageNumber="10" targetBox="[174,1406,1066,1809]" targetPageId="9">
<paragraph id="250036E5FF8EFF950CEB4A132AB2F84B" blockId="9.[151,1437,1902,2031]" pageId="9" pageNumber="10">
<emphasis id="17CBEAF7FF8EFF950CEB4A132F04F822" bold="true" box="[151,349,1902,1926]" pageId="9" pageNumber="10">FIGURES 1620.</emphasis>
<taxonomicName id="E2BF4D66FF8EFF950D1F4A132C82F821" authority="Thorell 1888" authorityName="Thorell" authorityYear="1888" box="[355,731,1902,1925]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8EFF950D1F4A132C6FF821" box="[355,566,1902,1925]" italics="true" pageId="9" pageNumber="10">Sarax sarawakensis</emphasis>
(Thorell 1888)
</taxonomicName>
, female holotype from Borneo unless stated otherwise: 16. Frontal carapace and right pedipalp; 17. External view of right chelicerae; 18. Pedipalpal tarsus, internal view; 19. Tibial and tarsal segments of leg I showing three most distal tibial segments, and five most proximal tarsal segments; 20. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV, (MZB.Ambl.149).
</paragraph>
</caption>
<subSubSection id="6DA5656EFF8DFF960CB94DEA2F74FB96" pageId="10" pageNumber="11" type="discussion">
<paragraph id="250036E5FF8DFF960CB94DEA2DCAFE86" blockId="10.[151,1437,151,1074]" pageId="10" pageNumber="11">
<emphasis id="17CBEAF7FF8DFF960CB94DEA2F1DFF15" bold="true" box="[197,324,151,177]" pageId="10" pageNumber="11">Remarks:</emphasis>
<taxonomicName id="E2BF4D66FF8DFF960D314DE52C19FF15" box="[333,576,152,177]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8DFF960D314DE52C19FF15" box="[333,576,152,177]" italics="true" pageId="10" pageNumber="11">Sarax sarawakensis</emphasis>
</taxonomicName>
was first described by
<bibRefCitation id="412E4B14FF8DFF960F224DE52A48FF16" author="Thorell" box="[862,1041,152,178]" pageId="10" pageNumber="11" refString="Thorell, T. (1888) Pedipalpi e Scorpioni dell'Arcipelago Malese conservati nel Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova, 26, 327 - 428." type="journal article" year="1888">Thorell (1888)</bibRefCitation>
in the genus
<taxonomicName id="E2BF4D66FF8DFF9608CB4DE52B4BFF15" box="[1207,1298,152,177]" class="Arachnida" family="Charontidae" genus="Charon" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF9608CB4DE52B4BFF15" box="[1207,1298,152,177]" italics="true" pageId="10" pageNumber="11">Charon</emphasis>
</taxonomicName>
based on a single female from
<collectingCountry id="5DA87675FF8DFF960D004DC02FBCFF73" box="[380,485,189,215]" name="Malaysia" pageId="10" pageNumber="11">Sarawak</collectingCountry>
. The species was later transferred to
<taxonomicName id="E2BF4D66FF8DFF960FEA4DC02D83FF72" box="[918,986,189,214]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960FEA4DC02D83FF72" box="[918,986,189,214]" italics="true" pageId="10" pageNumber="11">Sarax</emphasis>
</taxonomicName>
by
<bibRefCitation id="412E4B14FF8DFF96087B4DC02AF1FF73" author="Simon" box="[1031,1192,189,215]" pageId="10" pageNumber="11" refString="Simon, E. (1892) Arachnides. In: A. Raffray, Bolivar, I. and Simon, E. (Eds.) Etude sur les Arthropodes cavernicoles de ile Luzon, Voyage de M. E. Simon aux iles Philippines. Mars et avril 1890. Annales de la Societe Entomologique de France, 61, 35 - 52." type="journal article" year="1892">Simon (1892)</bibRefCitation>
when the genus was established. The
<typeStatus id="FA048847FF8DFF960D1F4D9F2FCEFF58" box="[355,407,226,252]" pageId="10" pageNumber="11">type</typeStatus>
species,
<taxonomicName id="E2BF4D66FF8DFF960E7A4D9F2DF0FF58" authority="Simon 1892" authorityName="Simon" authorityYear="1892" box="[518,937,226,252]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachydactylus">
<emphasis id="17CBEAF7FF8DFF960E7A4D9F2D52FF5F" box="[518,779,226,251]" italics="true" pageId="10" pageNumber="11">Sarax brachydactylus</emphasis>
<bibRefCitation id="412E4B14FF8DFF960F684D9F2DF0FF58" author="Simon" box="[788,937,226,252]" pageId="10" pageNumber="11" refString="Simon, E. (1892) Arachnides. In: A. Raffray, Bolivar, I. and Simon, E. (Eds.) Etude sur les Arthropodes cavernicoles de ile Luzon, Voyage de M. E. Simon aux iles Philippines. Mars et avril 1890. Annales de la Societe Entomologique de France, 61, 35 - 52." type="journal article" year="1892">Simon 1892</bibRefCitation>
</taxonomicName>
, was found in the
<collectingCountry id="5DA87675FF8DFF9608F44D9F2B56FF58" box="[1160,1295,226,252]" name="Philippines" pageId="10" pageNumber="11">Philippines</collectingCountry>
from caves situated near Antipolo, San-Mateo and Calapnitan (
<bibRefCitation id="412E4B14FF8DFF960E8D4C752DDFFE86" author="Simon" box="[753,902,264,290]" pageId="10" pageNumber="11" refString="Simon, E. (1892) Arachnides. In: A. Raffray, Bolivar, I. and Simon, E. (Eds.) Etude sur les Arthropodes cavernicoles de ile Luzon, Voyage de M. E. Simon aux iles Philippines. Mars et avril 1890. Annales de la Societe Entomologique de France, 61, 35 - 52." type="journal article" year="1892">Simon 1892</bibRefCitation>
).
</paragraph>
<paragraph id="250036E5FF8DFF960CB94C502CFCFDE8" blockId="10.[151,1437,151,1074]" pageId="10" pageNumber="11">
<bibRefCitation id="412E4B14FF8DFF960CB94C502FF7FEE3" box="[197,430,301,327]" pageId="10" pageNumber="11" refString="Delle Cave, L. (1986) Biospeleology of the Somaliland Amblypygi (Arachnida, Chelicerata) of the caves of the Showli Berdi and Mugdile (Bardera, Somaliland). Redia, 69, 143 - 170." type="journal article">Delle Cave (1986)</bibRefCitation>
provided additional information on the
<typeStatus id="FA048847FF8DFF960FD64C502A4EFEE3" box="[938,1047,301,327]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E2BF4D66FF8DFF9608384C502B54FEE2" box="[1092,1293,301,326]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8DFF9608384C502B54FEE2" box="[1092,1293,301,326]" italics="true" pageId="10" pageNumber="11">S. sarawakensis</emphasis>
</taxonomicName>
, which she compared with a
<taxonomicName id="E2BF4D66FF8DFF960D084C2F2FE4FECF" box="[372,445,338,363]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960D084C2F2FE4FECF" box="[372,445,338,363]" italics="true" pageId="10" pageNumber="11">Sarax</emphasis>
</taxonomicName>
specimen from
<collectingCountry id="5DA87675FF8DFF960EF04C2F2D55FEC8" box="[652,780,338,364]" name="Singapore" pageId="10" pageNumber="11">Singapore</collectingCountry>
in the collection of the British Museum. She noted differences between the
<typeStatus id="FA048847FF8DFF960DCB4C052C47FE36" box="[439,542,376,402]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
and specimen from
<collectingCountry id="5DA87675FF8DFF960F6C4C052DD3FE36" box="[784,906,376,402]" name="Singapore" pageId="10" pageNumber="11">Singapore</collectingCountry>
, where the pedipalpal patella possessed only one spine between
<emphasis id="17CBEAF7FF8DFF960DF44CE02FF2FE13" bold="true" box="[392,427,413,439]" pageId="10" pageNumber="11">P1</emphasis>
and the distal margin instead of two.
<bibRefCitation id="412E4B14FF8DFF960FF04CE02A12FE13" author="Gravely" box="[908,1099,413,439]" pageId="10" pageNumber="11" refString="Gravely, F. H. (1911) Notes on Pedipalpi in the collection of the Indian Museum. II. A preliminary note on a new Sarax from Singapore. Records of the Indian Museum, 6, 36 - 38." type="journal article" year="1911">Gravely (1911)</bibRefCitation>
reported this species from
<collectingCountry id="5DA87675FF8DFF960CEB4CBF2F4BFE78" box="[151,274,450,476]" name="Singapore" pageId="10" pageNumber="11">Singapore</collectingCountry>
which he named
<taxonomicName id="E2BF4D66FF8DFF960D964CBF2D34FE7F" box="[490,877,450,475]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="sarawakensis" subSpecies="singaporae">
<emphasis id="17CBEAF7FF8DFF960D964CBF2D34FE7F" box="[490,877,450,475]" italics="true" pageId="10" pageNumber="11">Sarax sarawakensis singaporae</emphasis>
</taxonomicName>
which was later elevated to species level and moved to the genus
<taxonomicName id="E2BF4D66FF8DFF960DED4C952D45FDA6" authority="Gravely 1915" authorityName="Gravely" authorityYear="1915" box="[401,796,488,514]" class="Arachnida" family="Charinidae" genus="Phrynichosarax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960DED4C952C0DFDA5" box="[401,596,488,513]" italics="true" pageId="10" pageNumber="11">Phrynichosarax</emphasis>
(
<bibRefCitation id="412E4B14FF8DFF960E184C952D4BFDA6" author="Gravely" box="[612,786,488,514]" pageId="10" pageNumber="11" refString="Gravely, F. H. (1915) A revision of the Oriental subfamilies of Tarantulidae order Pedipalpi. Records of the Indian Museum, 11, 433 - 455." type="journal article" year="1915">Gravely 1915</bibRefCitation>
)
</taxonomicName>
, forming the combination
<taxonomicName id="E2BF4D66FF8DFF9608184C952B54FDA5" box="[1124,1293,488,513]" class="Arachnida" family="Charinidae" genus="Phrynichosarax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="singaporae">
<emphasis id="17CBEAF7FF8DFF9608184C952B54FDA5" box="[1124,1293,488,513]" italics="true" pageId="10" pageNumber="11">P. singaporae</emphasis>
</taxonomicName>
. The genus
<taxonomicName id="E2BF4D66FF8DFF960CEB4F702F06FD82" box="[151,351,525,550]" class="Arachnida" family="Charinidae" genus="Phrynichosarax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960CEB4F702F06FD82" box="[151,351,525,550]" italics="true" pageId="10" pageNumber="11">Phrynichosarax</emphasis>
</taxonomicName>
was based on the presence of three basitibial segments of leg IV (
<bibRefCitation id="412E4B14FF8DFF9608D74F702B07FD83" author="Gravely" box="[1195,1374,525,551]" pageId="10" pageNumber="11" refString="Gravely, F. H. (1915) A revision of the Oriental subfamilies of Tarantulidae order Pedipalpi. Records of the Indian Museum, 11, 433 - 455." type="journal article" year="1915">Gravely 1915</bibRefCitation>
), but
<bibRefCitation id="412E4B14FF8DFF960CEB4F4F2F04FDE8" author="Weygoldt" box="[151,349,562,588]" pageId="10" pageNumber="11" refString="Weygoldt, P. (2000) Whip spiders: Their biology, morphology and systematics. Apollo Books, Stenstrup." type="book" year="2000">Weygoldt (2000)</bibRefCitation>
synonymised it with
<taxonomicName id="E2BF4D66FF8DFF960E264F4F2CC7FDEF" box="[602,670,562,587]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960E264F4F2CC7FDEF" box="[602,670,562,587]" italics="true" pageId="10" pageNumber="11">Sarax</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="250036E5FF8DFF960CB94F252A65FC38" blockId="10.[151,1437,151,1074]" pageId="10" pageNumber="11">
A single female specimen of
<taxonomicName id="E2BF4D66FF8DFF960E414F252CDEFDD5" box="[573,647,600,625]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960E414F252CDEFDD5" box="[573,647,600,625]" italics="true" pageId="10" pageNumber="11">Sarax</emphasis>
</taxonomicName>
sp. from Bacangan, Palawan Island, was found in the FMNH collection. It has small setiferous tubercles and one spine between spine
<emphasis id="17CBEAF7FF8DFF9608614F002A67FD33" bold="true" box="[1053,1086,637,663]" pageId="10" pageNumber="11">P1</emphasis>
and the distal margin of the pedipalpal patella, and 17 trichobothria on tibia of leg IV which conforms to
<taxonomicName id="E2BF4D66FF8DFF9608544FDF2ABDFD1F" box="[1064,1252,674,699]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8DFF9608544FDF2ABDFD1F" box="[1064,1252,674,699]" italics="true" pageId="10" pageNumber="11">S. sarawakensis</emphasis>
</taxonomicName>
(
<figureCitation id="BD842A60FF8DFF9608894FDF2B14FD18" box="[1269,1357,674,700]" captionStart="FIGURES 16 20" captionStartId="9.[151,269,1902,1926]" captionTargetBox="[174,1406,1066,1809]" captionTargetId="figure@9.[151,1436,1029,1881]" captionTargetPageId="9" captionText="FIGURES 16 20. Sarax sarawakensis (Thorell 1888), female holotype from Borneo unless stated otherwise: 16. Frontal carapace and right pedipalp; 17. External view of right chelicerae; 18. Pedipalpal tarsus, internal view; 19. Tibial and tarsal segments of leg I showing three most distal tibial segments, and five most proximal tarsal segments; 20. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV, (MZB. Ambl. 149)." httpUri="https://zenodo.org/record/197814/files/figure.png" pageId="10" pageNumber="11">Fig. 20</figureCitation>
).
<bibRefCitation id="412E4B14FF8DFF96091F4FDF2EBEFD46" author="Fage" pageId="10" pageNumber="11" refString="Fage, L. (1946) Scorpions et Pedipalpes de l'Indochine Francaise. Annales de la Societe Entomologique de France, 113, 71 - 81." type="journal article" year="1946">Fage (1946)</bibRefCitation>
compared the pedipalp spination of three species,
<taxonomicName id="E2BF4D66FF8DFF960F324FB52A3CFD46" authority="Fage, 1946" authorityName="Fage" authorityYear="1946" box="[846,1125,712,738]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="davidovi">
<emphasis id="17CBEAF7FF8DFF960F324FB52D8CFD45" box="[846,981,712,737]" italics="true" pageId="10" pageNumber="11">S. davidovi</emphasis>
<bibRefCitation id="412E4B14FF8DFF960FA14FB52A3CFD46" author="Fage" box="[989,1125,712,738]" pageId="10" pageNumber="11" refString="Fage, L. (1946) Scorpions et Pedipalpes de l'Indochine Francaise. Annales de la Societe Entomologique de France, 113, 71 - 81." type="journal article" year="1946">Fage, 1946</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E2BF4D66FF8DFF96080F4FB52B10FD45" box="[1139,1353,712,737]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachydactylus">
<emphasis id="17CBEAF7FF8DFF96080F4FB52B10FD45" box="[1139,1353,712,737]" italics="true" pageId="10" pageNumber="11">S. brachydactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="E2BF4D66FF8DFF9609F94FB52F6CFCA2" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8DFF9609F94FB52F6CFCA2" italics="true" pageId="10" pageNumber="11">S. sarawakensis</emphasis>
</taxonomicName>
. Among them,
<taxonomicName id="E2BF4D66FF8DFF960D924F902CE6FCA2" box="[494,703,749,774]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachydactylus">
<emphasis id="17CBEAF7FF8DFF960D924F902CE6FCA2" box="[494,703,749,774]" italics="true" pageId="10" pageNumber="11">S. brachydactylus</emphasis>
</taxonomicName>
can be distinguished from other congeners by the presence of two spines between
<emphasis id="17CBEAF7FF8DFF960DFE4E6F2FFBFC88" bold="true" box="[386,418,786,812]" pageId="10" pageNumber="11">P1</emphasis>
and the distal margin, five major spines on the pedipalpal patella and four major spines on the pedipalpal femur (see
<bibRefCitation id="412E4B14FF8DFF960D8D4E452C29FCF6" author="Fage" box="[497,624,824,850]" pageId="10" pageNumber="11" refString="Fage, L. (1946) Scorpions et Pedipalpes de l'Indochine Francaise. Annales de la Societe Entomologique de France, 113, 71 - 81." type="journal article" year="1946">Fage 1946</bibRefCitation>
, fig. 3).
<taxonomicName id="E2BF4D66FF8DFF960EA94E452D98FCF5" box="[725,961,824,849]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8DFF960EA94E452D98FCF5" box="[725,961,824,849]" italics="true" pageId="10" pageNumber="11">Sarax sarawakensis</emphasis>
</taxonomicName>
has four major spines on the pedipalpal patella (
<figureCitation id="BD842A60FF8DFF960C864E202F0DFCD3" box="[250,340,861,887]" captionStart="FIGURES 16 20" captionStartId="9.[151,269,1902,1926]" captionTargetBox="[174,1406,1066,1809]" captionTargetId="figure@9.[151,1436,1029,1881]" captionTargetPageId="9" captionText="FIGURES 16 20. Sarax sarawakensis (Thorell 1888), female holotype from Borneo unless stated otherwise: 16. Frontal carapace and right pedipalp; 17. External view of right chelicerae; 18. Pedipalpal tarsus, internal view; 19. Tibial and tarsal segments of leg I showing three most distal tibial segments, and five most proximal tarsal segments; 20. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV, (MZB. Ambl. 149)." httpUri="https://zenodo.org/record/197814/files/figure.png" pageId="10" pageNumber="11">Fig. 17</figureCitation>
) and three major spines on the pedipalpal femur with additional distal minor spines. The specimen from Raja Basa (southern Sumatra) shows similar pedipalp spination.
</paragraph>
<paragraph id="250036E5FF8DFF960CB94ED52F74FB96" blockId="10.[151,1437,151,1074]" pageId="10" pageNumber="11">
Although the number and arrangement of trichobothria on leg IV in the
<typeStatus id="FA048847FF8DFF9608384ED52A20FC66" box="[1092,1145,936,962]" pageId="10" pageNumber="11">type</typeStatus>
of
<taxonomicName id="E2BF4D66FF8DFF9608D84ED52B26FC65" box="[1188,1407,936,961]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachydactylus">
<emphasis id="17CBEAF7FF8DFF9608D84ED52B26FC65" box="[1188,1407,936,961]" italics="true" pageId="10" pageNumber="11">S. brachydactylus</emphasis>
</taxonomicName>
is unknown, the specimen of
<taxonomicName id="E2BF4D66FF8DFF960DAA4EB02C43FC42" box="[470,538,973,998]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="17CBEAF7FF8DFF960DAA4EB02C43FC42" box="[470,538,973,998]" italics="true" pageId="10" pageNumber="11">Sarax</emphasis>
</taxonomicName>
from Palawan is clearly similar to
<taxonomicName id="E2BF4D66FF8DFF960FC74EB02AD5FC42" box="[955,1164,973,998]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachydactylus">
<emphasis id="17CBEAF7FF8DFF960FC74EB02AD5FC42" box="[955,1164,973,998]" italics="true" pageId="10" pageNumber="11">S. brachydactylus</emphasis>
</taxonomicName>
based on the spination of the pedipalpal patella and femur as specified in the identification key by
<bibRefCitation id="412E4B14FF8DFF96087A4E8F2A97FBA8" author="Kraepelin" box="[1030,1230,1010,1036]" pageId="10" pageNumber="11" refString="Kraepelin, K. (1899) Scorpiones und Pedipalpi. Tierreich, 8, i - xviii, 1 - 265." type="journal article" year="1899">Kraepelin (1899)</bibRefCitation>
and the figure by Fage (1949).
</paragraph>
</subSubSection>
</treatment>
</document>