478 lines
63 KiB
XML
478 lines
63 KiB
XML
<document id="5C8248F405B1DC863A98514C825ED833" ID-DOI="10.5281/zenodo.197810" ID-GBIF-Dataset="3fed616e-ea51-496b-b522-145b87775b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="197810" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460279155734" checkinUser="plazi" docAuthor="Rahmadi, Cahyo, Harvey, Mark S. & Kojima, Jun-Ichi" docDate="2010" docId="AD1687F3FF82FF950CEB48172A95FE13" docLanguage="en" docName="zt02612p021.pdf" docOrigin="Zootaxa 2612" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sarax yayukae Rahmadi, Harvey & Kojima, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="10" masterDocId="512FFF8BFF87FF9C0C7C4D7D2E59FFA4" masterDocTitle="Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island" masterLastPageNumber="21" masterPageNumber="1" pageNumber="6" updateTime="1698249151004" updateUser="plazi">
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<mods:mods id="3F9D09F8F9333D410E839F142DB9ADBF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6F1704BC1C89B458ADBC554548FA3264">
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<mods:title id="5B129F07C1CD9FF3833F49C3F255BE1A">Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island</mods:title>
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</mods:titleInfo>
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<mods:name id="1F79E04BEB6DCA97A73D4B62F2C2137F" type="personal">
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<mods:role id="8A422137DCEFCE137D530F1810DF1E85">
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<mods:roleTerm id="6CA38053DBF75AC072EB329D360A2DA4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D6C51D7C2D8B6D720CDFEBE4D6722CF0">Rahmadi, Cahyo</mods:namePart>
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</mods:name>
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<mods:name id="6ED4C52BDCDBE63768D68953562E1576" type="personal">
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<mods:role id="95873C8F5C4B5449AC2CB2EB9C97094E">
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<mods:roleTerm id="7372AFC1BCAC35D3973A70DB267E632B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5BC6DBBF09BB5C6899605828AEFB5666">Harvey, Mark S.</mods:namePart>
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</mods:name>
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<mods:name id="2A516DE65664A15B4F25E267F5E89AD0" type="personal">
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<mods:role id="63DEBB08266A97424949E1A2425105C0">
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<mods:roleTerm id="E947606F2E5B706658A835FEFE5AE029">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FEE49965D603E3E1603966B531733FB8">Kojima, Jun-Ichi</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4317140484BC75A3DB9E8469ACD7A736">text</mods:typeOfResource>
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<mods:relatedItem id="F025D872825327BCDDF8C6F275ABB8B9" type="host">
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<mods:titleInfo id="6EAF65C92262457F4A0AAD2D573DD25E">
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<mods:title id="CF64804D0348AB7569CC016D8F370496">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3BBF4CB96534A933D906098634C3EF3E">
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<mods:date id="CEFF3CB600E15C76EB9055A552FEB90D">2010</mods:date>
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<mods:detail id="919B00D150F12F3555461567AE298319" type="volume">
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<mods:number id="82DD2C7CF415F473DEE396DFD27D7069">2612</mods:number>
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||
</mods:detail>
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<mods:extent id="B57878F7A969FA0E6033977D5A9B97EF" unit="page">
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<mods:start id="85319C266BAA47D30D6C5B0C52E4FD4C">1</mods:start>
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<mods:end id="D4FA78AC5B382E92629111363E4CFB3A">21</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="161404E67A063BFC9D2BEE3888676097">journal article</mods:classification>
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<mods:identifier id="9218A67EB7E9A27FC1C37FCF1FC0DF46" type="DOI">10.5281/zenodo.197810</mods:identifier>
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<mods:identifier id="12F5E8CE70EB267D9FE733718AC5527F" type="GBIF-Dataset">3fed616e-ea51-496b-b522-145b87775b5a</mods:identifier>
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<mods:identifier id="8D52643C4A90AC7A24A0B76108CDD038" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E2D565465031962890ABDC3A38056FBF" type="Zenodo-Dep">197810</mods:identifier>
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</mods:mods>
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<treatment id="AD1687F3FF82FF950CEB48172A95FE13" ID-DOI="http://doi.org/10.5281/zenodo.6207611" ID-GBIF-Taxon="119396513" ID-Zenodo-Dep="6207611" LSID="urn:lsid:zoobank.org:act:3B12FE29-A43B-48EE-A649-EDB99C2D2F0B" httpUri="http://treatment.plazi.org/id/AD1687F3FF82FF950CEB48172A95FE13" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
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<subSubSection id="6DA5656EFF82FF990CEB48172F26FA03" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="250036E5FF82FF990CEB48172FF7FA20" blockId="5.[151,430,1386,1447]" box="[151,430,1386,1412]" pageId="5" pageNumber="6">
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<heading id="7E488189FF82FF990CEB48172FF7FA20" bold="true" box="[151,430,1386,1412]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis id="17CBEAF7FF82FF990CEB48172FF7FA20" bold="true" box="[151,430,1386,1412]" pageId="5" pageNumber="6">
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<taxonomicName id="E2BF4D66FF82FF990CEB48172F10FA20" ID-CoL="79MHC" box="[151,329,1386,1412]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yayukae" status="sp. nov.">
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<emphasis id="17CBEAF7FF82FF990CEB48172F10FA20" bold="true" box="[151,329,1386,1412]" italics="true" pageId="5" pageNumber="6">Sarax yayukae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0CF8578CFF82FF990D2C48172FF7FA20" box="[336,430,1386,1412]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="250036E5FF82FF990CEB48F02F26FA03" blockId="5.[151,430,1386,1447]" box="[151,383,1421,1447]" pageId="5" pageNumber="6">
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(
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<figureCitation id="BD842A60FF82FF990CE348F02F47FA03" box="[159,286,1421,1447]" captionStart-0="FIGURES 8 – 12" captionStart-1="FIGURES 13 – 15" captionStartId-0="6.[151,273,1910,1934]" captionStartId-1="7.[151,269,1094,1118]" captionTargetBox-0="[173,1405,906,1859]" captionTargetBox-1="[201,1350,232,1051]" captionTargetId-0="figure@6.[151,1436,844,1890]" captionTargetId-1="figure@7.[151,1436,194,1072]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." captionText-1="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri-0="https://zenodo.org/record/197812/files/figure.png" httpUri-1="https://zenodo.org/record/197813/files/figure.png" pageId="5" pageNumber="6">
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Figs. 8
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<emphasis id="17CBEAF7FF82FF990C8C48F02EA6FA03" bold="true" box="[240,255,1421,1447]" italics="true" pageId="5" pageNumber="6">–</emphasis>
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15
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</figureCitation>
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,
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<figureCitation id="BD842A60FF82FF990D5748F02F2FFA03" box="[299,374,1421,1447]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="5" pageNumber="6">
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35
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<emphasis id="17CBEAF7FF82FF990D3448F02F0FFA03" bold="true" box="[328,342,1421,1447]" italics="true" pageId="5" pageNumber="6">–</emphasis>
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36
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</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF82FF990CEB48A52D21F8EA" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="250036E5FF82FF990CEB48A52D21F8EA" blockId="5.[151,1437,1496,2019]" pageId="5" pageNumber="6">
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<emphasis id="17CBEAF7FF82FF990CEB48A52FCCFA56" bold="true" box="[151,405,1496,1522]" pageId="5" pageNumber="6">Material examined:</emphasis>
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Male
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<typeStatus id="FA048847FF82FF990D9848A52C17FA56" box="[484,590,1496,1522]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="43AEAE20FF82FF990E2348A52CFAFA56" box="[607,675,1496,1522]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="6">MZB</collectionCode>
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.Ambl.056),
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<collectingCountry id="5DA87675FF82FF990F4148A52DBCFA56" box="[829,997,1496,1522]" name="Indonesia" pageId="5" pageNumber="6">INDONESIA</collectingCountry>
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:
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<emphasis id="17CBEAF7FF82FF990F8D48A52ABAFA55" box="[1009,1251,1496,1521]" italics="true" pageId="5" pageNumber="6">Central Kalimantan</emphasis>
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: Murung Raya Regency, Tumbang Topus, Liang Puruk, GPS coordinates
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<geoCoordinate id="408B5022FF82FF990F164B7D2A5FF9BE" box="[874,1030,1533,1562]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="0.46249998">0o27’45.0”N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF82FF9908684B7D2A95F9BE" box="[1044,1228,1533,1562]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="115.01542">115o00’55.5”E</geoCoordinate>
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,
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<date id="51011025FF82FF9908A64B7D2B2AF9BE" box="[1242,1395,1536,1562]" pageId="5" pageNumber="6" value="2004-06-09">9 June 2004</date>
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, C. Rahmadi, Y.R. Suhardjono & D. Silam.
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<typeStatus id="FA048847FF82FF990E0C4B5B2CBEF9E4" box="[624,743,1574,1600]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="33B9FD6CFF82FF990E8E4B5B2D3AF9E4" box="[754,867,1574,1600]" pageId="5" pageNumber="6" type="female">2 females</specimenCount>
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and
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<specimenCount id="33B9FD6CFF82FF990FE04B5B2A48F9E4" box="[924,1041,1574,1600]" pageId="5" pageNumber="6" type="juvenile">1 juvenile</specimenCount>
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(
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<collectionCode id="43AEAE20FF82FF99085D4B5B2A3DF9E4" box="[1057,1124,1574,1600]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="6">MZB</collectionCode>
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.Ambl. 057-059),
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<specimenCount id="33B9FD6CFF82FF99094A4B5B2BC5F9E4" box="[1334,1436,1574,1600]" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
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(MNHN-Am.09), Liang Puruk, GPS coordinates
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<geoCoordinate id="408B5022FF82FF990EAB4B332D34F9CC" box="[727,877,1611,1640]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="0.46249998">0o27’45.0”N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF82FF990F054B332A7EF9CC" box="[889,1063,1611,1640]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="115.01542">115o00’55.5”E</geoCoordinate>
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,
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<date id="51011025FF82FF99084F4B332A9AF9CC" box="[1075,1219,1614,1640]" pageId="5" pageNumber="6" value="2004-06-09">9 June 2004</date>
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, C. Rahmadi, Y.R. Suhardjono & D. Silam;
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<specimenCount id="33B9FD6CFF82FF990D944B0B2C19F934" box="[488,576,1654,1680]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
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(
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<collectionCode id="43AEAE20FF82FF990E2B4B0B2CC3F934" box="[599,666,1654,1680]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="6">MZB</collectionCode>
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.Ambl.0 93), Liang Hajuq, GPS coordinates
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<geoCoordinate id="408B5022FF82FF9908914B0B2BC1F934" box="[1261,1432,1651,1680]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="0.46052778">0o27’37.9”N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF82FF990CEB4BE32F1FF91C" box="[151,326,1691,1720]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="115.01322">115o00’47.6”E</geoCoordinate>
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,
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<date id="51011025FF82FF990D2E4BE32FA8F91C" box="[338,497,1694,1720]" pageId="5" pageNumber="6" value="2004-06-11">11 June 2004</date>
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, C. Rahmadi;
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<emphasis id="17CBEAF7FF82FF990EE64BE22D35F91C" box="[666,876,1695,1720]" italics="true" pageId="5" pageNumber="6">West Kalimantan:</emphasis>
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<specimenCount id="33B9FD6CFF82FF990F084BE32D9DF91C" box="[884,964,1694,1720]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
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(
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<collectionCode id="43AEAE20FF82FF990FA84BE32A40F91C" box="[980,1049,1694,1720]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="6">MZB</collectionCode>
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.Ambl.148), Gua Kelasi in buffer zone, Bukit Raya-Bukit Baka National Park, Menukung, Melawi,
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<date id="51011025FF82FF990FE04BB92A13F97A" box="[924,1098,1732,1758]" pageId="5" pageNumber="6" value="2008-08-07">7 August 2008</date>
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, K.P.G. Himakova.
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<specimenCount id="33B9FD6CFF82FF99094C4BB92BDBF97A" box="[1328,1410,1732,1758]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
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,
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<specimenCount id="33B9FD6CFF82FF9909F14BB92EAFF8A7" pageId="5" pageNumber="6" type="female">3 females</specimenCount>
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(
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<collectionCode id="43AEAE20FF82FF990D7B4B942F1FF8A7" LSID="urn:lsid:biocol.org:col:34700" box="[263,326,1769,1795]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="5" pageNumber="6">CAS</collectionCode>
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,
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<collectionCode id="43AEAE20FF82FF990D2F4B942F91F8A7" box="[339,456,1769,1795]" pageId="5" pageNumber="6">CASENT</collectionCode>
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9036127),
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<collectingCountry id="5DA87675FF82FF990E2A4B942CAEF8A7" box="[598,759,1769,1795]" name="Malaysia" pageId="5" pageNumber="6">MALAYSIA</collectingCountry>
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:
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<emphasis id="17CBEAF7FF82FF990F7F4B942D00F8A6" box="[771,857,1769,1794]" italics="true" pageId="5" pageNumber="6">Sabah:</emphasis>
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Kota Kinabalu, Manukan Island, under logs in forest,
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<date id="51011025FF82FF990C9B4A732F2EF88C" box="[231,375,1806,1832]" pageId="5" pageNumber="6" value="2006-05-06">6 May 2006</date>
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, T. Briggs;
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<specimenCount id="33B9FD6CFF82FF990D804A732C29F88C" box="[508,624,1806,1832]" pageId="5" pageNumber="6" type="juvenile">1 juvenile</specimenCount>
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(
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<collectionCode id="43AEAE20FF82FF990E034A732CE5F88C" LSID="urn:lsid:biocol.org:col:34700" box="[639,700,1806,1832]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="5" pageNumber="6">CAS</collectionCode>
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,
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<collectionCode id="43AEAE20FF82FF990EB44A732D63F88C" box="[712,826,1806,1832]" pageId="5" pageNumber="6">CASENT</collectionCode>
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9036126), Kota Kinabalu, Manukan Island, jogging track on leaf litter, under logs,
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<date id="51011025FF82FF990D834A492CC9F8EA" box="[511,656,1844,1870]" pageId="5" pageNumber="6" value="2006-05-06">6 May 2006</date>
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, H. Tu & T. Briggs.
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF82FF9A0CB94A252FFCFF58" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
|
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<paragraph id="250036E5FF82FF9A0CB94A252FFCFF58" blockId="5.[151,1437,1496,2019]" lastBlockId="6.[151,1437,152,812]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<emphasis id="17CBEAF7FF82FF990CB94A252F12F8D6" bold="true" box="[197,331,1880,1906]" pageId="5" pageNumber="6">Diagnosis:</emphasis>
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<taxonomicName id="E2BF4D66FF82FF990D284A242C5FF8D6" box="[340,518,1881,1906]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yayukae">
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<emphasis id="17CBEAF7FF82FF990D284A242C5FF8D6" box="[340,518,1881,1906]" italics="true" pageId="5" pageNumber="6">Sarax yayukae</emphasis>
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</taxonomicName>
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has distinct sexual dimorphism, the male with strong and slender pedipalp whereas the female shorter and stouter. The species is medium-sized with an adult body length of about
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<quantity id="E2479B00FF82FF9909114A032F5AF81A" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.18" metricValueMin="0.88" pageId="5" pageNumber="6" unit="mm" value="10.3" valueMax="11.8" valueMin="8.8">8.8- 11.8 mm</quantity>
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. Pedipalpal femur with three major spines on antero-dorsal and antero-ventral margins; patella with four major spines on antero-dorsal margin; tarsus with three spines on antero-dorsal margin in adult specimens; the juveniles of
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<taxonomicName id="E2BF4D66FF81FF9A0D864DE52CD9FF15" box="[506,640,152,177]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="yayukae">
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<emphasis id="17CBEAF7FF81FF9A0D864DE52CD9FF15" box="[506,640,152,177]" italics="true" pageId="6" pageNumber="7">S. yayukae</emphasis>
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</taxonomicName>
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have two spines on the pedipalpal tarsus. Tibia leg IV with 19 trichobothria with
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<emphasis id="17CBEAF7FF81FF9A0D0D4DC02FD4FF72" box="[369,397,189,214]" italics="true" pageId="6" pageNumber="7">bc</emphasis>
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||
much closer to
|
||
<emphasis id="17CBEAF7FF81FF9A0E344DC02C33FF72" box="[584,618,189,214]" italics="true" pageId="6" pageNumber="7">sbf</emphasis>
|
||
than to
|
||
<emphasis id="17CBEAF7FF81FF9A0EB64DC02CB8FF72" box="[714,737,189,214]" italics="true" pageId="6" pageNumber="7">bf</emphasis>
|
||
;
|
||
<emphasis id="17CBEAF7FF81FF9A0E8C4DC02D5FFF72" box="[752,774,189,214]" italics="true" pageId="6" pageNumber="7">bt</emphasis>
|
||
on fourth basitibial segment of leg IV close to the distal margin of the segment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF81FF940CB94C7A2B02FAF8" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="250036E5FF81FF9A0CB94C7A2D8BFE36" blockId="6.[151,1437,152,812]" pageId="6" pageNumber="7">
|
||
<emphasis id="17CBEAF7FF81FF9A0CB94C7A2F3FFE85" bold="true" box="[197,358,263,289]" pageId="6" pageNumber="7">Description:</emphasis>
|
||
<emphasis id="17CBEAF7FF81FF9A0D134C752FE1FE85" box="[367,440,264,289]" italics="true" pageId="6" pageNumber="7">Male:</emphasis>
|
||
Color in alcohol: Carapace brown, centrally with darker marks; pedipalp brown except for tarsus reddish-brown; legs yellowish-brown, but patella dark brown; abdomen yellowish-brown. The
|
||
<typeStatus id="FA048847FF81FF9A0CB24C2F2F6FFEC8" box="[206,310,338,364]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
|
||
male from Sabah is similar to the
|
||
<typeStatus id="FA048847FF81FF9A0EA74C2F2D10FEC8" box="[731,841,338,364]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
. The
|
||
<typeStatus id="FA048847FF81FF9A0FF04C2F2DADFEC8" box="[908,1012,338,364]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
|
||
male from West Kalimantan much darker than other specimens: carapace, pedipalps and legs dark brown.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF81FF9A0CB94CE02DF8FDD6" blockId="6.[151,1437,152,812]" pageId="6" pageNumber="7">
|
||
Carapace (
|
||
<figureCitation id="BD842A60FF81FF9A0D354CE02FCFFE13" box="[329,406,413,439]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
|
||
): Width about 1.4
|
||
<emphasis id="17CBEAF7FF81FF9A0E0B4CE02CDFFE13" bold="true" box="[631,646,413,439]" italics="true" pageId="6" pageNumber="7">–</emphasis>
|
||
1.5 times its length; surface finely granulate, without setiferous tubercles, with several short setae in frontal area; median sulcus deep in posterior one-third of the carapace; paired lateral sulci present. Flange wide and bent upward. Anterior margin of carapace rounded, with 5 (in
|
||
<typeStatus id="FA048847FF81FF9A0CEB4F702F59FD83" box="[151,256,525,551]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
) or 6 (in
|
||
<typeStatus id="FA048847FF81FF9A0D134F702FB8FD83" box="[367,481,525,551]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
) frontal setae and 11 fine setae. Median eye tubercle black, without apical setae, slightly emarginate antero-medially to form heart-shape; eyes facing antero-laterally. Lateral eyes close to lateral margin of carapace. Frontal process not visible from above.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF81FF9A0CB94F002ABDFC88" blockId="6.[151,1437,152,812]" pageId="6" pageNumber="7">
|
||
Chelicera (
|
||
<figureCitation id="BD842A60FF81FF9A0D394F002FD7FD33" box="[325,398,637,663]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
|
||
): Dorsal surface smooth, with 2 fine frontal setae and 2 fine setae. Basal segment with 4 teeth: lower-most tooth largest, upper-most tooth bicuspid, with upper cusp larger than lower cusp; inner surface with several setae arranged in vertical row; outer surface with small tooth opposite bicuspid tooth, ventrally with several setae near proximal margin. Movable article with 5 (
|
||
<typeStatus id="FA048847FF81FF9A081D4F902A8CFCA3" box="[1121,1237,749,775]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
) or 3 (
|
||
<typeStatus id="FA048847FF81FF9A09514F902BC4FCA3" box="[1325,1437,749,775]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
|
||
(MZB.Ambl.148)) teeth; basal-most tooth largest, subsequent teeth decreasing in size distally.
|
||
</paragraph>
|
||
<caption id="71C0666DFF81FF9A0CEB4A0B2F7CF84A" httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="6" pageNumber="7" targetBox="[173,1405,906,1859]" targetPageId="6">
|
||
<paragraph id="250036E5FF81FF9A0CEB4A0B2F7CF84A" blockId="6.[151,1436,1910,2030]" pageId="6" pageNumber="7">
|
||
<emphasis id="17CBEAF7FF81FF9A0CEB4A0B2F0FF82A" bold="true" box="[151,342,1910,1934]" pageId="6" pageNumber="7">FIGURES 8–12.</emphasis>
|
||
<taxonomicName id="E2BF4D66FF81FF9A0D224A0A2C59F82A" box="[350,512,1911,1934]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="yayukae" status="sp. nov.">
|
||
<emphasis id="17CBEAF7FF81FF9A0D224A0A2C59F82A" box="[350,512,1911,1934]" italics="true" pageId="6" pageNumber="7">Sarax yayukae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="17CBEAF7FF81FF9A0E744A0B2C3BF82A" bold="true" box="[520,610,1910,1934]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="0CF8578CFF81FF9A0E744A0B2C3BF82A" box="[520,610,1910,1934]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB.Ambl.148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="71C0666DFF80FF9B0CEB493B2F05FB39" httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="7" pageNumber="8" targetBox="[201,1350,232,1051]" targetPageId="7">
|
||
<paragraph id="250036E5FF80FF9B0CEB493B2F05FB39" blockId="7.[151,1437,1094,1181]" pageId="7" pageNumber="8">
|
||
<emphasis id="17CBEAF7FF80FF9B0CEB493B2F04FBFA" bold="true" box="[151,349,1094,1118]" pageId="7" pageNumber="8">FIGURES 13–15.</emphasis>
|
||
<taxonomicName id="E2BF4D66FF80FF9B0D1F493B2FA6FBF9" box="[355,511,1094,1117]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yayukae" status="sp. nov.">
|
||
<emphasis id="17CBEAF7FF80FF9B0D1F493B2FA6FBF9" box="[355,511,1094,1117]" italics="true" pageId="7" pageNumber="8">Sarax yayukae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="17CBEAF7FF80FF9B0E7A493B2C02FBFA" bold="true" box="[518,603,1094,1118]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="0CF8578CFF80FF9B0E7A493B2C02FBFA" box="[518,603,1094,1118]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 13. Left pedipalp, antero-dorsal view, female paratype (MZB.Ambl.057); 14. Female genitalia, paratype (MZB.Ambl.057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB.Ambl.059).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="250036E5FF80FF9B0CB949B62CA9FA94" blockId="7.[151,1437,1227,2037]" pageId="7" pageNumber="8">
|
||
Sternum (
|
||
<figureCitation id="BD842A60FF80FF9B0D4549B62FCAFB41" box="[313,403,1227,1253]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="7" pageNumber="8">Fig. 10</figureCitation>
|
||
): First sternite (= tritosternum) elongate, with paired apical and 17 other setae. Second and third sternites rounded and slightly elongate, with 5 and 2 setae, respectively, in addition to paired apical setae. Fourth sternite (= metasternum) with 5 setae.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF80FF9B0CB948462A23F851" blockId="7.[151,1437,1227,2037]" pageId="7" pageNumber="8">
|
||
Pedipalp (
|
||
<figureCitation id="BD842A60FF80FF9B0D4048462FCAFAF1" box="[316,403,1339,1365]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
|
||
): Strong and slender. Trochanter: antero-dorsal margin with several setiferous tubercles and 2 setae, antero-ventral margin with one spine medially and seven setiferous tubercles; distal margin with ventro-anterior apophysis equipped with several setiferous tubercles. Femur: antero-dorsal margin with 3 major spines (length
|
||
<emphasis id="17CBEAF7FF80FF9B0DE648D62C75FA61" bold="true" box="[410,556,1451,1477]" pageId="7" pageNumber="8">F1>F2>F3)</emphasis>
|
||
, several setiferous tubercles and small tubercles (
|
||
<figureCitation id="BD842A60FF80FF9B08ED48D62AB2FA61" box="[1169,1259,1451,1477]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
|
||
); area without setiferous tubercles or small tubercles forming narrow band running length-wise; antero-ventral margin with 3 major spines (length
|
||
<emphasis id="17CBEAF7FF80FF9B0DEE488B2C76F9B4" bold="true" box="[402,559,1526,1552]" pageId="7" pageNumber="8">FI>FII>FIII</emphasis>
|
||
), several minor spines and small tubercles; 1 spine present dorsally of
|
||
<emphasis id="17CBEAF7FF80FF9B0902488B2BC2F9B4" bold="true" box="[1406,1435,1526,1552]" pageId="7" pageNumber="8">FI</emphasis>
|
||
and as long as half length of
|
||
<emphasis id="17CBEAF7FF80FF9B0D914B662C4BF991" bold="true" box="[493,530,1563,1589]" pageId="7" pageNumber="8">FI,</emphasis>
|
||
minor spine present between
|
||
<emphasis id="17CBEAF7FF80FF9B0F084B662DC8F991" bold="true" box="[884,913,1563,1589]" pageId="7" pageNumber="8">FI</emphasis>
|
||
and
|
||
<emphasis id="17CBEAF7FF80FF9B0FB04B662DACF991" bold="true" box="[972,1013,1563,1589]" pageId="7" pageNumber="8">FII</emphasis>
|
||
and between
|
||
<emphasis id="17CBEAF7FF80FF9B08E54B662A9BF991" bold="true" box="[1177,1218,1563,1589]" pageId="7" pageNumber="8">FII</emphasis>
|
||
and
|
||
<emphasis id="17CBEAF7FF80FF9B08804B662B6EF991" bold="true" box="[1276,1335,1563,1589]" pageId="7" pageNumber="8">FIII,</emphasis>
|
||
3 minor spines between
|
||
<emphasis id="17CBEAF7FF80FF9B0D2B4B3D2FD4F9FE" bold="true" box="[343,397,1600,1626]" pageId="7" pageNumber="8">FIII</emphasis>
|
||
and distal margin of femur. Patella: antero-dorsal margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF80FF9B0CEB4B1B2F05F924" bold="true" box="[151,348,1638,1664]" pageId="7" pageNumber="8">P1>P2>P3>P4)</emphasis>
|
||
clumped distally, several minor spines, several setiferous tubercles and small tubercles; major spines located on distal half of patella, 1 minor spine between
|
||
<emphasis id="17CBEAF7FF80FF9B0FAC4BF62DABF901" bold="true" box="[976,1010,1675,1701]" pageId="7" pageNumber="8">P1</emphasis>
|
||
and distal margin of patella, and as long as half of
|
||
<emphasis id="17CBEAF7FF80FF9B0D364BCD2F32F96E" bold="true" box="[330,363,1712,1738]" pageId="7" pageNumber="8">P1</emphasis>
|
||
length, 1 spine between
|
||
<emphasis id="17CBEAF7FF80FF9B0EEF4BCD2C86F96E" bold="true" box="[659,735,1712,1738]" pageId="7" pageNumber="8">P3-P4</emphasis>
|
||
, three minor spines between
|
||
<emphasis id="17CBEAF7FF80FF9B08474BCD2A02F96E" bold="true" box="[1083,1115,1712,1738]" pageId="7" pageNumber="8">P4</emphasis>
|
||
and proximal margin (
|
||
<figureCitation id="BD842A60FF80FF9B09104BCC2EEFF954" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
|
||
); antero-ventral margin with 3 major spines (length
|
||
<emphasis id="17CBEAF7FF80FF9B0F5B4BAB2D97F954" bold="true" box="[807,974,1750,1776]" pageId="7" pageNumber="8">PI>PII>PIII)</emphasis>
|
||
, several setiferous tubercles and small tubercles present, 2 minor spines between
|
||
<emphasis id="17CBEAF7FF80FF9B0EE74B862C89F8B1" bold="true" box="[667,720,1787,1813]" pageId="7" pageNumber="8">PIII</emphasis>
|
||
and proximal margin. Tibia with outer surface smooth and with spines as follows: 2 major spines on antero-dorsal margin, proximal spine less than half as long as distal spine; 1 major spine on antero-ventral margin close to distal margin of tibia. Tarsus completely divided (claw clearly demarcated by articulation), with 3 spines on antero-dorsal margin: proximal spine shortest, medial spine about half as long as distal one, proximal and medial spines separated from each other by about one basal diameter of median one, medial spine separated proximal spine by about 3 basal diameters of medial one; cleaning organ ventrally with about 29 modified hairs; apotele present (
|
||
<figureCitation id="BD842A60FF80FF9B086A4AA62A35F851" box="[1046,1132,2011,2037]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8FFF940CB94DE52C1AFE13" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
Legs (
|
||
<figureCitation id="BD842A60FF8FFF940D734DE52F30FF16" box="[271,361,152,178]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
|
||
): Femora of legs I–IV with small tubercles bearing setae. Tibia and tarsus of leg I with 23 and 41 segments, respectively; tibiae of legs
|
||
<date id="51011025FF8FFF940ED34DC02D6BFF73" box="[687,818,189,215]" pageId="8" pageNumber="9">II and III 2</date>
|
||
-segmented; basitibia of leg
|
||
<date id="51011025FF8FFF9408F94DC02AE5FF73" box="[1157,1212,189,215]" pageId="8" pageNumber="9">IV 4</date>
|
||
-segmented, fourth segment with 1 trichobothrium (value in parentheses: ratio of the distance from the trichobothrium to the proximal margin of the segment against the length of the segment),
|
||
<emphasis id="17CBEAF7FF8FFF940FB64C752DBBFE85" box="[970,994,264,289]" italics="true" pageId="8" pageNumber="9">bt</emphasis>
|
||
(0.74); distitibiae of legs II–IV each with 18 trichobothria (
|
||
<figureCitation id="BD842A60FF8FFF940DDB4C502C58FEE3" box="[423,513,301,327]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
|
||
),
|
||
<emphasis id="17CBEAF7FF8FFF940E6B4C502C76FEE2" box="[535,559,301,326]" italics="true" pageId="8" pageNumber="9">bf</emphasis>
|
||
(0.14),
|
||
<emphasis id="17CBEAF7FF8FFF940EF34C502CEBFEE2" box="[655,690,301,326]" italics="true" pageId="8" pageNumber="9">sbf</emphasis>
|
||
(0.43),
|
||
<emphasis id="17CBEAF7FF8FFF940F6F4C502D76FEE2" box="[787,815,301,326]" italics="true" pageId="8" pageNumber="9">bc</emphasis>
|
||
(0.29),
|
||
<emphasis id="17CBEAF7FF8FFF940FEC4C502DF1FEE2" box="[912,936,301,326]" italics="true" pageId="8" pageNumber="9">bt</emphasis>
|
||
close to distal margin of fourth basitibial segment,
|
||
<emphasis id="17CBEAF7FF8FFF940D744C2F2F7DFECF" box="[264,292,338,363]" italics="true" pageId="8" pageNumber="9">bc</emphasis>
|
||
closer to
|
||
<emphasis id="17CBEAF7FF8FFF940DE44C2F2FE2FECF" box="[408,443,338,363]" italics="true" pageId="8" pageNumber="9">sbf</emphasis>
|
||
than to
|
||
<emphasis id="17CBEAF7FF8FFF940E604C2F2C6AFECF" box="[540,563,338,363]" italics="true" pageId="8" pageNumber="9">bf</emphasis>
|
||
. Tarsi of legs
|
||
<date id="51011025FF8FFF940EA04C2F2D6AFEC8" box="[732,819,338,364]" pageId="8" pageNumber="9">II–IV 4</date>
|
||
-segmented; first segment about as long as length of subsequent three segments combined; second segment with light-yellow transverse line; fourth segment without oblique slit; pulvilli present.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8FFF940CB94CBF2BCCFDE8" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
Genitalia (
|
||
<figureCitation id="BD842A60FF8FFF940D3D4CBF2F8BFE78" box="[321,466,450,476]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="8" pageNumber="9">Figs. 35–36</figureCitation>
|
||
): Covered ventrally by genital operculum rounded apically and with several setae; paired with 2 small lobes apically, 2 brown sclerotized curves, the apical curve larger than the basal in the white weakly sclerotized area, brownish sclerotized area with two projected point medially (
|
||
<figureCitation id="BD842A60FF8FFF9408B64F702B78FD83" box="[1226,1313,525,551]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="8" pageNumber="9">Fig. 36</figureCitation>
|
||
). In dorsal view, paired brown bands extend posteriorly, apical sclerotized area and small ventral lobes present (
|
||
<figureCitation id="BD842A60FF8FFF9409534F4F2BDEFDE8" box="[1327,1415,562,588]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="8" pageNumber="9">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8FFF940CB94F252F5CFCD3" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
<emphasis id="17CBEAF7FF8FFF940CB94F252F7BFDD5" box="[197,290,600,625]" italics="true" pageId="8" pageNumber="9">Female</emphasis>
|
||
(
|
||
<figureCitation id="BD842A60FF8FFF940D484F252F96FDD6" box="[308,463,600,626]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,269,1094,1118]" captionTargetBox="[201,1350,232,1051]" captionTargetId="figure@7.[151,1436,194,1072]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="8" pageNumber="9">Figs. 13–14</figureCitation>
|
||
): Color in alcohol much darker and distinctly different from male as follows: pedipalp distinctly shorter and stouter (
|
||
<figureCitation id="BD842A60FF8FFF940E1C4F002CEDFD33" box="[608,692,637,663]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,269,1094,1118]" captionTargetBox="[201,1350,232,1051]" captionTargetId="figure@7.[151,1436,194,1072]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="8" pageNumber="9">Fig. 13</figureCitation>
|
||
) (see
|
||
<figureCitation id="BD842A60FF8FFF940E864F002D15FD33" box="[762,844,637,663]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
|
||
for male); trochanter with 4 setiferous tubercles, 3 of them arranged in row along antero-dorsal margin; femur with single minor spine each between
|
||
<emphasis id="17CBEAF7FF8FFF9409634FDF2B33FD18" bold="true" box="[1311,1386,674,700]" pageId="8" pageNumber="9">F2-F3</emphasis>
|
||
and between
|
||
<emphasis id="17CBEAF7FF8FFF940D784FBA2F7FFD45" bold="true" box="[260,294,711,737]" pageId="8" pageNumber="9">F3</emphasis>
|
||
and distal margin; antero-ventral margin with 4 minor spines. Patella with four major spines distributed across most of patella. Tarsus with 3 spines: proximal spine the shortest, distal spine the longest, and medial spine about half as long as distal spine, medial and distal spines separated from each other by about three basal diameters of medial spine. Gonopods with paired tube-like, apically-pointed projections (
|
||
<figureCitation id="BD842A60FF8FFF940CE34E202EAEFCD3" box="[159,247,861,887]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,269,1094,1118]" captionTargetBox="[201,1350,232,1051]" captionTargetId="figure@7.[151,1436,194,1072]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8FFF940CB94EFF2FEEFC66" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
<emphasis id="17CBEAF7FF8FFF940CB94EFF2F7EFC3F" box="[197,295,898,923]" italics="true" pageId="8" pageNumber="9">Juvenile</emphasis>
|
||
(
|
||
<figureCitation id="BD842A60FF8FFF940D4A4EFF2FD4FC38" box="[310,397,898,924]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,269,1094,1118]" captionTargetBox="[201,1350,232,1051]" captionTargetId="figure@7.[151,1436,194,1072]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
|
||
): Similar to adult, but color in alcohol much paler, and pedipalpal tarsus with two instead of three spines (
|
||
<figureCitation id="BD842A60FF8FFF940D2E4ED52FF1FC66" box="[338,424,936,962]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,269,1094,1118]" captionTargetBox="[201,1350,232,1051]" captionTargetId="figure@7.[151,1436,194,1072]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8FFF940CB94EB02B02FAF8" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">Measurements (in mm): male (n =3) [female (n =5)]; values for segments of the appendages are their lengths. Body length (excluding chelicera) 9.80–10.60 [8.80–11.80]. Carapace: median length 3.19–4.20 [3.40–3.80]; width 5.40–5.80 [5.00–5.48]; median eyes to anterior margin of carapace 0.04 [0.04]; distance between lateral eyes 2.40–2.52 [2.40–2.60]; lateral eye to anterior margin of carapace 0.60 [0.52–0.60]; lateral eye to lateral margin of carapace 0.20 [0.12–0.20]. Pedipalps: trochanter 1.44–1.72 [1.40]; femur 5.60–6.80 [2.96–3.40]; patella 6.20–7.40 [3.60–3.80]; tibia 1.92–2.20 [1.40–2.00]; tarsus 1.60–2.00 [2.00–2.40]. Leg I: femur 14.80 [10.00–12.20]; patella 0.80 [0.80]. Leg II: femur 8.80 [6.20–7.20], patella 1.20 [1.20]; basitibia 6.40–8.00 [4.80–6.40]; distitibia 3.56–4.20 [3.04–3.40]; metatarsus + tarsus 3.00–3.60 [2.40–2.80]. Leg III: femur 8.00–11.20 [6.80–8.00]; patella 1.20–1.40 [1.20]; basitibia 8.80–10.00 [6.40–7.80]; distitibia 3.60–3.80 [3.20–3.80]; metatarsus+ tarsus 3.60 [2.00–2.80]. Leg IV: femur 8.00–9.60 [7.00–8.00]; patella 1.20 [1.00]; basitibia 9.20–10.80 [7.00–8.00]; distitibia 3.40–3.60 [3.00–3.40]; metatarsus + tarsus 3.20 [2.00–3.00].</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8FFF940CB9481A2B53FA03" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="250036E5FF8FFF940CB9481A2B53FA03" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
<emphasis id="17CBEAF7FF8FFF940CB9481A2F0CFA25" bold="true" box="[197,341,1383,1409]" pageId="8" pageNumber="9">Etymology:</emphasis>
|
||
This species is dedicated to Yayuk R. Suhardjono who collected the
|
||
<typeStatus id="FA048847FF8FFF94080648152AF4FA26" box="[1146,1197,1384,1410]" pageId="8" pageNumber="9">type</typeStatus>
|
||
material from caves in Central Kalimantan, and has contributed much to the knowledge on cave biology in
|
||
<collectingCountry id="5DA87675FF8FFF9408EC48F02B5FFA03" box="[1168,1286,1421,1447]" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8FFF940CB948CF2D78F923" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="250036E5FF8FFF940CB948CF2F83FA56" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
<emphasis id="17CBEAF7FF8FFF940CB948CF2F31FA68" bold="true" box="[197,360,1458,1484]" pageId="8" pageNumber="9">Distribution:</emphasis>
|
||
<taxonomicName id="E2BF4D66FF8FFF940D0C48CE2C45FA68" box="[368,540,1459,1484]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8FFF940D0C48CE2C45FA68" box="[368,540,1459,1484]" italics="true" pageId="8" pageNumber="9">Sarax yayukae</emphasis>
|
||
</taxonomicName>
|
||
is found in Central and West Kalimantan (
|
||
<collectingCountry id="5DA87675FF8FFF94086B48CF2AD6FA68" box="[1047,1167,1458,1484]" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
|
||
), and Manukan Island, Sabah (
|
||
<collectingCountry id="5DA87675FF8FFF940C9248A52F04FA56" box="[238,349,1496,1522]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
|
||
) (
|
||
<figureCitation id="BD842A60FF8FFF940D0948A52F94FA56" box="[373,461,1496,1522]" captionStart="FIGURES 41 – 42" captionStartId="16.[151,272,1727,1751]" captionTargetBox="[200,1385,221,1687]" captionTargetId="figure@16.[191,1395,194,1706]" captionTargetPageId="16" captionText="FIGURES 41 – 42. Map showing the distribution of Sarax species: 41. The distribution in Southeast Asia, except for Sarax species from India and New Guinea. 42. A detailed map showing the distribution in the Sangkulirang Karst, East Kalimantan." httpUri="https://zenodo.org/record/197818/files/figure.png" pageId="8" pageNumber="9">Fig. 41</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8FFF940CB948802D78F923" blockId="8.[151,1437,152,2007]" pageId="8" pageNumber="9">
|
||
<emphasis id="17CBEAF7FF8FFF940CB948802FC0F9B3" bold="true" box="[197,409,1533,1559]" pageId="8" pageNumber="9">Natural history:</emphasis>
|
||
This species inhabits caves (Liang Puruk, Liang Hajuq and Gua Kelasi), as well as occurring under logs in forest (Manukan Island, Sabah). Detailed descriptions of the caves in Central Kalimantan were given in
|
||
<bibRefCitation id="412E4B14FF8FFF940DB04B352D63F9C6" author="Rahmadi" box="[460,826,1608,1634]" pageId="8" pageNumber="9" refString="Rahmadi, C. & Suhardjono, Y. R. (2006) Gua-Gua di Topus, Hulu Sungai Barito Kalimantan Tengah, Tinjauan speleleologi. Gunung Sewu, 2 (1), 53 - 67." type="journal article" year="2006">Rahmadi & Suhardjono (2006)</bibRefCitation>
|
||
. The species was observed preying on cave crickets,
|
||
<emphasis id="17CBEAF7FF8FFF940CEB4B102F64F922" box="[151,317,1645,1670]" italics="true" pageId="8" pageNumber="9">Diestramenna</emphasis>
|
||
sp., in Liang Puruk, Central Kalimantan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8FFF950CB94BEF2C14FF73" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="250036E5FF8FFF950CB94BEF2C14FF73" blockId="8.[151,1437,152,2007]" lastBlockId="9.[151,1436,152,439]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<emphasis id="17CBEAF7FF8FFF940CB94BEF2F1BF908" bold="true" box="[197,322,1682,1708]" pageId="8" pageNumber="9">Remarks:</emphasis>
|
||
<taxonomicName id="E2BF4D66FF8FFF940D364BEE2FA3F908" box="[330,506,1683,1708]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8FFF940D364BEE2FA3F908" box="[330,506,1683,1708]" italics="true" pageId="8" pageNumber="9">Sarax yayukae</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from other congeners by its distinct sexual dimorphism, the presence of three spines on the pedipalpal tarsus in adults (
|
||
<figureCitation id="BD842A60FF8FFF940F0D4BC52D9DF976" box="[881,964,1720,1746]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
|
||
) and two in juveniles (
|
||
<figureCitation id="BD842A60FF8FFF9408AD4BC52B7FF976" box="[1233,1318,1720,1746]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,269,1094,1118]" captionTargetBox="[201,1350,232,1051]" captionTargetId="figure@7.[151,1436,194,1072]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059)." httpUri="https://zenodo.org/record/197813/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
|
||
), the tibia of leg IV having 19 trichobothria arranged with
|
||
<emphasis id="17CBEAF7FF8FFF940EB54BA02CBCF952" box="[713,741,1757,1782]" italics="true" pageId="8" pageNumber="9">bc</emphasis>
|
||
much closer to
|
||
<emphasis id="17CBEAF7FF8FFF940FE24BA02D98F952" box="[926,961,1757,1782]" italics="true" pageId="8" pageNumber="9">sbf</emphasis>
|
||
than to
|
||
<emphasis id="17CBEAF7FF8FFF9408624BA02A6DF952" box="[1054,1076,1757,1782]" italics="true" pageId="8" pageNumber="9">bf</emphasis>
|
||
, and
|
||
<emphasis id="17CBEAF7FF8FFF9408094BA02AD2F952" box="[1141,1163,1757,1782]" italics="true" pageId="8" pageNumber="9">bt</emphasis>
|
||
on the fourth basitibial segment of leg IV being close to the distal margin of the segment (
|
||
<figureCitation id="BD842A60FF8FFF940FB44A7F2A7DF8B8" box="[968,1060,1794,1820]" captionStart="FIGURES 8 – 12" captionStartId="6.[151,273,1910,1934]" captionTargetBox="[173,1405,906,1859]" captionTargetId="figure@6.[151,1436,844,1890]" captionTargetPageId="6" captionText="FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV." httpUri="https://zenodo.org/record/197812/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
|
||
). The juvenile of
|
||
<taxonomicName id="E2BF4D66FF8FFF9408814A7E2BD9F8B8" box="[1277,1408,1795,1820]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8FFF9408814A7E2BD9F8B8" box="[1277,1408,1795,1820]" italics="true" pageId="8" pageNumber="9">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="E2BF4D66FF8FFF940D6E4A552F88F8E5" box="[274,465,1832,1857]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="17CBEAF7FF8FFF940D6E4A552F88F8E5" box="[274,465,1832,1857]" italics="true" pageId="8" pageNumber="9">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
in having two spines on the pedipalpal tarsus, but the spines in
|
||
<taxonomicName id="E2BF4D66FF8FFF9408A84A552B0CF8E5" box="[1236,1365,1832,1857]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8FFF9408A84A552B0CF8E5" box="[1236,1365,1832,1857]" italics="true" pageId="8" pageNumber="9">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
(both juvenile and adult) are long while those in
|
||
<taxonomicName id="E2BF4D66FF8FFF940EF74A302D1FF8C2" box="[651,838,1869,1894]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="17CBEAF7FF8FFF940EF74A302D1FF8C2" box="[651,838,1869,1894]" italics="true" pageId="8" pageNumber="9">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
are short and minute.
|
||
<taxonomicName id="E2BF4D66FF8FFF9408304A302B6CF8C2" box="[1100,1333,1869,1894]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="17CBEAF7FF8FFF9408304A302B6CF8C2" box="[1100,1333,1869,1894]" italics="true" pageId="8" pageNumber="9">Sarax sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
has four spines on the antero-dorsal margin of the pedipalpal femur and three spines in the juvenile of
|
||
<taxonomicName id="E2BF4D66FF8FFF94089E4A0E2B06F828" box="[1250,1375,1907,1932]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8FFF94089E4A0E2B06F828" box="[1250,1375,1907,1932]" italics="true" pageId="8" pageNumber="9">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
. The pedipalpal patella of
|
||
<taxonomicName id="E2BF4D66FF8FFF940DEA4AE52C0EF815" box="[406,599,1944,1969]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="17CBEAF7FF8FFF940DEA4AE52C0EF815" box="[406,599,1944,1969]" italics="true" pageId="8" pageNumber="9">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
has four major spines on the antero-dorsal margin instead of three in juveniles of
|
||
<taxonomicName id="E2BF4D66FF8FFF940D504AC02FF4F872" box="[300,429,1981,2006]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8FFF940D504AC02FF4F872" box="[300,429,1981,2006]" italics="true" pageId="8" pageNumber="9">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
. The number and arrangement of the trichobothria on the tibia of leg IV also differ between both species:
|
||
<taxonomicName id="E2BF4D66FF8EFF950DDD4DE52C46FF15" box="[417,543,152,177]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8EFF950DDD4DE52C46FF15" box="[417,543,152,177]" italics="true" pageId="9" pageNumber="10">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
(both juvenile and adult) has 19 trichobothria while
|
||
<taxonomicName id="E2BF4D66FF8EFF9508F04DE52B11FF15" box="[1164,1352,152,177]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="17CBEAF7FF8EFF9508F04DE52B11FF15" box="[1164,1352,152,177]" italics="true" pageId="9" pageNumber="10">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
has 17 trichobothria (
|
||
<bibRefCitation id="412E4B14FF8EFF950D414DC02FACFF73" author="Weygoldt" box="[317,501,189,215]" pageId="9" pageNumber="10" refString="Weygoldt, P. (1996) Evolutionary morphology of whip spiders: towards a phylogenetic system (Chelicerata: Arachnida: Amblypygi). Journal of Zoological Systematics and Evolution Research, 34, 185 - 202." type="journal article" year="1996">Weygoldt 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="412E4B14FF8EFF950E7D4DC02C18FF73" author="Weygoldt" box="[513,577,189,215]" pageId="9" pageNumber="10" refString="Weygoldt, P. (2000) Whip spiders: Their biology, morphology and systematics. Apollo Books, Stenstrup." type="book" year="2000">2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8EFF950CB94D9F2A95FE13" pageId="9" pageNumber="10" type="reference_group">
|
||
<paragraph id="250036E5FF8EFF950CB94D9F2A95FE13" blockId="9.[151,1436,152,439]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="E2BF4D66FF8EFF950CB94D9F2FA2FF58" ID-CoL="6XMF6" authority="Fage 1946" authorityName="Fage" authorityYear="1946" box="[197,507,226,252]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="davidovi">
|
||
<emphasis id="17CBEAF7FF8EFF950CB94D9F2F2EFF5F" box="[197,375,226,251]" italics="true" pageId="9" pageNumber="10">Sarax davidovi</emphasis>
|
||
<bibRefCitation id="412E4B14FF8EFF950D034D9F2FA2FF58" author="Fage" box="[383,507,226,252]" pageId="9" pageNumber="10" refString="Fage, L. (1946) Scorpions et Pedipalpes de l'Indochine Francaise. Annales de la Societe Entomologique de France, 113, 71 - 81." type="journal article" year="1946">Fage 1946</bibRefCitation>
|
||
</taxonomicName>
|
||
from Indochina shows distinct sexual dimorphism and has three spines on the antero-dorsal margin of the pedipalpal tarsus as in
|
||
<taxonomicName id="E2BF4D66FF8EFF950F784C752DDFFE85" ID-CoL="79MHC" box="[772,902,264,289]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8EFF950F784C752DDFFE85" box="[772,902,264,289]" italics="true" pageId="9" pageNumber="10">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
. They differ in the number of spines on the antero-dorsal margin of the male pedipalpal patella (4 spines decreasing in size from distal to proximal in
|
||
<taxonomicName id="E2BF4D66FF8EFF9509FA4C502EAEFECF" ID-CoL="79MHC" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8EFF9509FA4C502EAEFECF" italics="true" pageId="9" pageNumber="10">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
; 2 long spines in
|
||
<taxonomicName id="E2BF4D66FF8EFF950DB04C2F2C09FECF" ID-CoL="6XMF6" box="[460,592,338,363]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="davidovi">
|
||
<emphasis id="17CBEAF7FF8EFF950DB04C2F2C09FECF" box="[460,592,338,363]" italics="true" pageId="9" pageNumber="10">S. davidovi</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="412E4B14FF8EFF950EF34C2F2D57FEC8" author="Fage" box="[655,782,338,364]" pageId="9" pageNumber="10" refString="Fage, L. (1946) Scorpions et Pedipalpes de l'Indochine Francaise. Annales de la Societe Entomologique de France, 113, 71 - 81." type="journal article" year="1946">Fage 1946</bibRefCitation>
|
||
, fig. 4)) and the shape of the carapace (anterior margin rounded in
|
||
<taxonomicName id="E2BF4D66FF8EFF950D5C4C052FC6FE35" ID-CoL="79MHC" box="[288,415,376,401]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF8EFF950D5C4C052FC6FE35" box="[288,415,376,401]" italics="true" pageId="9" pageNumber="10">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
, with broad flange on the antero-lateral corners in
|
||
<taxonomicName id="E2BF4D66FF8EFF95087F4C052ADEFE35" ID-CoL="6XMF6" box="[1027,1159,376,401]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="davidovi">
|
||
<emphasis id="17CBEAF7FF8EFF95087F4C052ADEFE35" box="[1027,1159,376,401]" italics="true" pageId="9" pageNumber="10">S. davidovi</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="412E4B14FF8EFF9508C24C052B67FE36" author="Fage" box="[1214,1342,376,402]" pageId="9" pageNumber="10" refString="Fage, L. (1946) Scorpions et Pedipalpes de l'Indochine Francaise. Annales de la Societe Entomologique de France, 113, 71 - 81." type="journal article" year="1946">Fage 1946</bibRefCitation>
|
||
, fig. 2). The number and arrangement of the trichobothria on tibia leg IV of
|
||
<taxonomicName id="E2BF4D66FF8EFF950FCF4CE02A6CFE12" ID-CoL="6XMF6" box="[947,1077,413,438]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="davidovi">
|
||
<emphasis id="17CBEAF7FF8EFF950FCF4CE02A6CFE12" box="[947,1077,413,438]" italics="true" pageId="9" pageNumber="10">S. davidovi</emphasis>
|
||
</taxonomicName>
|
||
is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |