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<document id="9F7A33F8F37AE59D530EE87C70C95E26" ID-CLB-Dataset="7433" ID-DOI="10.5852/ejt.2021.744.1303" ID-GBIF-Dataset="f62caa9b-9794-41d0-8551-a6ccb1d2e092" ID-ISSN="2118-9773" ID-Zenodo-Dep="4673914" ID-ZooBank="BBB5D83C-54BE-43AB-B4B8-8B896B5909BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617944647591" checkinUser="plazi" docAuthor="Chatzimanolis, Stylianos &amp; Brunke, Adam J." docDate="2021" docId="752487D7FF81067AFDDDFB89FB493451" docLanguage="en" docName="ejt-744_67-82.pdf" docOrigin="European Journal of Taxonomy 744" docSource="http://dx.doi.org/10.5852/ejt.2021.744.1303" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ikaros paramo Chatzimanolis &amp; Brunke 2021, gen. et sp. nov." docType="treatment" docUuid="8FE14A94-3312-4BA8-901A-DAE9542DF2B4" docUuidSource="ZooBank" docVersion="7" lastPageNumber="77" masterDocId="891DFFAFFF890670FFB8FFBBFFDC3525" masterDocTitle="A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position" masterLastPageNumber="82" masterPageNumber="67" pageNumber="75" updateTime="1698936220334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D8C1B41E71812BA742D71A5E22E6B799" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="04020DFB656AFB1E93F8D78F6B92D0A9">
<mods:title id="16EC07843A83DB4A683A403731FF38D2">A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position</mods:title>
</mods:titleInfo>
<mods:name id="1CFE83C0936DFED37688A1D71432C200" type="personal">
<mods:role id="FB0CD1F1A24604414A1D9A140E65F239">
<mods:roleTerm id="99B7020B16FE84A122F906FF6EB6D95B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2259FB36830F63BA21E34B297FF1A746">Chatzimanolis, Stylianos</mods:namePart>
<mods:nameIdentifier id="33423A50ED514C80E73EE98A39E20855" type="LSID">0BFB18A5-6424-4827-B500-DD36E99E1123</mods:nameIdentifier>
<mods:affiliation id="FF401CB8121D8CFA8F761FB0DAF85A54">Department of Biology, Geology and Environmental Science, University of Tennessee at Chattanooga, 615 McCallie Ave. Dept 2653, Chattanooga, TN 37403, USA.</mods:affiliation>
<mods:nameIdentifier id="03C621E7750A14D4E2E3CECF4C282079" type="email">stylianos-chatzimanolis@utc.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="60319C94FAEAB239A247B69C954FB8F9" type="personal">
<mods:role id="E003E2653BD0C8CC1462BF0675913B3E">
<mods:roleTerm id="791427721E4EA6C65C37D040686491E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5607D91B96334C90AAF082B5FD495805">Brunke, Adam J.</mods:namePart>
<mods:nameIdentifier id="848479F3926EF3731389C2CB6D6EBD4B" type="LSID">8AEFAA65-7FA7-422C-86ED-5CD4FE611985</mods:nameIdentifier>
<mods:affiliation id="0618B6BE3EEAE023A1B95E8EC7B0B91D">Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Avenue, K. W. Neatby Building, Ottawa, ON K 1 A 0 C 6, Canada.</mods:affiliation>
<mods:nameIdentifier id="51A8AACCC081164058AE55B5E0B020B5" type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4AD5102C7A5E5218E997AE9C7209C8CC">text</mods:typeOfResource>
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<mods:titleInfo id="6583F1587B8D7B01A63901EBF0B98F57">
<mods:title id="08CDCDAA3A175500580FED887F24C8B9">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="A398435B0745BAA643BB97827D1C0316">
<mods:date id="CAD5E72AA8E285A04634E2D61673B3A3">2021</mods:date>
<mods:detail id="327516A992AA700269B75DCDDFEABB85" type="pubDate">
<mods:number id="6353D6A1748573ED615E70575C41FB99">2021-04-08</mods:number>
</mods:detail>
<mods:detail id="965122675A16A4FF10CCD5616536D05C" type="volume">
<mods:number id="F34348D1661263337D1FC61752987C91">744</mods:number>
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<mods:start id="064EC68296AD8E8D7559D28ABEB9F117">67</mods:start>
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<mods:url id="4FF26A4EFDCAACF8FE4C57AF12DB4B61">http://dx.doi.org/10.5852/ejt.2021.744.1303</mods:url>
</mods:location>
<mods:classification id="7F645BD44F394F5891619DF5716DDE82">journal article</mods:classification>
<mods:identifier id="54CD221318052042D259990C15A3D8CF" type="CLB-Dataset">7433</mods:identifier>
<mods:identifier id="7805EC8FC30ACA20D0F8A1F992414496" type="DOI">10.5852/ejt.2021.744.1303</mods:identifier>
<mods:identifier id="D335B95355B4797194560C05896AB412" type="GBIF-Dataset">f62caa9b-9794-41d0-8551-a6ccb1d2e092</mods:identifier>
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<mods:identifier id="635CBF7957223B8D6610B493D065D219" type="Zenodo-Dep">4673914</mods:identifier>
<mods:identifier id="19EDB38FE181488716FF774587D246AB" type="ZooBank">BBB5D83C-54BE-43AB-B4B8-8B896B5909BF</mods:identifier>
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<treatment id="752487D7FF81067AFDDDFB89FB493451" ID-DOI="http://doi.org/10.5281/zenodo.4673922" ID-GBIF-Taxon="180887974" ID-Zenodo-Dep="4673922" LSID="urn:lsid:zoobank.org:act:8FE14A94-3312-4BA8-901A-DAE9542DF2B4" httpUri="http://treatment.plazi.org/id/752487D7FF81067AFDDDFB89FB493451" lastPageId="10" lastPageNumber="77" pageId="8" pageNumber="75">
<subSubSection id="B597654AFF810678FDDDFB89FC133168" box="[613,975,1074,1101]" pageId="8" pageNumber="75" type="nomenclature">
<paragraph id="FD3236C1FF810678FDDDFB89FC133168" blockId="8.[363,1224,1074,1171]" box="[613,975,1074,1101]" pageId="8" pageNumber="75">
<heading id="A67A81ADFF810678FDDDFB89FC133168" box="[613,975,1074,1101]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="75" reason="2">
<taxonomicName id="3A8D4D42FF810678FDDDFB89FCC43168" ID-CoL="8Y4VQ" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[613,792,1074,1101]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="paramo" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF810678FDDDFB89FCC43168" bold="true" box="[613,792,1074,1101]" italics="true" pageId="8" pageNumber="75">Ikaros paramo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF810678FCA7FB88FC133168" box="[799,975,1075,1101]" pageId="8" pageNumber="75" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B597654AFF810678FED3FBEEFC8B31B6" pageId="8" pageNumber="75" type="description">
<paragraph id="FD3236C1FF810678FED3FBEEFB143155" blockId="8.[363,1224,1074,1171]" box="[363,1224,1109,1136]" pageId="8" pageNumber="75">
<uri id="891C3AC3FF810678FED3FBEEFB143155" box="[363,1224,1109,1136]" pageId="8" pageNumber="75">
urn:lsid:zoobank.org:act:
<uuid id="892B0C14FF810678FD32FBEEFB143155" box="[650,1224,1109,1136]" pageId="8" pageNumber="75">8FE14A94-3312-4BA8-901A-DAE9542DF2B4</uuid>
</uri>
</paragraph>
<paragraph id="FD3236C1FF810678FD65FBC2FC8B31B6" blockId="8.[363,1224,1074,1171]" box="[733,855,1145,1171]" pageId="8" pageNumber="75">
<figureCitation id="65B62A44FF810678FD65FBC2FCE031B6" box="[733,828,1145,1171]" captionStart="Fig" captionStartId="6.[189,232,1913,1939]" captionTargetBox="[201,1380,691,1837]" captionTargetId="figure-114@6.[191,1399,688,1877]" captionTargetPageId="6" captionText="Fig. 2. Habitus photographs of species of Ikaros gen. nov. A. I. apteros gen. et sp. nov. Holotype, ♂ (SDEI). B. I. paramo gen. et sp. nov. Paratype, ♂ (barcode: SM05500251, UTCI). C. I. polygonos gen. et sp. nov. Holotype, ♂ (barcode: SM0114364, MIZA). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4673920" httpUri="https://zenodo.org/record/4673920/files/figure.png" pageId="8" pageNumber="75">Figs 2B</figureCitation>
,
<figureCitation id="65B62A44FF810678FCF0FBC2FC8B31B6" box="[840,855,1145,1171]" captionStart="Fig" captionStartId="9.[189,232,1952,1978]" captionTargetBox="[189,1386,1199,1915]" captionTargetId="figure-272@9.[189,1398,1199,1927]" captionTargetPageId="9" captionText="Fig. 4. Aedeagus ofIkaros paramo gen. et sp. nov.A. Dorsal view.B. Lateral view.C. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673926" httpUri="https://zenodo.org/record/4673926/files/figure.png" pageId="8" pageNumber="75">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B597654AFF810678FF05FB01FC96306F" pageId="8" pageNumber="75" type="diagnosis">
<paragraph id="FD3236C1FF810678FF05FB01FEEA31F1" blockId="8.[189,310,1210,1236]" box="[189,310,1210,1236]" pageId="8" pageNumber="75">
<heading id="A67A81ADFF810678FF05FB01FEEA31F1" bold="true" box="[189,310,1210,1236]" fontSize="11" level="3" pageId="8" pageNumber="75" reason="3">
<emphasis id="CFF9EAD3FF810678FF05FB01FEEA31F1" bold="true" box="[189,310,1210,1236]" pageId="8" pageNumber="75">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF810678FF05FB52FC96306F" blockId="8.[189,1399,1256,1354]" pageId="8" pageNumber="75">
Among species of
<taxonomicName id="3A8D4D42FF810678FE19FB52FE373026" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[417,491,1257,1283]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CFF9EAD3FF810678FE19FB52FE373026" box="[417,491,1257,1283]" italics="true" pageId="8" pageNumber="75">Ikaros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF810678FE4AFB52FD833026" box="[498,607,1257,1283]" pageId="8" pageNumber="75" rank="genus">gen. nov.</taxonomicNameLabel>
that lack arch-like carina on terga 3–5,
<taxonomicName id="3A8D4D42FF810678FBFBFB52FB653027" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[1091,1209,1257,1282]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="paramo" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF810678FBFBFB52FB653027" box="[1091,1209,1257,1282]" italics="true" pageId="8" pageNumber="75">I. paramo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF810678FB78FB52FAAB3026" box="[1216,1399,1257,1283]" pageId="8" pageNumber="75" rank="species">gen. et sp. nov.</taxonomicNameLabel>
has punctures arranged in multiple rows, whereas
<taxonomicName id="3A8D4D42FF810678FCB5FAB6FCA13003" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[781,893,1293,1318]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="apteros" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF810678FCB5FAB6FCA13003" box="[781,893,1293,1318]" italics="true" pageId="8" pageNumber="75">I. apteros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF810678FC3CFAB7FBEA3003" box="[900,1078,1292,1318]" pageId="8" pageNumber="75" rank="species">gen. et sp. nov.</taxonomicNameLabel>
lacks any punctures on the pronotum, beyond the punctures along the median area.
</paragraph>
</subSubSection>
<subSubSection id="B597654AFF810678FF05FACAFCCD30F9" pageId="8" pageNumber="75" type="etymology">
<paragraph id="FD3236C1FF810678FF05FACAFE9830AE" blockId="8.[189,324,1393,1419]" box="[189,324,1393,1419]" pageId="8" pageNumber="75">
<heading id="A67A81ADFF810678FF05FACAFE9830AE" bold="true" box="[189,324,1393,1419]" fontSize="11" level="3" pageId="8" pageNumber="75" reason="3">
<emphasis id="CFF9EAD3FF810678FF05FACAFE9830AE" bold="true" box="[189,324,1393,1419]" pageId="8" pageNumber="75">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF810678FF05FA24FCCD30F9" blockId="8.[189,1398,1439,1501]" pageId="8" pageNumber="75">The specific epithet is derived from the name of the ecosystem (páramo) where the specimens were collected. It is treated here as a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="B597654AFF810679FF05F9BFFE423463" lastPageId="9" lastPageNumber="76" pageId="8" pageNumber="75" type="materials_examined">
<paragraph id="FD3236C1FF810678FF05F9BFFEB03338" blockId="8.[189,364,1539,1566]" box="[189,364,1539,1566]" pageId="8" pageNumber="75">
<heading id="A67A81ADFF810678FF05F9BFFEB03338" bold="true" box="[189,364,1539,1566]" fontSize="11" level="3" pageId="8" pageNumber="75" reason="3">
<emphasis id="CFF9EAD3FF810678FF05F9BFFEB03338" bold="true" box="[189,364,1539,1566]" pageId="8" pageNumber="75">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF810678FF52F989FBC33393" blockId="8.[189,1399,1586,1718]" pageId="8" pageNumber="75">
<materialsCitation id="4DE53C9CFF810678FF52F989FBC33393" ID-GBIF-Occurrence="3069587307" collectingDate="2001-03-09" collectingDateMax="2001-03-13" collectingDateMin="2001-03-09" collectingMethod="Pitfall" collectionCode="IAVH" collectorName="P. Reina Leg." country="Colombia" elevation="2860" latitude="5.4166665" location="SFF Iguaque Qda. Los Francos" longLatPrecision="1305" longitude="-73.45" municipality="Colombia" pageId="8" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca" typeStatus="holotype">
<emphasis id="CFF9EAD3FF810678FF52F989FE803369" bold="true" box="[234,348,1586,1612]" pageId="8" pageNumber="75">
<typeStatus id="22368863FF810678FF52F989FE803369" box="[234,348,1586,1612]" pageId="8" pageNumber="75" type="holotype">Holotype</typeStatus>
</emphasis>
(here designated)
<collectingCountry id="859A7651FF810678FF05F9EDFE843355" box="[189,344,1622,1648]" name="Colombia" pageId="8" pageNumber="75">COLOMBIA</collectingCountry>
•
<specimenCount id="EB8BFD48FF810678FEC1F9EDFE4E3355" box="[377,402,1622,1648]" pageId="8" pageNumber="75" type="male">♂</specimenCount>
; “
<collectingMunicipality id="1D56ACBBFF810678FE15F9EEFDF4334A" box="[429,552,1621,1647]" country="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" name="Colombia" pageId="8" pageNumber="75" stateProvince="Boyaca">Colombia</collectingMunicipality>
:
<collectingRegion id="3F49F823FF810678FD8FF9EDFD4E3355" box="[567,658,1622,1648]" country="Colombia" name="Boyaca" pageId="8" pageNumber="75">Boyacá</collectingRegion>
,
<location id="F852601AFF810678FD1AF9EDFBF83355" LSID="urn:lsid:plazi:treatment:752487D7FF81067AFDDDFB89FB493451:F852601AFF810678FD1AF9EDFBF83355" box="[674,1060,1621,1648]" country="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Colombia" name="SFF Iguaque Qda. Los Francos" pageId="8" pageNumber="75" stateProvince="Boyaca">SFF Iguaque Qda. Los Francos</location>
,
<geoCoordinate id="98B95006FF810678FB8CF9EEFB573355" box="[1076,1163,1621,1648]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="8" pageNumber="75" precision="925" value="5.4166665">5°25’N</geoCoordinate>
<geoCoordinate id="98B95006FF810678FB2EF9EDFADD3355" box="[1174,1281,1622,1648]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="8" pageNumber="75" precision="925" value="-73.45">73°27’W</geoCoordinate>
,
<quantity id="3A759B24FF810678FAAAF9EEFAAD3355" box="[1298,1393,1621,1648]" metricMagnitude="3" metricUnit="m" metricValue="2.86" pageId="8" pageNumber="75" unit="m" value="2860.0">
<elevation id="76A0D1F2FF810678FAAAF9EEFAAD3355" box="[1298,1393,1621,1648]" metricMagnitude="3" metricUnit="m" metricValue="2.86" pageId="8" pageNumber="75" unit="m" value="2860.0">2860 m</elevation>
</quantity>
,
<collectingMethod id="24CC4ED6FF810678FF05F9C2FED433B6" box="[189,264,1657,1683]" pageId="8" pageNumber="75">Pitfall</collectingMethod>
,
<date id="89331001FF810678FEAAF9C2FE6D33B6" box="[274,433,1657,1683]" pageId="8" pageNumber="75" value="2001-03-09" valueMax="2001-03-13" valueMin="2001-03-09">
<collectingDate id="9977E9E9FF810678FEAAF9C2FE6D33B6" box="[274,433,1657,1683]" pageId="8" pageNumber="75" value="2001-03-09" valueMax="2001-03-13" valueMin="2001-03-09">9–13.iii.2001</collectingDate>
</date>
,
<collectorName id="50785317FF810678FE03F9C2FD8B33B6" box="[443,599,1657,1683]" pageId="8" pageNumber="75">P. Reina Leg.</collectorName>
, M.1345 / SM0650483 [barcode label] / HOLOTYPE
<taxonomicName id="3A8D4D42FF810678FB72F9C2FE243393" authority="Chatzimanolis and Brunke" authorityName="Chatzimanolis and Brunke" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="paramo">
<emphasis id="CFF9EAD3FF810678FB72F9C2FAA933B7" box="[1226,1397,1657,1683]" italics="true" pageId="8" pageNumber="75">Ikaros paramo</emphasis>
Chatzimanolis and Brunke
</taxonomicName>
, des. Chatzimanolis and Brunke 2020”;
<collectionCode id="9B9CAE04FF810678FC6AF927FBC73393" box="[978,1051,1692,1718]" country="Colombia" httpUri="http://grbio.org/cool/6phm-mi1y" name="Instituto de Ivestigacion de los Recursos Biologicos Alexander von Humboldt" pageId="8" pageNumber="75">IAVH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="FD3236C1FF810679FF52F966FE423463" blockId="8.[189,1399,1757,1995]" lastBlockId="9.[189,1398,265,326]" lastPageId="9" lastPageNumber="76" pageId="8" pageNumber="75">
<materialsCitation id="4DE53C9CFF810678FF52F966FBB2321B" ID-GBIF-Occurrence="3069587301" collectingDate="2000-09-03" collectingDateMax="2000-09-05" collectingDateMin="2000-09-03" collectingMethod="Pitfall" collectionCode="UTCI" collectorName="P. Reina Leg." country="Colombia" elevation="2850" latitude="5.4166665" location="SFF Iguaque Qda. Los Francos" longLatPrecision="1305" longitude="-73.45" municipality="Colombia" pageId="8" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca" typeStatus="paratype">
<emphasis id="CFF9EAD3FF810678FF52F966FEBB33D2" bold="true" box="[234,359,1757,1783]" pageId="8" pageNumber="75">
<typeStatus id="22368863FF810678FF52F966FEBB33D2" box="[234,359,1757,1783]" pageId="8" pageNumber="75" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="859A7651FF810678FF05F8BAFE84323E" box="[189,344,1793,1819]" name="Colombia" pageId="8" pageNumber="75">COLOMBIA</collectingCountry>
•
<specimenCount id="EB8BFD48FF810678FECEF8BAFE74323E" box="[374,424,1793,1819]" pageId="8" pageNumber="75" type="male">1 ♂</specimenCount>
; “
<collectingMunicipality id="1D56ACBBFF810678FE7AF8BBFDE1323F" box="[450,573,1792,1818]" country="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" name="Colombia" pageId="8" pageNumber="75" stateProvince="Boyaca">Colombia</collectingMunicipality>
:
<collectingRegion id="3F49F823FF810678FDF3F8BAFD7A323E" box="[587,678,1793,1819]" country="Colombia" name="Boyaca" pageId="8" pageNumber="75">Boyacá</collectingRegion>
,
<location id="F852601AFF810678FD0CF8BAFBEC323E" LSID="urn:lsid:plazi:treatment:752487D7FF81067AFDDDFB89FB493451:F852601AFF810678FD0CF8BAFBEC323E" box="[692,1072,1792,1819]" country="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Colombia" name="SFF Iguaque Qda. Los Francos" pageId="8" pageNumber="75" stateProvince="Boyaca">SFF Iguaque Qda. Los Francos</location>
,
<geoCoordinate id="98B95006FF810678FB87F944FB4E323E" box="[1087,1170,1791,1819]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="8" pageNumber="75" precision="925" value="5.4166665">5°25′N</geoCoordinate>
<geoCoordinate id="98B95006FF810678FB24F944FAD9323E" box="[1180,1285,1791,1819]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="8" pageNumber="75" precision="925" value="-73.45">73°27′W</geoCoordinate>
,
<quantity id="3A759B24FF810678FAABF8BBFAAD323E" box="[1299,1393,1792,1819]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="8" pageNumber="75" unit="m" value="2850.0">
<elevation id="76A0D1F2FF810678FAABF8BBFAAD323E" box="[1299,1393,1792,1819]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="8" pageNumber="75" unit="m" value="2850.0">2850 m</elevation>
</quantity>
,
<collectingMethod id="24CC4ED6FF810678FF05F89FFED4321B" box="[189,264,1828,1854]" pageId="8" pageNumber="75">Pitfall</collectingMethod>
,
<date id="89331001FF810678FEABF898FE7D321B" box="[275,417,1827,1854]" pageId="8" pageNumber="75" value="2000-09-03" valueMax="2000-09-05" valueMin="2000-09-03">
<collectingDate id="9977E9E9FF810678FEABF898FE7D321B" box="[275,417,1827,1854]" pageId="8" pageNumber="75" value="2000-09-03" valueMax="2000-09-05" valueMin="2000-09-03">3–5.ix.2000</collectingDate>
</date>
,
<collectorName id="50785317FF810678FE15F89FFD96321B" box="[429,586,1828,1854]" pageId="8" pageNumber="75">P. Reina Leg.</collectorName>
, M.756 / SM05500251 [barcode label]”;
<collectionCode id="9B9CAE04FF810678FB92F89FFBB2321B" box="[1066,1134,1828,1854]" pageId="8" pageNumber="75">UTCI</collectionCode>
</materialsCitation>
•
<materialsCitation id="4DE53C9CFF810678FB3EF89FFC483244" ID-GBIF-Occurrence="3069587308" collectingDate="2000-09-03" collectingDateMax="2000-09-05" collectingDateMin="2000-09-03" collectingMethod="Pitfall" collectionCode="SEMC" collectorName="P. Reina Leg." country="Colombia" elevation="2850" latitude="5.4166665" location="SFF Iguaque Qda. Los Francos" longLatPrecision="1305" longitude="-73.45" municipality="Colombia" pageId="8" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca" typeStatus="paratype">
<specimenCount id="EB8BFD48FF810678FB3EF89FFB68321B" box="[1158,1204,1828,1854]" pageId="8" pageNumber="75" type="male">1 ♂</specimenCount>
; same collection data as for preceding; “SM05500250 [barcode label]”;
<collectionCode id="9B9CAE04FF810678FCFCF8FCFC483244" box="[836,916,1863,1889]" collectionName="USA, Kansas, Lawrence, University of Kansas, Snow Entomological Museum" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="8" pageNumber="75" type="Museum">SEMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="4DE53C9CFF810678FC15F8FCFDA5328D" ID-GBIF-Occurrence="3069587303" collectingDate="2000-04-19" collectingDateMax="2001-03-13" collectingDateMin="2000-04-19" collectingMethod="Malaise / Pitfall" collectionCode="SEMC" collectorName="P. Reina Leg." country="Colombia" county="Colombia" elevation="2855" latitude="5.4166665" location="SFF Iguaque Cabana Mamaramos" longLatPrecision="1305" longitude="-73.45" municipality="Iguaque Cabana Mamaramos" pageId="8" pageNumber="75" specimenCount="2" specimenCount-female="2" stateProvince="Boyaca" typeStatus="paratype">
<specimenCount id="EB8BFD48FF810678FC15F8FCFC063244" box="[941,986,1863,1889]" pageId="8" pageNumber="75" type="female">1 ♀</specimenCount>
; “
<collectingCountry id="859A7651FF810678FC4BF8FCFBB23244" box="[1011,1134,1863,1889]" name="Colombia" pageId="8" pageNumber="75">Colombia</collectingCountry>
:
<collectingRegion id="3F49F823FF810678FBC1F8FCFB083244" box="[1145,1236,1863,1889]" country="Colombia" name="Boyaca" pageId="8" pageNumber="75">Boyacá</collectingRegion>
,
<location id="F852601AFF810678FB58F8FCFE6A32A0" LSID="urn:lsid:plazi:treatment:752487D7FF81067AFDDDFB89FB493451:F852601AFF810678FB58F8FCFE6A32A0" country="Colombia" county="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Iguaque Cabana Mamaramos" name="SFF Iguaque Cabana Mamaramos" pageId="8" pageNumber="75" stateProvince="Boyaca">SFF Iguaque Cabanā Mamaramos</location>
, m4,
<geoCoordinate id="98B95006FF810678FE44F8D3FD9332A1" box="[508,591,1896,1924]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="8" pageNumber="75" precision="925" value="5.4166665">5°25′N</geoCoordinate>
<geoCoordinate id="98B95006FF810678FDE2F8D3FD1F32A1" box="[602,707,1896,1924]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="8" pageNumber="75" precision="925" value="-73.45">73°27′W</geoCoordinate>
,
<quantity id="3A759B24FF810678FD6AF8D1FCEC32A0" box="[722,816,1898,1925]" metricMagnitude="3" metricUnit="m" metricValue="2.855" pageId="8" pageNumber="75" unit="m" value="2855.0">
<elevation id="76A0D1F2FF810678FD6AF8D1FCEC32A0" box="[722,816,1898,1925]" metricMagnitude="3" metricUnit="m" metricValue="2.855" pageId="8" pageNumber="75" unit="m" value="2855.0">2855 m</elevation>
</quantity>
,
<collectingMethod id="24CC4ED6FF810678FCF8F8D1FC7C32A1" box="[832,928,1898,1924]" pageId="8" pageNumber="75">Malaise</collectingMethod>
,
<date id="89331001FF810678FC17F8D1FBBC32A1" box="[943,1120,1898,1924]" pageId="8" pageNumber="75" value="2000-04-19" valueMax="2000-05-06" valueMin="2000-04-19">
<collectingDate id="9977E9E9FF810678FC17F8D1FBBC32A1" box="[943,1120,1898,1924]" pageId="8" pageNumber="75" value="2000-04-19" valueMax="2000-05-06" valueMin="2000-04-19">19.iv–6.v.2000</collectingDate>
</date>
,
<collectorName id="50785317FF810678FBC8F8D0FACA32A0" box="[1136,1302,1898,1925]" pageId="8" pageNumber="75">P. Reina Leg.</collectorName>
, M.55 / SM0548719 [barcode label];
<collectionCode id="9B9CAE04FF810678FDAFF836FDBB3282" box="[535,615,1933,1959]" collectionName="USA, Kansas, Lawrence, University of Kansas, Snow Entomological Museum" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="8" pageNumber="75" type="Museum">SEMC</collectionCode>
•
</materialsCitation>
<materialsCitation id="4DE53C9CFF810679FD39F836FE463463" ID-GBIF-Occurrence="3069587305" collectingDate="2000-04-19" collectingDateMax="2001-03-13" collectingDateMin="2000-04-19" collectingMethod="Malaise / Pitfall" collectionCode="SEMC" collectorName="P. Reina Leg." country="Colombia" county="Colombia" elevation="2855" lastPageId="9" lastPageNumber="76" latitude="5.4166665" location="SFF Iguaque Cabana Mamaramos" longLatPrecision="1305" longitude="-73.45" municipality="Iguaque Cabana Mamaramos" pageId="8" pageNumber="75" specimenCount="2" specimenCount-female="2" stateProvince="Boyaca" typeStatus="paratype">
<specimenCount id="EB8BFD48FF810678FD39F836FD73328D" box="[641,687,1933,1960]" pageId="8" pageNumber="75" type="female">1 ♀</specimenCount>
; “
<collectingCountry id="859A7651FF810678FD70F836FC9F3282" box="[712,835,1933,1959]" name="Colombia" pageId="8" pageNumber="75">Colombia</collectingCountry>
:
<collectingRegion id="3F49F823FF810678FCF7F836FC763282" box="[847,938,1933,1959]" country="Colombia" name="Boyaca" pageId="8" pageNumber="75">Boyacá</collectingRegion>
,
<location id="F852601AFF810678FC0EF836FACA3282" LSID="urn:lsid:plazi:treatment:752487D7FF81067AFDDDFB89FB493451:F852601AFF810678FC0EF836FACA3282" box="[950,1302,1933,1960]" country="Colombia" county="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Iguaque Cabana Mamaramos" name="SFF Iguaque Cabana Carrizal" pageId="8" pageNumber="75" stateProvince="Boyaca">SFF Iguaque Cabanā Carrizal</location>
,
<geoCoordinate id="98B95006FF810678FA9BF837FAAA328D" box="[1315,1398,1932,1960]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="8" pageNumber="75" precision="925" value="5.4166665">5°25′N</geoCoordinate>
<geoCoordinate id="98B95006FF810678FF05F814FEFA32EE" box="[189,294,1967,1995]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="8" pageNumber="75" precision="925" value="-73.45">73°27′W</geoCoordinate>
,
<quantity id="3A759B24FF810678FE8AF80BFE5132EE" box="[306,397,1968,1995]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="8" pageNumber="75" unit="m" value="2850.0">
<elevation id="76A0D1F2FF810678FE8AF80BFE5132EE" box="[306,397,1968,1995]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="8" pageNumber="75" unit="m" value="2850.0">2850 m</elevation>
</quantity>
,
<collectingMethod id="24CC4ED6FF810678FE21F80BFE3832EF" box="[409,484,1968,1994]" pageId="8" pageNumber="75">Pitfall</collectingMethod>
,
<date id="89331001FF810678FE48F80BFD5332EF" box="[496,655,1968,1994]" pageId="8" pageNumber="75" value="2001-03-09" valueMax="2001-03-13" valueMin="2001-03-09">
<collectingDate id="9977E9E9FF810678FE48F80BFD5332EF" box="[496,655,1968,1994]" pageId="8" pageNumber="75" value="2001-03-09" valueMax="2001-03-13" valueMin="2001-03-09">9–13.iii.2001</collectingDate>
</date>
,
<collectorName id="50785317FF810678FD23F80AFCE632EE" box="[667,826,1968,1995]" pageId="8" pageNumber="75">P. Reina Leg.</collectorName>
, M.1346 / SM0645316 [barcode label]”;
<collectionCode id="9B9CAE04FF810678FAA7F80AFAAF32EE" box="[1311,1395,1969,1995]" collectionName="USA, Kansas, Lawrence, University of Kansas, Snow Entomological Museum" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="8" pageNumber="75" type="Museum">SEMC</collectionCode>
. All paratypes with label “PARATYPE
<taxonomicName id="3A8D4D42FF800679FD37FEB2FB5B3406" authority="Chatzimanolis and Brunke" authorityName="Chatzimanolis and Brunke" box="[655,1159,265,291]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="paramo">
<emphasis id="CFF9EAD3FF800679FD37FEB2FCE13406" box="[655,829,265,291]" italics="true" pageId="9" pageNumber="76">Ikaros paramo</emphasis>
Chatzimanolis and Brunke
</taxonomicName>
, des. Chatzimanolis and Brunke 2020”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B597654AFF800679FF05FED6FD9C36E3" pageId="9" pageNumber="76" type="description">
<paragraph id="FD3236C1FF800679FF05FED6FE9234A2" blockId="9.[189,334,365,391]" box="[189,334,365,391]" pageId="9" pageNumber="76">
<heading id="A67A81ADFF800679FF05FED6FE9234A2" bold="true" box="[189,334,365,391]" fontSize="11" level="3" pageId="9" pageNumber="76" reason="3">
<emphasis id="CFF9EAD3FF800679FF05FED6FE9234A2" bold="true" box="[189,334,365,391]" pageId="9" pageNumber="76">Description</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF800679FF05FE27FD9C36E3" blockId="9.[187,1401,412,967]" pageId="9" pageNumber="76">
Forebody length
<quantity id="3A759B24FF800679FE3EFE27FDC83492" box="[390,532,412,439]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.9" metricValueMin="5.3" pageId="9" pageNumber="76" unit="mm" value="5.6" valueMax="5.9" valueMin="5.3">5.3–5.9 mm</quantity>
long. Coloration reddish brown with head and mouthparts slightly darker brown. Head transverse, HW/HL ratio = 1.21. Epicranium with numerous large punctures, except impunctate centre; punctures not contiguous, distance between punctures varies; with faint polygonshaped microsculpture. Labial palpus with palpomere 3 widest apically, subparallel-sided.Antennomeres with crown-like macrosetae shorter than length of antennomere. Pronotum longer than wide, PW/PL ratio = 0.88; surface of pronotum with a median impunctate area as wide as 4–5 punctures; with 2–3 rows of disorganized punctures in addition to rows flanking impunctate centre; with faint polygonshaped microsculpture. Elytra shorter than pronotum, EL/PL ratio = 0.79. Elytra with large, deep punctures, distance between punctures equals to width of 0.5–1 punctures; elytra with faint polygonshaped microsculpture. Abdominal terga 3–5 without arch-like carina. Male secondary sexual structures with shallow emargination on sternum 7; with shallow, narrow U-shaped emargination on sternum 8; borders of emargination on sternum 7 and 8 appearing ‘shaved’ (with no setae), but less so than in
<taxonomicName id="3A8D4D42FF800679FF05FCF8FEEC3679" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[189,304,835,860]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="apteros" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF800679FF05FCF8FEEC3679" box="[189,304,835,860]" italics="true" pageId="9" pageNumber="76">I. apteros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF800679FE82FCF8FE283678" box="[314,500,835,861]" pageId="9" pageNumber="76" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Aedeagus as in
<figureCitation id="65B62A44FF800679FD06FCF9FCD53678" box="[702,777,834,861]" captionStart="Fig" captionStartId="9.[189,232,1952,1978]" captionTargetBox="[189,1386,1199,1915]" captionTargetId="figure-272@9.[189,1398,1199,1927]" captionTargetPageId="9" captionText="Fig. 4. Aedeagus ofIkaros paramo gen. et sp. nov.A. Dorsal view.B. Lateral view.C. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673926" httpUri="https://zenodo.org/record/4673926/files/figure.png" pageId="9" pageNumber="76">Fig. 4</figureCitation>
; in dorsal view paramere subequal to median lobe; paramere broad, converging to rounded tip; in lateral view paramere more or less straight. Median lobe in dorsal view broad, narrowing to rounded apex; in lateral view median lobe becoming narrower near apex, with small subapical tooth.
</paragraph>
</subSubSection>
<subSubSection id="B597654AFF800679FF05FC56FD95317F" pageId="9" pageNumber="76" type="distribution">
<paragraph id="FD3236C1FF800679FF05FC56FE8A3122" blockId="9.[189,342,1005,1031]" box="[189,342,1005,1031]" pageId="9" pageNumber="76">
<heading id="A67A81ADFF800679FF05FC56FE8A3122" bold="true" box="[189,342,1005,1031]" fontSize="11" level="3" pageId="9" pageNumber="76" reason="3">
<emphasis id="CFF9EAD3FF800679FF05FC56FE8A3122" bold="true" box="[189,342,1005,1031]" pageId="9" pageNumber="76">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF800679FF05FBA7FD95317F" blockId="9.[189,1399,1052,1114]" pageId="9" pageNumber="76">
This species is only known from Iguaque Fauna and Flora Sanctuary (SFF Iguaque) in the
<collectingRegion id="3F49F823FF800679FAA7FBA7FE9B317F" country="Colombia" name="Boyaca" pageId="9" pageNumber="76">Boyacá Department</collectingRegion>
of
<collectingCountry id="859A7651FF800679FED7FB84FE3F317C" box="[367,483,1087,1113]" name="Colombia" pageId="9" pageNumber="76">Colombia</collectingCountry>
(
<figureCitation id="65B62A44FF800679FE4AFB84FDE6317F" box="[498,570,1087,1114]" captionStart="Fig" captionStartId="12.[191,234,1945,1971]" captionTargetBox="[189,1398,680,1912]" captionTargetId="figure-145@12.[189,1398,677,1912]" captionTargetPageId="12" captionText="Fig. 6. Distribution map of Ikaros apteros gen. et sp. nov. (square), I. paramo gen. et sp. nov. (circle) and I. polygonos gen. et sp. nov. (triangle)." figureDoi="http://doi.org/10.5281/zenodo.4673930" httpUri="https://zenodo.org/record/4673930/files/figure.png" pageId="9" pageNumber="76">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="A9F26649FF800679FF05F81BFE8F32F8" ID-DOI="http://doi.org/10.5281/zenodo.4673926" ID-Zenodo-Dep="4673926" httpUri="https://zenodo.org/record/4673926/files/figure.png" pageId="9" pageNumber="76" startId="9.[189,232,1952,1978]" targetBox="[189,1386,1199,1915]" targetPageId="9">
<paragraph id="FD3236C1FF800679FF05F81BFE8F32F8" blockId="9.[187,1398,1952,2013]" pageId="9" pageNumber="76">
<emphasis id="CFF9EAD3FF800679FF05F81BFED4329F" bold="true" box="[189,264,1952,1978]" pageId="9" pageNumber="76">Fig. 4.</emphasis>
Aedeagus of
<taxonomicName id="3A8D4D42FF800679FE27F81BFD95329F" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[415,585,1952,1978]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="paramo" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF800679FE27F81BFD95329F" box="[415,585,1952,1978]" italics="true" pageId="9" pageNumber="76">Ikaros paramo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF800679FDEAF81AFD20329E" box="[594,764,1953,1979]" pageId="9" pageNumber="76" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="CFF9EAD3FF800679FD47F81BFCC8329F" bold="true" box="[767,788,1952,1978]" pageId="9" pageNumber="76">A</emphasis>
. Dorsal view.
<emphasis id="CFF9EAD3FF800679FC0FF81BFC17329F" bold="true" box="[951,971,1952,1978]" pageId="9" pageNumber="76">B</emphasis>
. Lateral view.
<emphasis id="CFF9EAD3FF800679FBCCF81BFB55329F" bold="true" box="[1140,1161,1952,1978]" pageId="9" pageNumber="76">C</emphasis>
. Detail of paramere, ventral view.
</paragraph>
</caption>
<subSubSection id="B597654AFF83067AFF05FEB2FB493451" pageId="10" pageNumber="77" type="discussion">
<paragraph id="FD3236C1FF83067AFF05FEB2FEF33406" blockId="10.[189,303,265,291]" box="[189,303,265,291]" pageId="10" pageNumber="77">
<heading id="A67A81ADFF83067AFF05FEB2FEF33406" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="10" pageNumber="77" reason="3">
<emphasis id="CFF9EAD3FF83067AFF05FEB2FEF33406" bold="true" box="[189,303,265,291]" pageId="10" pageNumber="77">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF83067AFF05FE8CFB493451" blockId="10.[189,1398,311,373]" pageId="10" pageNumber="77">
The
<typeStatus id="22368863FF83067AFF4EFE8CFE813474" box="[246,349,311,337]" pageId="10" pageNumber="77" type="holotype">holotype</typeStatus>
is currently in the collection of SEMC but due to the collecting permit requirements (Z. Falin pers. com.), it will be transferred to the IAVH collection in the near future.
</paragraph>
</subSubSection>
</treatment>
</document>
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