210 lines
24 KiB
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210 lines
24 KiB
XML
<document id="D3019F3DAFC0DCA3204ED45B5BC72466" ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23683" ID-GBIF-Dataset="6ef290f2-f65d-4dfa-afbd-2fbb505860ab" ID-PMC="PMC5904508" ID-Pensoft-Pub="1313-2970-745-1" ID-PubMed="29670448" ID-ZooBank="16F1AE595650485F9D8C6149E962D461" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-745-1" ModsDocOrigin="ZooKeys 745" ModsDocTitle="Synopsis of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae)" checkinTime="1521799043702" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. & Branham, Marc A." docDate="2018" docId="5E6B8BC3BB8C5B646B85600F2BB84EDD" docLanguage="en" docName="ZooKeys 745: 1-99" docOrigin="ZooKeys 745" docSource="http://dx.doi.org/10.3897/zookeys.745.23683" docTitle="Arriguttia Martinez 1960" docType="treatment" docVersion="5" lastPageNumber="43" masterDocId="C24FFFFC630B3774FFCFDB4EFFAFAD27" masterDocTitle="Synopsis of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae)" masterLastPageNumber="99" masterPageNumber="1" pageNumber="41" updateTime="1701315738437" updateUser="plazi">
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<mods:title id="982B4C66FAB3B8838FD5D92CDF3F5291">Synopsis of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae)</mods:title>
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<mods:namePart id="53A30C60C9CB9FF83CFD4FE488281A1C">Moore, Matthew R.</mods:namePart>
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<mods:namePart id="9965ADE4B76CC635EC0E2A1AD3F9D9AB">Cave, Ronald D.</mods:namePart>
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<mods:namePart id="6659D15F8FDA158C049D27F7CD7DD37B">Branham, Marc A.</mods:namePart>
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<mods:typeOfResource id="2A9923DB0CB4F32B4B440C495EEF57DC">text</mods:typeOfResource>
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<mods:date id="83430B28AE617CDF2F32553C6DABC982">2018</mods:date>
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<mods:detail id="DBA2DDE533A8C755CC05C505FE323AA8" type="volume">
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<mods:number id="F8A3DF82C2FAACDBEBC09E5431E24A6F">745</mods:number>
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<mods:classification id="03412AFA81CC7585637B8CAC8FA03F75">journal article</mods:classification>
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<mods:identifier id="D4F98FE8E9DD6AF42CC9944BFC734970" type="DOI">http://dx.doi.org/10.3897/zookeys.745.23683</mods:identifier>
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<treatment id="5E6B8BC3BB8C5B646B85600F2BB84EDD" ID-GBIF-Taxon="142479967" LSID="urn:lsid:plazi:treatment:5E6B8BC3BB8C5B646B85600F2BB84EDD" httpUri="http://treatment.plazi.org/id/5E6B8BC3BB8C5B646B85600F2BB84EDD" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
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<subSubSection id="FC63DC333C54099B76D27E7DA910A229" pageId="40" pageNumber="41" type="nomenclature">
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<paragraph id="6B57236B7C5284197CD4D939EE1F7B3B" pageId="40" pageNumber="41">
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<taxonomicName id="40B9D8775BD3F80D1CFAD749305E2FC7" ID-CoL="338N" authority="Martinez, 1960" authorityName="Martinez" authorityYear="1960" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
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Arriguttia
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<normalizedToken id="546B9105F0BC4C9224444467730B393F" originalValue="Martínez">Martinez</normalizedToken>
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, 1960
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="40902F0772DBE276A1654CC09C19C110" pageId="40" pageNumber="41" type="type species">
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<paragraph id="0B10EABA5A19C1FFECB8180EDE48F733" pageId="40" pageNumber="41">Type species.</paragraph>
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<paragraph id="1706FFABF57EBEF6660FDB14165C6D6D" pageId="40" pageNumber="41">
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<taxonomicName id="BAD575FDB7264F6A29BB2BD084A410EC" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala brevissima" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="brevissima">Cyclocephala brevissima</taxonomicName>
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Arrow, 1911, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="EA027779B2D0A5E90A3B550C3C5B8379" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="valid taxa">
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<paragraph id="DCC0F826EF3B2B7CB57D307D34709A97" pageId="40" pageNumber="41">Valid taxa.</paragraph>
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<paragraph id="F7CCCDC6854F7739E5C399D41E6BA37D" pageId="40" pageNumber="41">Two species.</paragraph>
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<paragraph id="04242A60789404C6836B76A75C6DBE50" pageId="40" pageNumber="41">
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<taxonomicName id="0884B143B369B46D2220C30A0F198E17" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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contains two South American species known only from the Brazilian Amazon, Guyana, and French Guiana (
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<bibRefCitation id="CC77B04BB9CEC854C95185E02A7B7999" pageId="40" pageNumber="41">Arrow 1911</bibRefCitation>
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,
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<bibRefCitation id="F15C57B64F8FC43B6C7CFC9F8FFFD451" pageId="40" pageNumber="41">1937b</bibRefCitation>
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,
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<bibRefCitation id="C499FF9C85B304C46D60397AFA1853B9" author="Blackwelder, RE" journalOrPublisher="Bulletin of the United States National Museum" pageId="64" pageNumber="65" pagination="189 - 341" title="Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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,
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<bibRefCitation id="69D16CF4AB5D0D068AF7547569ED9B09" author="Martinez, A" journalOrPublisher="Ciencia (Mexico)" pageId="79" pageNumber="80" pagination="97 - 98" title="Un nuevo genero de Cyclocephalini (Col., Scarab., Dyasti.)." volume="20" year="1960 a">
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<normalizedToken id="C05117B86FBBEBEC43C2499446E49F06" originalValue="Martínez">Martinez</normalizedToken>
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1960a
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</bibRefCitation>
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,
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<bibRefCitation id="0D698500CBD5666BAD7DC5E2EA236DA2" author="Martinez, A" journalOrPublisher="Ciencia (Mexico)" pageId="79" pageNumber="80" pagination="185 - 190" title="Notas sobre Cyclocephalini americanos con descripcion de dos nuevas especies (Col. Scarab., Dynast.)." volume="26" year="1968 a">1968a</bibRefCitation>
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,
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<bibRefCitation id="A444012F5A81C54861BABD25DD838298" author="Endrodi, S" journalOrPublisher="Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden" pageId="70" pageNumber="71" pagination="1 - 460" title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil." volume="33" year="1966">
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<normalizedToken id="4FB34D53869588FE216DC11093E2E63E" originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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,
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<bibRefCitation id="42D32306087CA9A3F429F3FB87C79908" author="Endrodi, S" journalOrPublisher="Junk Publisher, Dordrecht" pageId="71" pageNumber="72" title="The Dynastinae of the World. Dr. W." year="1985 a">1985a</bibRefCitation>
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,
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<bibRefCitation id="AC9A710B74D8154FED092749A18C862C" author="Ponchel, Y" journalOrPublisher="United States Department of Agriculture Department Bulletin" pageId="82" pageNumber="83" url="http://dynastidae.voila.net/biologie.html" year="2006">Ponchel 2006</bibRefCitation>
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,
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<bibRefCitation id="FD0EC5B3A87D3042EE519E6E08623638" author="Ponchel, Y" journalOrPublisher="Contribution a L'Etude des Coleopteres de Guyane" pageId="82" pageNumber="83" pagination="60 - 61" title="Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae)." volume="4" year="2011">2011</bibRefCitation>
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,
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<bibRefCitation id="8AD839588EAFF43034866A1ADC08D8F5" author="Ponchel, Y" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="82" pageNumber="83" pagination="36 - 38" title="Note sur les Dynastinae floricoles de Guyane (Coleoptera, Scarabaeidae)." volume="9" year="2015">2015</bibRefCitation>
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) (Fig. 53). Very little is known about the biology of
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<taxonomicName id="6684B030DB3A386D3B7443A299C7B328" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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species.
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<taxonomicName id="6AB08B828DA7E46CD6C278AAFB85FB90" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia brevissima" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="brevissima">Arriguttia brevissima</taxonomicName>
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(Arrow) feeds within the inflorescences of
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<taxonomicName id="F01268D9663166FF73417CBA554E4A8B" class="Insecta" family="Geometridae" genus="Victoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Victoria" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Victoria</taxonomicName>
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sp. in Brazil (
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<bibRefCitation id="38F7AC7EF247CCB75263E9C6995FE82F" author="Martinez, A" journalOrPublisher="Ciencia (Mexico)" pageId="79" pageNumber="80" pagination="185 - 190" title="Notas sobre Cyclocephalini americanos con descripcion de dos nuevas especies (Col. Scarab., Dynast.)." volume="26" year="1968 a">
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<normalizedToken id="244D7D5DFC400830DA345964D626BD15" originalValue="Martínez">Martinez</normalizedToken>
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1968a
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</bibRefCitation>
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). In French Guiana,
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<taxonomicName id="2EE4A4D91329F44588E0A7779698BA3C" lsidName="A. brevissima" pageId="40" pageNumber="41" rank="species" species="brevissima">A. brevissima</taxonomicName>
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was found in the spathes of an unidentified terrestrial aroid (
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<taxonomicName id="3F35DBE1176CBDBB871813F690223274" family="Araceae" lsidName="Oryctomorphina" pageId="40" pageNumber="41" rank="family">Araceae</taxonomicName>
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) (
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<bibRefCitation id="C1EF06268C82B0EFDDCBD89C30B6DB86" author="Ponchel, Y" journalOrPublisher="United States Department of Agriculture Department Bulletin" pageId="82" pageNumber="83" url="http://dynastidae.voila.net/biologie.html" year="2006">Ponchel 2006</bibRefCitation>
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,
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<bibRefCitation id="03074F626BD3051A54507CE4982A1176" author="Ponchel, Y" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="82" pageNumber="83" pagination="36 - 38" title="Note sur les Dynastinae floricoles de Guyane (Coleoptera, Scarabaeidae)." volume="9" year="2015">2015</bibRefCitation>
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). In Brazilian cerrado habitat,
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<taxonomicName id="6F09EF42D1C41FE3751D5602B8490C19" lsidName="A. brevissima" pageId="40" pageNumber="41" rank="species" species="brevissima">A. brevissima</taxonomicName>
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are floral visitors of
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<taxonomicName id="3008F16CE607975FF7D547B070C69A9F" class="Magnoliopsida" family="Annonaceae" genus="Annona" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Annona coriacea" order="Magnoliales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="coriacea">Annona coriacea</taxonomicName>
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Mart. and are likely late-season, secondary pollinators of this species (
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<bibRefCitation id="F11C27A0D982FBF7EFAA15409CB36F59" author="Costa, MS" journalOrPublisher="Journal of Economic Entomology" pageId="67" pageNumber="68" url="https://doi.org/10.1371/journal.pone.0171092" year="2017">Costa et al. 2017</bibRefCitation>
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). Specimens of
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<taxonomicName id="5587C655927EB0CEBE3AB958E8237048" lsidName="A. brevissima" pageId="40" pageNumber="41" rank="species" species="brevissima">A. brevissima</taxonomicName>
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have been collected at lights at night (
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<bibRefCitation id="7C52AA9D02AA629F5A66B986B7A5B649" author="Martinez, A" journalOrPublisher="Ciencia (Mexico)" pageId="79" pageNumber="80" pagination="185 - 190" title="Notas sobre Cyclocephalini americanos con descripcion de dos nuevas especies (Col. Scarab., Dynast.)." volume="26" year="1968 a">
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<normalizedToken id="AF992262B64E4CCE12A771AD64FEF6DB" originalValue="Martínez">Martinez</normalizedToken>
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1968a
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</bibRefCitation>
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). The immature stages are undescribed and unknown.
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</paragraph>
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<caption id="915DA64468FC461F753241BF75A20378" pageId="40" pageNumber="41">
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<paragraph id="0E72C3B4643B13F66D0F476690562F84" pageId="40" pageNumber="41">
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Figure 53. Country-level distribution of
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<taxonomicName id="E7907A334B9C6E8E8766ECC2A0061244" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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taxa in South America. Numbers indicate taxa per country.
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</paragraph>
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</caption>
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<paragraph id="E406B46AA3D0D396271AD917C5B61690" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<taxonomicName id="6002E4F1490EC09067782CED2D65AAEC" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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was compared to
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<taxonomicName id="4822FB113FC2751420B378A66ABA8F9E" class="Insecta" family="Scarabaeidae" genus="Surutu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Surutu" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Surutu</taxonomicName>
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in the original description of the genus (
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<bibRefCitation id="C0B6128B303B2C413EC6FE2B987C59BD" author="Martinez, A" journalOrPublisher="Ciencia (Mexico)" pageId="79" pageNumber="80" pagination="97 - 98" title="Un nuevo genero de Cyclocephalini (Col., Scarab., Dyasti.)." volume="20" year="1960 a">
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<normalizedToken id="EF0880B7AD2356318239D6C2554F4FA9" originalValue="Martínez">Martinez</normalizedToken>
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1960a
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</bibRefCitation>
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). This is possibly confusing for identification purposes.
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<taxonomicName id="91B31ED3ABD71FD7006D899405483E5F" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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shares many more characters with
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<taxonomicName id="C66CC4E01DC6AB610DFE5490BA28CA31" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
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and
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<taxonomicName id="489C29A371FFDA800A9874A941FF1320" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
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than with
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<taxonomicName id="728C7D117060D051BD4792FE1AF00A74" class="Insecta" family="Scarabaeidae" genus="Surutu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Surutu" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Surutu</taxonomicName>
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.
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<taxonomicName id="C7243095B4844F4F57916438464347AC" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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species can be recognized by the following combination of characters: 1) dorsal coloration varying from all black or with variable dark, reddish coloration on the elytra; 2) body convex and anteroposteriorly compressed, creating a relatively round gestalt; 3) clypeus quadrate in dorsal view, with sides nearly parallel, and the apex distinctly reflexed upwards (most obvious in lateral view); 4) frontoclypeal suture complete medially; 5) males with anterolateral margin of the mandibles weakly toothed; 6) mandibular molar area with rows of circular micropunctures; 7) apical margin of mentum weakly emarginated; 8) galea of the maxilla on inner surface with 3 fused basal teeth,
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<pageBreakToken id="C6C61BBCAEBF3721449AF178A7A2AE92" pageId="41" pageNumber="42" start="start">a</pageBreakToken>
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free median tooth, and 2 fused apical teeth (3-1-2 arrangement); 9) pronotum with broadly incomplete beaded basal margin; 10) males and females with 3 protibial teeth, basal tooth reduced, removed from the more apical 2 teeth, and oriented anteriorly; 11) protibial spur straight to weakly deflexed; 12) males with inner protarsal claw enlarged and narrowly cleft at apex; 13) mesocoxae widely separated; 14) meso- and metatibiae with distal, transverse carinae; 15) metacoxae with lateral edge perpendicular to ventral surface; 16) anterior edge of hindwing distal to apical hinge lacking setae and with produced, membranous border; 17) vein RA with 2 rows of pegs extending distally nearly to margin of apical hinge.
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</paragraph>
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<paragraph id="E7231E98BBBB5F257CA9EF13735DB87D" pageId="42" pageNumber="43">
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<pageBreakToken id="DD38E1EA4E26536D3149424A2D608649" pageId="42" pageNumber="43" start="start">The</pageBreakToken>
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relationships of
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<taxonomicName id="B3079AF2D1EFA52721AD8F3B7955BA56" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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to other cyclocephaline genera have not been clearly discussed in the literature.
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<bibRefCitation id="D51B9E888BDCCBDCED89024702567510" author="Martinez, A" journalOrPublisher="Ciencia (Mexico)" pageId="79" pageNumber="80" pagination="185 - 190" title="Notas sobre Cyclocephalini americanos con descripcion de dos nuevas especies (Col. Scarab., Dynast.)." volume="26" year="1968 a">
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<normalizedToken id="5693B03790C2382915654BD2FD1AAFD0" originalValue="Martínez">Martinez</normalizedToken>
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(1968a)
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</bibRefCitation>
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stated that
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<taxonomicName id="D8CAB1504D8C03EFC5BEE798050C36F6" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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should be "placed next to"
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<taxonomicName id="E9461DC53D7E7ECF77468AC7D1348C9F" class="Insecta" family="Scarabaeidae" genus="Surutu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Surutu" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Surutu</taxonomicName>
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, but he did not offer any character justifications for this hypothesis.
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<bibRefCitation id="F20CBE20E584D01DDB3889D385AF91A6" author="Endrodi, S" journalOrPublisher="Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden" pageId="70" pageNumber="71" pagination="1 - 460" title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil." volume="33" year="1966">
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<normalizedToken id="2CDDEEDD66B80B2AAF0B964D7C2B6C59" originalValue="Endrődi">Endrodi</normalizedToken>
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(1966)
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</bibRefCitation>
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considered
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<taxonomicName id="BB1537DD67914692F1810F6F87DE0E4E" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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to be a
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<normalizedToken id="0B9A6AFB77B3D0CA3366D1FB43A35079" originalValue="“primitive”">"primitive"</normalizedToken>
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cyclocephaline based on his poorly justified character analysis.
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<taxonomicName id="CF462B8F2A3D5D7A9F0EA4FFB97F7D81" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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shares hindwing characters (two rows of pegs on vein RA and a membrane on the leading edge of the hindwing distal to the apical hinge) with
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<taxonomicName id="0EEA4F47401869E75350F34772BBA33B" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
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,
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<taxonomicName id="C009BCAD073B24CD5FFD00E96071A01C" class="Insecta" family="Scarabaeidae" genus="Aspidolea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspidolea" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Aspidolea</taxonomicName>
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, and
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<taxonomicName id="D36EA20A10EC9F79430F0BDBABB63699" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
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. The form of the maxilla (3-1-2 teeth arrangement), the mandibular form (males with anterolateral margin weakly toothed and the molar area with rows of circular micropunctures), the incomplete bead on the basal margin of the pronotum, and the shape and arrangement of the protibial teeth are shared among
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|
<taxonomicName id="B006817A012503E0BC5FBCDB8798838E" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
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|
,
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|
<taxonomicName id="5745F11A5711E5F55C51FBDFC7E367D0" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
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|
, and some
|
|
<taxonomicName id="62855C3F9E92A5F90B6CE58500E022BB" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
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|
(especially species like
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|
<taxonomicName id="596745A58AC063CB0B036189B0199DCD" lsidName="C. sexpunctata" pageId="42" pageNumber="43" rank="species" species="sexpunctata">C. sexpunctata</taxonomicName>
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|
Laporte and species formerly placed in
|
|
<taxonomicName id="B231A53C61D2A791CD730C1D00644D62" class="Insecta" family="Scarabaeidae" genus="Stigmalia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Stigmalia</taxonomicName>
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|
Casey). Future analyses should focus on comparing characters in these
|
|
<taxonomicName id="D7B04A3FD5DE7C48AAAE75D1AA5374BA" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
|
|
species-groups and genera to
|
|
<taxonomicName id="9FB9960EDFDD40B134A39C379CCBB52A" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
|
|
, rather than
|
|
<taxonomicName id="648505B3A5BB20E49355A69DDBA4F1AA" class="Insecta" family="Scarabaeidae" genus="Surutu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Surutu" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Surutu</taxonomicName>
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.
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|
</paragraph>
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|
</subSubSection>
|
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</treatment>
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</document> |