693 lines
50 KiB
XML
693 lines
50 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.96724" ID-Pensoft-Pub="1313-2970-1145-1" ID-Pensoft-UUID="ACBBD138B7375B0D9F63CE792A82F653" ID-ZooBank="830628C276CD4641BFC6144CD775ED6B" ModsDocID="1313-2970-1145-1" checkinTime="1675506069289" checkinUser="pensoft" docAuthor="Hedin, Marshal & Milne, Marc A." docDate="2023" docId="5E146B38C2B85BBBA51EB91AB2934E26" docLanguage="en" docName="ZooKeys 1145: 1-130" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.96724" docTitle="Nesticus nasicus Coyle & McGarity 1992" docType="treatment" docVersion="1" id="ACBBD138B7375B0D9F63CE792A82F653" lastPageNumber="1" masterDocId="ACBBD138B7375B0D9F63CE792A82F653" masterDocTitle="New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1675506069289" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedin, Marshal</mods:namePart>
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<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182 - 4614, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mhedin@sdsu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Milne, Marc A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1943-0161</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, University of Indianapolis, Indianapolis, Indiana 46227, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1145</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>130</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1145.96724</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.96724</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1145-1</mods:identifier>
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<mods:identifier type="ZooBank">830628C276CD4641BFC6144CD775ED6B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ACBBD138B7375B0D9F63CE792A82F653</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26" httpUri="http://treatment.plazi.org/id/5E146B38C2B85BBBA51EB91AB2934E26" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="478YH" ID-ENA="60733" LSID="5E146B38-C2B8-5BBB-A51E-B91AB2934E26" authority="Coyle & McGarity, 1992" authorityName="Coyle & McGarity" authorityYear="1992" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus nasicus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nasicus">Nesticus nasicus Coyle & McGarity, 1992</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Nesticus nasicus ♂ palps. North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen # 1155, dorsal (A), ventral (B). North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02 _ 190, dorsal (C), ventral (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure31" httpUri="https://binary.pensoft.net/fig/804482" pageId="0" pageNumber="1">Figs 31A-D</figureCitation>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Nesticus nasicus epigynal variation. North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02 _ 190, ventral (A), dorsal (B). North Carolina, Jackson Co., Coward Mountain, E of Wolfpen Gap, MCH 02 _ 177, ventral (C), dorsal (D). North Carolina, Transylvania Co., Hwy 276 at Davidson River, MCH 02 _ 192, ventral (E), dorsal (F). North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen # 1157, ventral (G), dorsal (H). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure32" httpUri="https://binary.pensoft.net/fig/804483" pageId="0" pageNumber="1">, 32A-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Coyle & McGarity" authorityYear="1992" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus nasicus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nasicus">Nesticus nasicus</taxonomicName>
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Coyle & McGarity, 1992: 162, figs 1-4, 7-14.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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Type material:
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<materialsCitation collectingDate="1990-10-28" collectingDateMax="2007-08-19" collectingDateMin="1990-10-28" collectorName="T McGarity, M. Hedin, Non, NE Mt. Pisgah, M. Lowder, P. Paquin, R. Keith, J. Starrett, S. Thomas, Blue Ridge Parkway, Richland Balsam, F. Coyle, N Pigeon Gap, Jackie Spring Gap, Coward Mountain, E, Mountain, SE, B. Dellinger, Cullowhee Mountain Road, Silverstein Road, M. McCormack, S. Derkarabetian, S Brevard, Bald Knob, S Pinhook Gap, Stillwater Branch, N Looking Glass Falls, E Owens Gap, N Fork French Broad, FR" country="USA" county="Jackson Co." latitude="35.2503" location="Cowee Mountain" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" specimenCount="71" specimenCount-female="65" specimenCount-male="6" stateProvince="North Carolina" typeStatus="Holotype">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="United States of America">USA</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="North Carolina">North Carolina</collectingRegion>
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,
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<collectingCounty>Jackson Co.</collectingCounty>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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<typeStatus>holotype</typeStatus>
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;
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<locationDeviation location="Dillsboro">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.609344" unit="mi" value="1.0">1 mile</quantity>
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W of Dillsboro
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</locationDeviation>
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at
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<collectingMunicipality>
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:A6BA023B1EBCDAEF83AC476D167CF50A" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Cowee Mountain" stateProvince="North Carolina">
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Cowee
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:350DF0854185E06D65505A74B2F19DB3" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Mountain Train Tunnel" stateProvince="North Carolina">Mountain Train Tunnel</location>
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</location>
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</collectingMunicipality>
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, rock bank;
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<collectingDate value="1990-10-28">28 Oct. 1990</collectingDate>
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;
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<collectorName>T McGarity</collectorName>
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leg. AMNH;
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:92B4CF6198CAF3BFD342FCAC2771FEDB" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="New" stateProvince="North Carolina">New</location>
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collections from type locality
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</emphasis>
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: -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:0764261FCC5E740AF1191E69CEF3BC39" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Jackson Co." stateProvince="North Carolina">Jackson Co.</location>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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,
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<specimenCount type="female">17♀</specimenCount>
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;
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:1AD05025666518B348BB111D7EE5F241" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Cowee Mountain Train Tunnel" stateProvince="North Carolina">
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:32087C209CBE615CB1FE566CA22DB99A" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Cowee Mountain" stateProvince="North Carolina">
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Cowee
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:0F9FF9227137C464EBD7F7EFAEDE35E5" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Mountain Train Tunnel" stateProvince="North Carolina">Mountain Train Tunnel</location>
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</location>
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</location>
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,
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<locationDeviation location="Dillsboro">NW of Dillsboro</locationDeviation>
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;
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<geoCoordinate degrees="35.3768" direction="north" orientation="latitude" precision="5" value="35.3768">35.3768°N</geoCoordinate>
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, -
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<geoCoordinate degrees="83.268" direction="west" orientation="longitude" precision="55" value="-83.268">83.268°W</geoCoordinate>
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;
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<collectingDate value="1992-08-14">14 Aug. 1992</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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leg.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectorName>Non</collectorName>
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type material
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</emphasis>
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: -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:BCEFAE01A3FEFDD1D334E45EFAEA1912" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Buncombe Co." stateProvince="North Carolina">Buncombe Co.</location>
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</emphasis>
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•
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<specimenCount>♀</specimenCount>
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;
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<collectorName>
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NE
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:90E5C8A86BFE93A7F5DDD431B05F2BBB" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Mt. Pisgah" stateProvince="North Carolina">Mt. Pisgah</location>
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</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:AF23FC32991730110A05D4F815FF6A38" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
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151, head of
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:7867060EAC5CD4E99556CFEC8D429D5B" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="McKinney Creek" stateProvince="North Carolina">McKinney Creek</location>
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;
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<geoCoordinate degrees="35.4448" direction="north" orientation="latitude" precision="5" value="35.4448">35.4448°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.7225" direction="west" orientation="longitude" precision="5" value="-82.7225">82.7225°W</geoCoordinate>
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;
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<collectingDate value="2002-09-05">5 Sep. 2002</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>M. Lowder</collectorName>
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,
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<collectorName>P. Paquin</collectorName>
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leg.; MCH 02_194; •
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<specimenCount>♀</specimenCount>
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;
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<collectorName>
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NE
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:FB82AB7092C32777C8A62D8B8B0A0518" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Mt. Pisgah" stateProvince="North Carolina">Mt. Pisgah</location>
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</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:8459F1D706FA268047AE1D3F6C9DB285" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
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151, head of
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:ECA3FE742F5287F73E62054D5EB68DEC" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="McKinney Creek" stateProvince="North Carolina">McKinney Creek</location>
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;
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<geoCoordinate degrees="35.4448" direction="north" orientation="latitude" precision="5" value="35.4448">35.4448°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.7225" direction="west" orientation="longitude" precision="5" value="-82.7225">82.7225°W</geoCoordinate>
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;
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<collectingDate value="2004-08-22">22 Aug. 2004</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>R. Keith</collectorName>
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,
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<collectorName>J. Starrett</collectorName>
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,
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<collectorName>S. Thomas</collectorName>
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leg.; MCH 04_074; -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:507AC9A7D4E0BEEBC7BD34A298AABCE2" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Haywood Co." stateProvince="North Carolina">Haywood Co.</location>
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</emphasis>
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•
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<specimenCount>♀</specimenCount>
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;
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<collectorName>Blue Ridge Parkway</collectorName>
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, vicinity
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<collectorName>
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Richland
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:3EDEACF29153457F44381D3DF2082B9C" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Balsam" stateProvince="North Carolina">Balsam</location>
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</collectorName>
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;
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<geoCoordinate degrees="35.3666" direction="north" orientation="latitude" precision="5" value="35.3666">35.3666°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.9915" direction="west" orientation="longitude" precision="5" value="-82.9915">82.9915°W</geoCoordinate>
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;
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<collectingDate value="1992-08-04">4 Aug. 1992</collectingDate>
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;
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<collectorName>F. Coyle</collectorName>
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leg.; •
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<specimenCount type="male">4♂</specimenCount>
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,
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<specimenCount type="female">2♀</specimenCount>
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, 7 imm;
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:6CC9C30C891F3A0CD02B2331552E63F7" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
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215, along
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:57751AF1B7F5A9C094D2699FC5A79BE7" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="West Fork Pigeon River" stateProvince="North Carolina">West Fork Pigeon River</location>
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;
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<geoCoordinate degrees="35.339" direction="north" orientation="latitude" precision="55" value="35.339">35.339°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.9016" direction="west" orientation="longitude" precision="5" value="-82.9016">82.9016°W</geoCoordinate>
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;
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<collectingDate value="2002-09-04">4 Sep. 2002</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>F. Coyle</collectorName>
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,
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<collectorName>P. Paquin</collectorName>
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leg.; MCH 02_190; •
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<specimenCount type="male">2♂</specimenCount>
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,
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<specimenCount type="female">8♀</specimenCount>
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, 5 imm;
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<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:20E6E0237DB3064D30BFFC0D9D69FC47" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
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276,
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<collectorName>N Pigeon Gap</collectorName>
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;
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<geoCoordinate degrees="35.3677" direction="north" orientation="latitude" precision="5" value="35.3677">35.3677°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.7958" direction="west" orientation="longitude" precision="5" value="-82.7958">82.7958°W</geoCoordinate>
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;
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<collectingDate value="2002-09-04">4 Sep. 2002</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>M. Lowder</collectorName>
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,
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<collectorName>P. Paquin</collectorName>
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leg.; MCH 02_193; -
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<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:47EAD373FA323F7CEEE54004482E9040" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Jackson Co." stateProvince="North Carolina">Jackson Co.</location>
|
|
</emphasis>
|
|
•
|
|
<specimenCount type="female">3♀</specimenCount>
|
|
, 1 imm;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:AB13C106DEB1CCC6F590BFD4208B14B0" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Coward Mountain" stateProvince="North Carolina">Coward Mountain</location>
|
|
near
|
|
<collectorName>Jackie Spring Gap</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.3352" direction="north" orientation="latitude" precision="5" value="35.3352">35.3352°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.0897" direction="west" orientation="longitude" precision="5" value="-83.0897">83.0897°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2002-09-01">1 Sep. 2002</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>P. Paquin</collectorName>
|
|
leg.; MCH 02_178; •
|
|
<specimenCount type="female">4♀</specimenCount>
|
|
, 1 imm;
|
|
<collectorName>
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:8C6C0970ABBA9E3700F41CEB2680FE64" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Coward Mountain" stateProvince="North Carolina">Coward Mountain</location>
|
|
, E
|
|
</collectorName>
|
|
Wolfpen Gap;
|
|
<geoCoordinate degrees="35.3606" direction="north" orientation="latitude" precision="5" value="35.3606">35.3606°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.1037" direction="west" orientation="longitude" precision="5" value="-83.1037">83.1037°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2002-09-01">1 Sep. 2002</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>M. Lowder</collectorName>
|
|
leg.; MCH 02_177; •
|
|
<specimenCount type="female">9♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:09D9D515DAB86F489F1C4D2813A484F7" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Mull Creek on Caney Fork Road" stateProvince="North Carolina">Mull Creek on Caney Fork Road</location>
|
|
,
|
|
<locationDeviation location="Hwy">
|
|
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.7702784" unit="mi" value="11.0">11 mi.</quantity>
|
|
E Hwy
|
|
</locationDeviation>
|
|
107;
|
|
<geoCoordinate degrees="35.3417" direction="north" orientation="latitude" precision="5" value="35.3417">35.3417°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.0292" direction="west" orientation="longitude" precision="5" value="-83.0292">83.0292°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="1992-08-11">11 Aug. 1992</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
leg.; •
|
|
<specimenCount type="female">2♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:B473222C32AA51683D7733FAD5214FA5" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Rich Mountain" stateProvince="North Carolina">
|
|
Rich
|
|
<collectorName>Mountain, SE</collectorName>
|
|
</location>
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:9F518F147FA1776AAA3C2D54B0278888" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Sugar Creek Gap" stateProvince="North Carolina">Sugar Creek Gap</location>
|
|
;
|
|
<geoCoordinate degrees="35.2907" direction="north" orientation="latitude" precision="5" value="35.2907">35.2907°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.004" direction="west" orientation="longitude" precision="55" value="-83.004">83.004°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2002-09-01">1 Sep. 2002</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>P. Paquin</collectorName>
|
|
leg.; MCH 02_179; •
|
|
<specimenCount>♂</specimenCount>
|
|
;
|
|
<locationDeviation location="Rich Mountain Bald">SW of Rich Mountain Bald</locationDeviation>
|
|
,
|
|
<locationDeviation location="Sugar Creek Gap">
|
|
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.04672" unit="mi" value="0.5">0.5 mi</quantity>
|
|
E Sugar Creek Gap
|
|
</locationDeviation>
|
|
;
|
|
<geoCoordinate degrees="35.2915" direction="north" orientation="latitude" precision="5" value="35.2915">35.2915°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.006" direction="west" orientation="longitude" precision="55" value="-83.006">83.006°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="1992-06-26">26 Jun. 1992</collectingDate>
|
|
;
|
|
<collectorName>B. Dellinger</collectorName>
|
|
leg.; •
|
|
<specimenCount>♀</specimenCount>
|
|
;
|
|
<locationDeviation location="Rich Mountain Bald">SW of Rich Mountain Bald</locationDeviation>
|
|
,
|
|
<locationDeviation location="Sugar Creek Gap">
|
|
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.04672" unit="mi" value="0.5">0.5 mi</quantity>
|
|
E Sugar Creek Gap
|
|
</locationDeviation>
|
|
;
|
|
<geoCoordinate degrees="35.2915" direction="north" orientation="latitude" precision="5" value="35.2915">35.2915°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.006" direction="west" orientation="longitude" precision="55" value="-83.006">83.006°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="1994-04-17">17 Apr. 1994</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>B. Dellinger</collectorName>
|
|
leg.; •
|
|
<specimenCount type="female">7♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:647D8BD814FA5617F6653DBD499399E0" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Wolf Creek" stateProvince="North Carolina">Wolf Creek</location>
|
|
at
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:3D177F13EA791182E7044FF675E0B071" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Cullowhee Creek" stateProvince="North Carolina">Cullowhee Creek</location>
|
|
, off
|
|
<collectorName>
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:5FE9638BC4BEE9F55F25360B1AB34D2D" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Cullowhee Mountain" stateProvince="North Carolina">
|
|
Cullowhee
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:67D77539B1BCEE905D8608CA877EB787" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Mountain Road" stateProvince="North Carolina">Mountain Road</location>
|
|
</location>
|
|
</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.2468" direction="north" orientation="latitude" precision="5" value="35.2468">35.2468°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.1843" direction="west" orientation="longitude" precision="5" value="-83.1843">83.1843°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="1992-08-11">11 Aug. 1992</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
leg.; -
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:8A01AD94321EA6E253DEFDF07F230148" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Transylvania Co." stateProvince="North Carolina">Transylvania Co.</location>
|
|
</emphasis>
|
|
•
|
|
<specimenCount>♀</specimenCount>
|
|
; along
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:E1831057B0D1A826BF72E285B89183C3" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="West Fork French Broad River" stateProvince="North Carolina">West Fork French Broad River</location>
|
|
,
|
|
<collectorName>Silverstein Road</collectorName>
|
|
,
|
|
<locationDeviation location="Hwy">
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.218688" unit="mi" value="2.0">2 mi.</quantity>
|
|
N Hwy
|
|
</locationDeviation>
|
|
64;
|
|
<geoCoordinate degrees="35.1573" direction="north" orientation="latitude" precision="5" value="35.1573">35.1573°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.8758" direction="west" orientation="longitude" precision="5" value="-82.8758">82.8758°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2007-08-19">19 Aug. 2007</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>M. McCormack</collectorName>
|
|
,
|
|
<collectorName>S. Derkarabetian</collectorName>
|
|
leg.; MCH 07_128; •
|
|
<specimenCount>♀</specimenCount>
|
|
; below
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:0D1C23166A1825E905311437E0AE4DA5" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Connestee Falls" stateProvince="North Carolina">Connestee Falls</location>
|
|
, off
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:C9315B549BD5813BA5A319BE241EDFDA" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
276
|
|
<collectorName>S Brevard</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.1647" direction="north" orientation="latitude" precision="5" value="35.1647">35.1647°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.7319" direction="west" orientation="longitude" precision="5" value="-82.7319">82.7319°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="1992-10-02">2 Oct. 1992</collectingDate>
|
|
;
|
|
<collectorName>B. Dellinger</collectorName>
|
|
leg.; •
|
|
<specimenCount>♂</specimenCount>
|
|
,
|
|
<specimenCount type="female">2♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:16F0D7FEC909A4B5E752D77CFBB6D73A" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
215,
|
|
<locationDeviation location="Balsam Grove">
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.5639168" unit="mi" value="4.7">4.7 mi.</quantity>
|
|
NW Balsam Grove
|
|
</locationDeviation>
|
|
along
|
|
<collectorName>Bald Knob</collectorName>
|
|
branch;
|
|
<geoCoordinate degrees="35.2568" direction="north" orientation="latitude" precision="5" value="35.2568">35.2568°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.9098" direction="west" orientation="longitude" precision="5" value="-82.9098">82.9098°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="1992-08-13">13 Aug. 1992</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
leg.; •
|
|
<specimenCount type="female">5♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:DF945C6B029B9EF7188AD2F762A6B7FA" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
215,
|
|
<collectorName>S Pinhook Gap</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.2575" direction="north" orientation="latitude" precision="5" value="35.2575">35.2575°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.9204" direction="west" orientation="longitude" precision="5" value="-82.9204">82.9204°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2004-08-22">22 Aug. 2004</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>R. Keith</collectorName>
|
|
,
|
|
<collectorName>J. Starrett</collectorName>
|
|
,
|
|
<collectorName>S. Thomas</collectorName>
|
|
leg.; MCH 04_073; •
|
|
<specimenCount type="female">2♀</specimenCount>
|
|
, 1 imm;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:D8FC3C5087F3686F838804DDC1A5FF04" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
276 at
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:BF21B943A9FB1474227F5EADB6BBE4CA" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Davidson River" stateProvince="North Carolina">Davidson River</location>
|
|
, opposite
|
|
<collectorName>Stillwater Branch</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.284" direction="north" orientation="latitude" precision="55" value="35.284">35.284°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.7591" direction="west" orientation="longitude" precision="5" value="-82.7591">82.7591°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2001-08-28">28 Aug. 2001</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>M. Lowder</collectorName>
|
|
,
|
|
<collectorName>P. Paquin</collectorName>
|
|
leg.; MCH 01_180; •
|
|
<specimenCount type="female">4♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:BC203BC0AB8BD91F0B0D66377561DC33" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
276 at
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:16D6D2CD625F5C6B6AF81E88C718CF98" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Davidson River" stateProvince="North Carolina">Davidson River</location>
|
|
, opposite
|
|
<collectorName>Stillwater Branch</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.284" direction="north" orientation="latitude" precision="55" value="35.284">35.284°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.7591" direction="west" orientation="longitude" precision="5" value="-82.7591">82.7591°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2002-09-04">4 Sep. 2002</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>M. Lowder</collectorName>
|
|
,
|
|
<collectorName>P. Paquin</collectorName>
|
|
leg.; MCH 02_192; •
|
|
<specimenCount>♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:F6557EA49C0C9071CA5BE2FDB8C26D1B" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
276,
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:36A0FC0BC29D1BE0199DDE92BD3A7AB5" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Looking Glass Creek" stateProvince="North Carolina">Looking Glass Creek</location>
|
|
just
|
|
<collectorName>
|
|
N
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:F1840F3DECCCD6FADBCF669D821329C7" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Looking Glass Falls" stateProvince="North Carolina">Looking Glass Falls</location>
|
|
</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.2978" direction="north" orientation="latitude" precision="5" value="35.2978">35.2978°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.7676" direction="west" orientation="longitude" precision="5" value="-82.7676">82.7676°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2007-08-19">19 Aug. 2007</collectingDate>
|
|
;
|
|
<collectorName>M. McCormack</collectorName>
|
|
,
|
|
<collectorName>S. Derkarabetian</collectorName>
|
|
leg.; MCH 07_129; •
|
|
<specimenCount>♀</specimenCount>
|
|
;
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:7C6CAE936E0348BA804CB590DF7BB910" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
281,
|
|
<collectorName>E Owens Gap</collectorName>
|
|
;
|
|
<geoCoordinate degrees="35.1957" direction="north" orientation="latitude" precision="5" value="35.1957">35.1957°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.9608" direction="west" orientation="longitude" precision="5" value="-82.9608">82.9608°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2004-08-20">20 Aug. 2004</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>R. Keith</collectorName>
|
|
,
|
|
<collectorName>J. Starrett</collectorName>
|
|
,
|
|
<collectorName>S. Thomas</collectorName>
|
|
leg.; MCH 04_071; •
|
|
<specimenCount>♀</specimenCount>
|
|
, 1 imm;
|
|
<collectorName>N Fork French Broad, FR</collectorName>
|
|
140 off
|
|
<location LSID="urn:lsid:plazi:treatment:5E146B38C2B85BBBA51EB91AB2934E26:328DE6481A924AECB8C14F7E3090634F" country="USA" county="Jackson Co." latitude="35.2503" longLatPrecision="45" longitude="-82.889" municipality="Cowee Mountain Train Tunnel" name="Hwy" stateProvince="North Carolina">Hwy</location>
|
|
215;
|
|
<geoCoordinate degrees="35.2503" direction="north" orientation="latitude" precision="5" value="35.2503">35.2503°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="82.889" direction="west" orientation="longitude" precision="55" value="-82.889">82.889°W</geoCoordinate>
|
|
;
|
|
<collectingDate value="2002-09-04">4 Sep. 2002</collectingDate>
|
|
;
|
|
<collectorName>M. Hedin</collectorName>
|
|
,
|
|
<collectorName>M. Lowder</collectorName>
|
|
,
|
|
<collectorName>P. Paquin</collectorName>
|
|
leg.; MCH 02_189.
|
|
</materialsCitation>
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Males may be distinguished from other members of the species group by the palp with a uniquely shaped tegular apophysis (except in comparison to
|
|
<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus brimleyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="brimleyi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus brimleyi</emphasis>
|
|
</taxonomicName>
|
|
), combined with a narrow-based paracymbial dorsal process (Fig.
|
|
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Nesticus nasicus ♂ palps. North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen # 1155, dorsal (A), ventral (B). North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02 _ 190, dorsal (C), ventral (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure31" httpUri="https://binary.pensoft.net/fig/804482" pageId="0" pageNumber="1">31A, C</figureCitation>
|
|
) which differs greatly from
|
|
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. brimleyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="brimleyi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N. brimleyi</emphasis>
|
|
</taxonomicName>
|
|
. The
|
|
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. nasicus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nasicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N. nasicus</emphasis>
|
|
</taxonomicName>
|
|
epigynum is very similar to
|
|
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. brimleyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="brimleyi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N. brimleyi</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. templetoni" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="templetoni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N. templetoni</emphasis>
|
|
</taxonomicName>
|
|
. When viewed dorsally all possess
|
|
<normalizedToken originalValue="“crinkled”">"crinkled"</normalizedToken>
|
|
sac-shaped structures above (anterior to) the main epigynal plate, which we hypothesize are homologous to vulval pockets (Vp) as seen in Japanese
|
|
<taxonomicName authorityName="Thorell" authorityYear="1869" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation DOI="https://doi.org/10.3897/subtbiol.35.53933" author="Suzuki, Y" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="1" pagination="79 - 96" refId="B59" refString="Suzuki, Y, Ballarin, F, 2020. Nesticus kosodensis Yaginuma, 1972 bona species. Molecular and morphological separation from N. latiscapus Yaginuma, 1972 with notes on cave scaffold-web spiders subspecies in Japan (Araneae, Nesticidae). Subterranean Biology 35: 79 - 96, DOI: https://doi.org/10.3897/subtbiol.35.53933" title="Nesticus kosodensis Yaginuma, 1972 bona species. Molecular and morphological separation from N. latiscapus Yaginuma, 1972 with notes on cave scaffold-web spiders subspecies in Japan (Araneae, Nesticidae)." url="https://doi.org/10.3897/subtbiol.35.53933" volume="35" year="2020">Suzuki and Ballarin 2020</bibRefCitation>
|
|
). Further diagnostic features for
|
|
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. nasicus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nasicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N. nasicus</emphasis>
|
|
</taxonomicName>
|
|
are discussed in
|
|
<bibRefCitation author="Coyle, FA" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="1" pagination="161 - 168" publicationUrl="https://www.jstor.org/stable/3705889" refId="B8" refString="Coyle, FA, McGarity, AC, 1992. Two new species of Nesticus spiders from the southern Appalachians (Araneae, Nesticidae). The Journal of Arachnology 19: 161 - 168, https://www.jstor.org/stable/3705889" title="Two new species of Nesticus spiders from the southern Appalachians (Araneae, Nesticidae)." url="https://www.jstor.org/stable/3705889" volume="19" year="1992">Coyle and McGarity (1992)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1145.96724.figure31" httpUri="https://binary.pensoft.net/fig/804482" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
|
|
<taxonomicName authorityName="Coyle & McGarity" authorityYear="1992" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus nasicus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nasicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus nasicus</emphasis>
|
|
</taxonomicName>
|
|
♂ palps. North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen #1155, dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
). North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02_190, dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
). Scale bar: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="variation">
|
|
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Notable variation exists in the shape of the dorsal process of the paracymbium, which is sometimes narrow and finger-like (
|
|
<bibRefCitation author="Coyle, FA" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="1" pagination="161 - 168" publicationUrl="https://www.jstor.org/stable/3705889" refId="B8" refString="Coyle, FA, McGarity, AC, 1992. Two new species of Nesticus spiders from the southern Appalachians (Araneae, Nesticidae). The Journal of Arachnology 19: 161 - 168, https://www.jstor.org/stable/3705889" title="Two new species of Nesticus spiders from the southern Appalachians (Araneae, Nesticidae)." url="https://www.jstor.org/stable/3705889" volume="19" year="1992">Coyle and McGarity 1992</bibRefCitation>
|
|
: figs 1, 2), or fishtailed (Fig.
|
|
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Nesticus nasicus ♂ palps. North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen # 1155, dorsal (A), ventral (B). North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02 _ 190, dorsal (C), ventral (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure31" httpUri="https://binary.pensoft.net/fig/804482" pageId="0" pageNumber="1">31A-D</figureCitation>
|
|
), with variation in the shape of the end of the process. Both the distomedial and dorsomedial paracymbial processes vary in presence across populations, with a distomedial process found in West Fork Pigeon River and Cowee Mountain males (
|
|
<bibRefCitation author="Coyle, FA" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="1" pagination="161 - 168" publicationUrl="https://www.jstor.org/stable/3705889" refId="B8" refString="Coyle, FA, McGarity, AC, 1992. Two new species of Nesticus spiders from the southern Appalachians (Araneae, Nesticidae). The Journal of Arachnology 19: 161 - 168, https://www.jstor.org/stable/3705889" title="Two new species of Nesticus spiders from the southern Appalachians (Araneae, Nesticidae)." url="https://www.jstor.org/stable/3705889" volume="19" year="1992">Coyle and McGarity 1992</bibRefCitation>
|
|
: fig. 2), and a dorsomedial process found only in Cowee Mountain males; these processes are lacking in males from other populations. As discussed below similar population-level variation in these processes is observed in
|
|
<taxonomicName authorityName="Hedin & Milne" authorityYear="2023" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus templetoni" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="templetoni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus templetoni</emphasis>
|
|
</taxonomicName>
|
|
. The shape of both the lateral and apical processes of the median apophysis also varies across populations (Fig.
|
|
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Nesticus nasicus ♂ palps. North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen # 1155, dorsal (A), ventral (B). North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02 _ 190, dorsal (C), ventral (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure31" httpUri="https://binary.pensoft.net/fig/804482" pageId="0" pageNumber="1">31A-D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Epigyna vary across sample locations in the length of the projection of the median septum, the shape of the epigynal pockets (though generally spherical), the width of epigynal pocket lateral hoods, and the length of the spermathecae (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Nesticus nasicus epigynal variation. North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02 _ 190, ventral (A), dorsal (B). North Carolina, Jackson Co., Coward Mountain, E of Wolfpen Gap, MCH 02 _ 177, ventral (C), dorsal (D). North Carolina, Transylvania Co., Hwy 276 at Davidson River, MCH 02 _ 192, ventral (E), dorsal (F). North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen # 1157, ventral (G), dorsal (H). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure32" httpUri="https://binary.pensoft.net/fig/804483" pageId="0" pageNumber="1">32A-H</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1145.96724.figure32" httpUri="https://binary.pensoft.net/fig/804483" pageId="0" pageNumber="1" start="Figure 32" startId="F32">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 32.</emphasis>
|
|
<taxonomicName authorityName="Coyle & McGarity" authorityYear="1992" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus nasicus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nasicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus nasicus</emphasis>
|
|
</taxonomicName>
|
|
epigynal variation. North Carolina, Haywood Co., along West Fork Pigeon River, MCH 02_190, ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
). North Carolina, Jackson Co., Coward Mountain, E of Wolfpen Gap, MCH 02_177, ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
). North Carolina, Transylvania Co., Hwy 276 at Davidson River, MCH 02_192, ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
), dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
). North Carolina, Transylvania Co., Hwy 215, NW of Balsam Grove, MCH specimen #1157, ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
|
), dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
|
). Scale bar: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph pageId="0" pageNumber="1">Distribution and natural history.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Previously known only from two locations but now known to be reasonably widespread in the Great Balsam and Pisgah Mountains southwest of Asheville North Carolina, west of the Asheville Basin (Fig.
|
|
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Distribution of Nesticus nasicus group species. Type localities designated with yellow circles. Primary cities shown for geographic context. Dashed lines circumscribe known species distributions; possible distribution of Nesticus secretus is tentative, see text for details." figureDoi="10.3897/zookeys.1145.96724.figure30" httpUri="https://binary.pensoft.net/fig/804481" pageId="0" pageNumber="1">30</figureCitation>
|
|
). The southeastern Connestee Falls population, found east of the French Broad River, is a geographic outlier.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
No obvious phylogeographic trends are apparent in the mitochondrial data, with geographically separate locations seemingly less genetically divergent than in other similarly widespread taxa (Fig.
|
|
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. COI IQT gene tree. Distant outgroups trimmed. Specimen numbers correspond to those in Suppl. material 3 (with detailed location provided in Suppl. material 1). Sequences from UCE capture denoted with " NODE ". Populations with a " Nesticus cooperi - like " male paracymbium denoted by small circles. Not all tip bootstrap values shown (see Suppl. material 4 for. tre file with all bootstrap values)." figureDoi="10.3897/zookeys.1145.96724.figure6" httpUri="https://binary.pensoft.net/fig/804457" pageId="0" pageNumber="1">6</figureCitation>
|
|
). This is particularly striking considering the notable male morphological variation observed across populations.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |