treatments-xml/data/46/20/6C/46206C74FFD01D44FF78FF4FFB8DFE5A.xml
2024-06-21 12:35:10 +02:00

479 lines
57 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="36DF4254705379989E66A966525AC173" ID="10.11646/zootaxa.4132.4.9" ID-DOI="10.11646/zootaxa.4132.4.9" ID-GBIF-Dataset="9c2ca859-fb7a-46b9-b78e-e471eed708c2" ID-ISSN="1175-5326" ID-Zenodo-Dep="271908" ID-ZooBank="5F9B951D-26EB-4CA7-B4FC-F89526081CAD" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467372578577" checkinUser="plazi" docAuthor="Pilato, Giovanni, Sabella, Giorgio &amp; Lisi, Oscar" docDate="2016" docId="46206C74FFD01D44FF78FF4FFB8DFE5A" docLanguage="en" docName="zootaxa.4132.4.9.pdf" docOrigin="Zootaxa 4132 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Milnesium vorax Pilato, Sabella &amp; Lisi, 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="579" masterDocId="BA19140CFFD11D40FFEFFF93FFCEFFAE" masterDocTitle="Two new species of Milnesium (Tardigrada: Milnesiidae)" masterLastPageNumber="587" masterPageNumber="575" pageNumber="576" updateTime="1698363095221" updateUser="plazi">
<mods:mods id="A22762747EA32F487B8B141AD44ECF2D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="798641AE3019AA445ED0612B3A043B0E">
<mods:title id="870B4612B2A775C3111FC94837C8EAE4">Two new species of Milnesium (Tardigrada: Milnesiidae)</mods:title>
</mods:titleInfo>
<mods:name id="B97E09C66BB28F08621B45F664FBF2E5" type="personal">
<mods:role id="2D5E666CC3C9833FC646B51525FF8493">
<mods:roleTerm id="2ED07C0A94C937980872FD693E8ADCAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65E93E03A7D7FB4A8B20D796EFE86176">Pilato, Giovanni</mods:namePart>
</mods:name>
<mods:name id="9CCF9904C0311CCCC6F44BAFA6C93C9D" type="personal">
<mods:role id="4470F11F871C61B87C8C239847990675">
<mods:roleTerm id="F9BEAB832C876F2EC1568AE35B7C6A59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2BEB38AB72166A2A847D4E5BB61065E">Sabella, Giorgio</mods:namePart>
</mods:name>
<mods:name id="27C66EFDCDF2B96DC883EA21D9D12AB9" type="personal">
<mods:role id="E2C861DA3CB77B914D582E5977638779">
<mods:roleTerm id="1B81D5F693A04C7C8C1CC8D461094974">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECE1373C0A2142CB55D618B0D295A18B">Lisi, Oscar</mods:namePart>
</mods:name>
<mods:typeOfResource id="FD1A79703BF0264A5BBEC74B3B1F33AF">text</mods:typeOfResource>
<mods:relatedItem id="2708C8BB6E7CA9BD06FA5B1E6C447533" type="host">
<mods:titleInfo id="1103FA54CC53442715256593B057946F">
<mods:title id="423FAF196B5CBF99248C66ACDEFC1DAF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="753BDDEC01DD91CC1C2449924ACB33F3">
<mods:date id="7B028820E598990404A6ED524DEFDE07">2016</mods:date>
<mods:detail id="0029AE554F94E600796885684EE69AEB" type="volume">
<mods:number id="E9CE387DE64ED1F950BFBDDC51CB2A9A">4132</mods:number>
</mods:detail>
<mods:detail id="CE0F67FE3F67BDED7D1A77138DD93A9C" type="issue">
<mods:number id="40B5971D61157EC571199D920A08700C">4</mods:number>
</mods:detail>
<mods:extent id="4A0A7D284ADFA80A824E1A5AF6FC1ADE" unit="page">
<mods:start id="6311EB10346E9D2FDF76189F57A3B8A0">575</mods:start>
<mods:end id="BFAAF86780858FB9B047E5F58D2C4F06">587</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8443E4ED595D56BCDE2C8242865490EC">journal article</mods:classification>
<mods:identifier id="C224872A8AFA5A7AD68F8D03A513B2B0" type="DOI">10.11646/zootaxa.4132.4.9</mods:identifier>
<mods:identifier id="BA8AFCBA773DA51C50839DC1E935CCFB" type="GBIF-Dataset">9c2ca859-fb7a-46b9-b78e-e471eed708c2</mods:identifier>
<mods:identifier id="F6087548EEC287CEC0E83DBB916DAF6F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="32EC5D6050C829BDB170A91F2FDE0EC9" type="Zenodo-Dep">271908</mods:identifier>
<mods:identifier id="0A9EE29193C4C4B8A0083665223C0C60" type="ZooBank">5F9B951D-26EB-4CA7-B4FC-F89526081CAD</mods:identifier>
</mods:mods>
<treatment id="46206C74FFD01D44FF78FF4FFB8DFE5A" ID-DOI="http://doi.org/10.5281/zenodo.5611931" ID-GBIF-Taxon="121268799" ID-Zenodo-Dep="5611931" LSID="urn:lsid:plazi:treatment:46206C74FFD01D44FF78FF4FFB8DFE5A" httpUri="http://treatment.plazi.org/id/46206C74FFD01D44FF78FF4FFB8DFE5A" lastPageId="4" lastPageNumber="579" pageId="1" pageNumber="576">
<subSubSection id="86938EE9FFD01D41FF78FF4FFF16FEB6" pageId="1" pageNumber="576" type="nomenclature">
<paragraph id="CE36DD62FFD01D41FF78FF4FFE07FF59" blockId="1.[151,457,220,280]" box="[151,457,220,247]" pageId="1" pageNumber="576">
<heading id="957E6A0EFFD01D41FF78FF4FFE07FF59" bold="true" box="[151,457,220,247]" fontSize="11" level="1" pageId="1" pageNumber="576" reason="1">
<emphasis id="FCFD0170FFD01D41FF78FF4FFE07FF59" bold="true" box="[151,457,220,247]" pageId="1" pageNumber="576">
<taxonomicName id="0989A6E1FFD01D41FF78FF4FFEABFF58" box="[151,357,220,246]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="576" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD01D41FF78FF4FFEABFF58" bold="true" box="[151,357,220,246]" italics="true" pageId="1" pageNumber="576">Milnesium vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD01D41FE83FF4EFE07FF59" box="[364,457,221,247]" pageId="1" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CE36DD62FFD01D41FF78FF6CFF16FEB6" blockId="1.[151,457,220,280]" box="[151,216,255,280]" pageId="1" pageNumber="576">
<figureCitation id="56B2C1E7FFD01D41FF78FF6CFF16FEB6" box="[151,216,255,280]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="1" pageNumber="576">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="86938EE9FFD01D41FF78FED4FDE2FD96" pageId="1" pageNumber="576" type="materials_examined">
<paragraph id="CE36DD62FFD01D41FF78FED4FE2DFE2B" blockId="1.[151,1436,327,568]" pageId="1" pageNumber="576">
<emphasis id="FCFD0170FFD01D41FF78FED4FEF9FECE" bold="true" box="[151,311,327,352]" pageId="1" pageNumber="576">
<typeStatus id="113263C0FFD01D41FF78FED4FF1CFECE" box="[151,210,327,352]" pageId="1" pageNumber="576">Type</typeStatus>
locality.
</emphasis>
Sicily, neighbourhood of Ramacca (Catania),
<geoCoordinate id="ABBDBBA5FFD01D41FCA6FED4FC1CFECE" box="[841,978,327,352]" direction="north" orientation="latitude" pageId="1" pageNumber="576" precision="15" value="37.383335">37°2300″N</geoCoordinate>
,
<geoCoordinate id="ABBDBBA5FFD01D41FC30FED4FBADFECE" box="[991,1123,327,352]" direction="east" orientation="longitude" pageId="1" pageNumber="576" precision="15" value="14.7">14°4200″E</geoCoordinate>
,
<quantity id="09717087FFD01D41FB9EFEDBFB74FEF1" box="[1137,1210,328,352]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="576" unit="m" value="270.0">270 m</quantity>
a.s.l. (
<typeStatus id="113263C0FFD01D41FAE8FED4FAA5FECE" box="[1287,1387,327,352]" pageId="1" pageNumber="576" type="holotype">holotype</typeStatus>
and one
<typeStatus id="113263C0FFD01D41FF29FEFEFEE4FE2B" box="[198,298,365,389]" pageId="1" pageNumber="576" type="paratype">paratype</typeStatus>
, slide no. 2775).
</paragraph>
<paragraph id="CE36DD62FFD01D41FF28FE1CFD8EFE62" blockId="1.[151,1436,327,568]" pageId="1" pageNumber="576">
<emphasis id="FCFD0170FFD01D41FF28FE1CFE7BFE06" bold="true" box="[199,437,399,424]" pageId="1" pageNumber="576">Material examined.</emphasis>
<typeStatus id="113263C0FFD01D41FE2EFE03FE37FE06" box="[449,505,400,424]" pageId="1" pageNumber="576">Type</typeStatus>
material from a moss on soil sample collected by our colleague Vera DUrso (University of Catania) in
<date id="BA37FBA2FFD01D41FE51FE27FDF5FE62" box="[446,571,436,461]" pageId="1" pageNumber="576" value="1998-04">April 1998</date>
.
</paragraph>
<paragraph id="CE36DD62FFD01D41FF28FE44FDE2FD96" blockId="1.[151,1436,327,568]" pageId="1" pageNumber="576">
<emphasis id="FCFD0170FFD01D41FF28FE44FE45FE5E" bold="true" box="[199,395,471,496]" pageId="1" pageNumber="576">
<typeStatus id="113263C0FFD01D41FF28FE44FECCFE5E" box="[199,258,471,496]" pageId="1" pageNumber="576">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="113263C0FFD01D41FE7AFE44FE33FE5E" box="[405,509,471,496]" pageId="1" pageNumber="576" type="holotype">Holotype</typeStatus>
and
<typeStatus id="113263C0FFD01D41FDD5FE4BFD54FE5E" box="[570,666,472,496]" pageId="1" pageNumber="576" type="paratype">paratype</typeStatus>
are deposited in the collection of Binda &amp; Pilato, Museum of the Department of Biological, Geological and Environmental Sciences, Section of Animal Biology “Marcello La Greca”, University of Catania,
<collectingCountry id="B69E9DF2FFD01D41FE1CFD8CFDE6FD96" box="[499,552,543,568]" name="Italy" pageId="1" pageNumber="576">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="9AF68DEAFFD01D41FF78F8A3FCDBF82D" httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="1" pageNumber="576" targetBox="[213,1369,629,1808]" targetPageId="1">
<paragraph id="CE36DD62FFD01D41FF78F8A3FCDBF82D" blockId="1.[151,1436,1840,1923]" pageId="1" pageNumber="576">
<emphasis id="FCFD0170FFD01D41FF78F8A3FE84F8EB" bold="true" box="[151,330,1840,1862]" pageId="1" pageNumber="576">FIGURE 1. AD</emphasis>
,
<taxonomicName id="0989A6E1FFD01D41FEB8F8A3FDCCF8E8" box="[343,514,1840,1862]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="576" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD01D41FEB8F8A3FDCCF8E8" box="[343,514,1840,1862]" italics="true" pageId="1" pageNumber="576">Milnesium vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD01D41FDE7F8A3FD97F8EB" box="[520,601,1840,1861]" pageId="1" pageNumber="576" rank="species">
<emphasis id="FCFD0170FFD01D41FDE7F8A3FD97F8EB" bold="true" box="[520,601,1840,1861]" pageId="1" pageNumber="576">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype).
<emphasis id="FCFD0170FFD01D41FD3AF8A3FD29F8EB" bold="true" box="[725,743,1840,1861]" pageId="1" pageNumber="576">A</emphasis>
, Buccal tube.
<emphasis id="FCFD0170FFD01D41FC96F8A3FC44F8EB" bold="true" box="[889,906,1840,1861]" pageId="1" pageNumber="576">B</emphasis>
, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening).
<emphasis id="FCFD0170FFD01D41FDBEF8DCFDADF8CB" bold="true" box="[593,611,1871,1893]" pageId="1" pageNumber="576">C</emphasis>
, Claws of legs III (the arrow indicates the long cuticular thickening).
<emphasis id="FCFD0170FFD01D41FACCF8DCFAFBF8CA" bold="true" box="[1315,1333,1871,1892]" pageId="1" pageNumber="576">D</emphasis>
, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm.
</paragraph>
</caption>
<subSubSection id="86938EE9FFD31D42FF28FF04FE2FFE91" pageId="2" pageNumber="577" type="diagnosis">
<paragraph id="CE36DD62FFD31D42FF28FF04FE2FFE91" blockId="2.[151,1436,151,1077]" pageId="2" pageNumber="577">
<emphasis id="FCFD0170FFD31D42FF28FF04FE68FF1E" bold="true" box="[199,422,151,176]" pageId="2" pageNumber="577">Specific diagnosis.</emphasis>
Colourless; cuticle smooth; eye spots present; six peribuccal and two lateral papillae present; mouth terminal with six triangular peribuccal lamellae with basal stripes; buccal tube wide; stylet supports inserted on the buccal tube at nearly 62 % of its length; claws of the
<taxonomicName id="0989A6E1FFD31D42FC7EFF73FBC9FF56" box="[913,1031,224,248]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="genus">
<emphasis id="FCFD0170FFD31D42FC7EFF73FBC9FF56" box="[913,1031,224,248]" italics="true" pageId="2" pageNumber="577">Milnesium</emphasis>
</taxonomicName>
<typeStatus id="113263C0FFD31D42FBE1FF73FBF0FF56" box="[1038,1086,224,248]" pageId="2" pageNumber="577">type</typeStatus>
with configuration [2-3]-[3-2]; main branches with thin accessory points; secondary branches with rounded basal thickenings; a long cuticular bar present under the claws IIII.
</paragraph>
</subSubSection>
<subSubSection id="86938EE9FFD31D42FF28FEDFFEB4FBBE" pageId="2" pageNumber="577" type="description">
<paragraph id="CE36DD62FFD31D42FF28FEDFFC01FC42" blockId="2.[151,1436,151,1077]" pageId="2" pageNumber="577">
<emphasis id="FCFD0170FFD31D42FF28FEDFFDDBFECB" bold="true" box="[199,533,332,357]" pageId="2" pageNumber="577">
Description of the
<typeStatus id="113263C0FFD31D42FE48FEDFFDDEFECB" box="[423,528,332,357]" pageId="2" pageNumber="577" type="holotype">holotype</typeStatus>
.
</emphasis>
Body uncoloured 585 µm long; cuticle smooth without granulation or reticular design, and without pseudopores; eye spots present. Six peribuccal and two lateral papillae present. Buccopharyngeal apparatus of the
<taxonomicName id="0989A6E1FFD31D42FE38FE07FD80FE02" box="[471,590,404,428]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="genus">
<emphasis id="FCFD0170FFD31D42FE38FE07FD80FE02" box="[471,590,404,428]" italics="true" pageId="2" pageNumber="577">Milnesium</emphasis>
</taxonomicName>
<typeStatus id="113263C0FFD31D42FDBAFE06FD4BFE03" box="[597,645,405,429]" pageId="2" pageNumber="577">type</typeStatus>
(rigid buccal tube without ventral lamina, apophyses for the insertion of the stylet muscles in the shape of very short and flat ridges symmetrical with respect to the frontal plane and without caudal processes; pharyngeal bulb elongated, piriform and without apophyses, placoids or septulum); six triangular peribuccal lamellae present with basal stripes. Stylet furcae wide and triangular in shape. The buccal tube (
<figureCitation id="56B2C1E7FFD31D42FF39FDB7FEE8FD93" box="[214,294,548,573]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="2" pageNumber="577">Fig.1A</figureCitation>
) is cylindrical, 35.0 µm long (measured according to
<bibRefCitation id="AA18A093FFD31D42FC94FDB6FBE2FD93" author="Tumanov" box="[891,1068,548,573]" pageId="2" pageNumber="577" refString="Tumanov, D. V. (2006) Five new species of the genus Milnesium (Tardigrada, Eutardigrada, Milnesiidae). Zootaxa, 1122, 1 - 23." type="journal article" year="2006">Tumanov, 2006</bibRefCitation>
, i.e. including the caudal flexible portion); the external width at the level of the stylet insertion point is 21.1 µm (
<emphasis id="FCFD0170FFD31D42FBC5FDDAFB8EFDCE" box="[1066,1088,585,608]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 60.3). Stylet supports short, inserted on the buccal tube at 62.2% of its length (
<emphasis id="FCFD0170FFD31D42FD1AFDFEFCC5FD2A" box="[757,779,621,644]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 62.2). Claws of the
<taxonomicName id="0989A6E1FFD31D42FBF0FDFFFB5BFD2A" box="[1055,1173,620,644]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="genus">
<emphasis id="FCFD0170FFD31D42FBF0FDFFFB5BFD2A" box="[1055,1173,620,644]" italics="true" pageId="2" pageNumber="577">Milnesium</emphasis>
</taxonomicName>
<typeStatus id="113263C0FFD31D42FB4EFDFEFB1FFD2B" box="[1185,1233,621,645]" pageId="2" pageNumber="577">type</typeStatus>
(
<figureCitation id="56B2C1E7FFD31D42FB0AFDFFFA97FD2B" box="[1253,1369,620,645]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="2" pageNumber="577">Fig.1BD</figureCitation>
). The external claws of the first three pairs of legs have two points, the internal claws three points, on leg pair four the anterior claws with two points and posterior claws with three points, (configuration [2-3]-[3-2] according to the code suggested by
<bibRefCitation id="AA18A093FFD31D42FE6AFD44FD6BFD5E" author="Michalczyk" box="[389,677,727,752]" pageId="2" pageNumber="577" refString="Michalczyk, L., Welnicz, W., Frohme, M. &amp; Kaczmarek, L. (2012) Redescription of three Milnesium Doyere, 1840 taxa (Tardigrada: Eutardigrada: Milnesiidae), including the nominal species for the genus. Zootaxa, 3154, 1 - 20." type="journal article" year="2012">
Michalczyk
<emphasis id="FCFD0170FFD31D42FDF7FD4AFD96FD5E" box="[536,600,728,752]" italics="true" pageId="2" pageNumber="577">et al.</emphasis>
2012
</bibRefCitation>
). External main branch of legs I, 16.3 µm long (
<emphasis id="FCFD0170FFD31D42FAFAFD4AFAE5FD5E" box="[1301,1323,729,752]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 46.6); base+secondary branch of leg I, 12.5 µm long (
<emphasis id="FCFD0170FFD31D42FD41FD6EFD0AFCBA" box="[686,708,765,788]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 35.7); the base+secondary branch length is 76.7% of the main branch length. External main branch on legs II 17.4 µm (
<emphasis id="FCFD0170FFD31D42FCCFFCB2FCF3FC96" box="[800,829,801,824]" italics="true" pageId="2" pageNumber="577">pt.</emphasis>
= 19.7). External main branches on legs III, 18.1 µm long (
<emphasis id="FCFD0170FFD31D42FF35FCD6FF21FCF2" box="[218,239,837,860]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 51.7) and the base+secondary branch, 13.5 µm (
<emphasis id="FCFD0170FFD31D42FCDDFCD6FC86FCF2" box="[818,840,837,860]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 38.6); the base+secondary branch length is 74.6% of the main branch length. Posterior main branch on legs IV, 20.8 µm long (
<emphasis id="FCFD0170FFD31D42FC12FCFAFBDDFC2E" box="[1021,1043,873,896]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 59.4), posterior base+secondary branch, 14.9 µm (
<emphasis id="FCFD0170FFD31D42FEB0FC1EFEBAFC0A" box="[351,372,909,932]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
= 42.6); the base+secondary branch length is 71.6% of the main branch length. Main branches with thin accessory points; claw bases with rounded basal thickenings (
<figureCitation id="56B2C1E7FFD31D42FC37FC3CFBFDFC66" box="[984,1075,943,968]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="2" pageNumber="577">Fig. 1B</figureCitation>
. arrow a, and
<figureCitation id="56B2C1E7FFD31D42FB30FC3CFAF7FC66" box="[1247,1337,943,968]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="2" pageNumber="577">Fig. 1D</figureCitation>
); a long cuticular bar present under the claws IIII (
<figureCitation id="56B2C1E7FFD31D42FD93FC47FD05FC43" box="[636,715,980,1005]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="2" pageNumber="577">Fig.1B</figureCitation>
, arrow b, and
<figureCitation id="56B2C1E7FFD31D42FC83FC47FC0CFC42" box="[876,962,980,1005]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="2" pageNumber="577">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="CE36DD62FFD31D42FF28FC6BFEB4FBBE" blockId="2.[151,1436,151,1077]" box="[199,378,1015,1040]" pageId="2" pageNumber="577">Eggs not found.</paragraph>
</subSubSection>
<subSubSection id="86938EE9FFD31D42FF28FB8FFAA5FB9A" box="[199,1387,1052,1077]" pageId="2" pageNumber="577" type="discussion">
<paragraph id="CE36DD62FFD31D42FF28FB8FFAA5FB9A" blockId="2.[151,1436,151,1077]" box="[199,1387,1052,1077]" pageId="2" pageNumber="577">
<emphasis id="FCFD0170FFD31D42FF28FB8FFEF5FB9B" bold="true" box="[199,315,1052,1077]" pageId="2" pageNumber="577">Remarks.</emphasis>
The
<typeStatus id="113263C0FFD31D42FE9BFB8EFE1AFB9B" box="[372,468,1053,1077]" pageId="2" pageNumber="577" type="paratype">paratype</typeStatus>
is similar to the
<typeStatus id="113263C0FFD31D42FD7FFB8FFD3CFB9B" box="[656,754,1052,1077]" pageId="2" pageNumber="577" type="holotype">holotype</typeStatus>
in both qualitative and quantitative characters (
<tableCitation id="830BE8D9FFD31D42FAE9FB8FFA94FB9A" box="[1286,1370,1052,1077]" captionStart="TABLE 1" captionStartId="2.[151,239,1124,1147]" captionTargetBox="[159,1370,1206,2007]" captionTargetPageId="2" captionText="TABLE 1. Measurements in µm, pt index values relative to some structures, and percent ratio between the base + secondary and the main claw branches of Milnesium vorax sp. nov.." httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="2" pageNumber="577" tableUuid="9AF68DEAFFD31D42FF78FBF7FC64FB35">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="9AF68DEAFFD31D42FF78FBF7FC64FB35" ID-Table-UUID="9AF68DEAFFD31D42FF78FBF7FC64FB35" httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="2" pageNumber="577" targetBox="[159,1370,1206,2007]" targetIsTable="true" targetPageId="2">
<paragraph id="CE36DD62FFD31D42FF78FBF7FC64FB35" blockId="2.[151,1435,1124,1179]" pageId="2" pageNumber="577">
<emphasis id="FCFD0170FFD31D42FF78FBF7FEC2FBD5" bold="true" box="[151,268,1124,1147]" pageId="2" pageNumber="577">TABLE 1</emphasis>
. Measurements in µm,
<emphasis id="FCFD0170FFD31D42FDC2FBF7FD8FFBD5" box="[557,577,1124,1147]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
index values relative to some structures, and percent ratio between the base+secondary and the main claw branches of
<taxonomicName id="0989A6E1FFD31D42FD7BFB10FC86FB35" box="[660,840,1155,1179]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD31D42FD7BFB10FC86FB35" box="[660,840,1155,1179]" italics="true" pageId="2" pageNumber="577">Milnesium vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD31D42FCA1FB17FC6AFB35" box="[846,932,1156,1179]" pageId="2" pageNumber="577" rank="species">
<emphasis id="FCFD0170FFD31D42FCA1FB17FC6AFB35" bold="true" box="[846,932,1156,1179]" pageId="2" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="CE36DD62FFD31D42FF70FB72FB2EF879" pageId="2" pageNumber="577">
<table id="BC892FC2FFD3E2BFFF70FB25FA94F879" box="[159,1370,1206,2007]" gridcols="5" gridrows="7" pageId="2" pageNumber="577">
<tr id="70B9DF20FFD3E2BFFF70FB25FA94FABA" box="[159,1370,1206,1300]" gridrow="0" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70FB25FE01FABA" box="[159,463,1206,1300]" gridcol="0" gridrow="0" pageId="2" pageNumber="577">No. slide</th>
<th id="3368B65CFFD3E2BFFE1DFB25FD6AFABA" box="[498,676,1206,1300]" gridcol="1" gridrow="0" pageId="2" pageNumber="577">
<taxonomicName id="0989A6E1FFD31D42FE1DFB2BFD80FB63" box="[498,590,1208,1229]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD31D42FE1DFB2BFD80FB63" box="[498,590,1208,1229]" italics="true" pageId="2" pageNumber="577">M. vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD31D42FDBBFB24FD6AFB62" box="[596,676,1207,1228]" pageId="2" pageNumber="577" rank="species">
<emphasis id="FCFD0170FFD31D42FDBBFB24FD6AFB62" bold="true" box="[596,676,1207,1228]" pageId="2" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
2775 Ramacca paratype
</th>
<th id="3368B65CFFD3E2BFFD22FB25FCBDFABA" box="[717,883,1206,1300]" gridcol="2" gridrow="0" pageId="2" pageNumber="577">2775 Ramacca holotype</th>
<th id="3368B65CFFD3E2BFFC48FB25FBA9FABA" box="[935,1127,1206,1300]" gridcol="3" gridrow="0" pageId="2" pageNumber="577">
<taxonomicName id="0989A6E1FFD31D42FC48FB2BFB86FB62" box="[935,1096,1206,1229]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="species" species="tardigradum">
<emphasis id="FCFD0170FFD31D42FC48FB2BFB86FB62" box="[935,1096,1206,1229]" italics="true" pageId="2" pageNumber="577">M. tardigradum</emphasis>
</taxonomicName>
3436 Tunisia
</th>
<th id="3368B65CFFD3E2BFFB5BFB25FA94FABA" box="[1204,1370,1206,1300]" gridcol="4" gridrow="0" pageId="2" pageNumber="577">
<taxonomicName id="0989A6E1FFD31D42FB5AFB2BFAFAFB62" box="[1205,1332,1206,1229]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="577" phylum="Annelida" rank="species" species="reductum">
<emphasis id="FCFD0170FFD31D42FB5AFB2BFAFAFB62" box="[1205,1332,1206,1229]" italics="true" pageId="2" pageNumber="577">M. reductum</emphasis>
</taxonomicName>
5102 paratype
</th>
</tr>
<tr id="70B9DF20FFD3E2BFFF70FABFFA94FA13" box="[159,1370,1324,1469]" gridrow="1" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70FABFFE01FA13" box="[159,463,1324,1469]" gridcol="0" gridrow="1" pageId="2" pageNumber="577">Body length Buccal tube length Buccal tube width</th>
<td id="3368B65CFFD3E2BFFE1DFABFFD6AFA13" box="[498,676,1324,1469]" gridcol="1" gridrow="1" pageId="2" pageNumber="577">
µm
<emphasis id="FCFD0170FFD31D42FD83FABEFD4EFAEC" box="[620,640,1325,1346]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
578 35.8 21.3 59.5
</td>
<td id="3368B65CFFD3E2BFFD22FABFFCBDFA13" box="[717,883,1324,1469]" gridcol="2" gridrow="1" pageId="2" pageNumber="577">
µm
<emphasis id="FCFD0170FFD31D42FCA8FABEFC95FAEC" box="[839,859,1325,1346]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
585 35.0 21.1 60.3
</td>
<td id="3368B65CFFD3E2BFFC48FABFFBA9FA13" box="[935,1127,1324,1469]" gridcol="3" gridrow="1" pageId="2" pageNumber="577">
µm
<emphasis id="FCFD0170FFD31D42FBD4FABEFB80FAEC" box="[1083,1102,1325,1346]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
492 40.4 19.6 48.5
</td>
<td id="3368B65CFFD3E2BFFB5BFABFFA94FA13" box="[1204,1370,1324,1469]" gridcol="4" gridrow="1" pageId="2" pageNumber="577">
µm
<emphasis id="FCFD0170FFD31D42FAC1FABEFA8FFAEC" box="[1326,1345,1325,1346]" italics="true" pageId="2" pageNumber="577">pt</emphasis>
594 46.0 13.3 28.9
</td>
</tr>
<tr id="70B9DF20FFD3E2BFFF70FA42FA94F9BE" box="[159,1370,1489,1552]" gridrow="2" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70FA42FE01F9BE" box="[159,463,1489,1552]" gridcol="0" gridrow="2" pageId="2" pageNumber="577">Stylet supports insertion point Ext. main branch I</th>
<td id="3368B65CFFD3E2BFFE1DFA42FD6AF9BE" box="[498,676,1489,1552]" gridcol="1" gridrow="2" pageId="2" pageNumber="577">62.0 16.1 45.0</td>
<td id="3368B65CFFD3E2BFFD22FA42FCBDF9BE" box="[717,883,1489,1552]" gridcol="2" gridrow="2" pageId="2" pageNumber="577">62.2 16.3 46.6</td>
<td id="3368B65CFFD3E2BFFC48FA42FBA9F9BE" box="[935,1127,1489,1552]" gridcol="3" gridrow="2" pageId="2" pageNumber="577">60.7 16.5 40.8</td>
<td id="3368B65CFFD3E2BFFB5BFA42FA94F9BE" box="[1204,1370,1489,1552]" gridcol="4" gridrow="2" pageId="2" pageNumber="577">65.5 18.2 39.6</td>
</tr>
<tr id="70B9DF20FFD3E2BFFF70F9B7FA94F91B" box="[159,1370,1572,1717]" gridrow="3" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70F9B7FE01F91B" box="[159,463,1572,1717]" gridcol="0" gridrow="3" pageId="2" pageNumber="577">Base+sec. branch I % Base+sec/main branch I Ext. main branch II Base+sec. branch II</th>
<td id="3368B65CFFD3E2BFFE1DF9B7FD6AF91B" box="[498,676,1572,1717]" gridcol="1" gridrow="3" pageId="2" pageNumber="577">12.5 34.9 77.6 17.2 48.0 13.4 37.4</td>
<td id="3368B65CFFD3E2BFFD22F9B7FCBDF91B" box="[717,883,1572,1717]" gridcol="2" gridrow="3" pageId="2" pageNumber="577">12.5 35.7 76.7 17.4 49.7??</td>
<td id="3368B65CFFD3E2BFFC48F9B7FBA9F91B" box="[935,1127,1572,1717]" gridcol="3" gridrow="3" pageId="2" pageNumber="577">12.2 30.2 73.9 18.1 44.8 13.1 32.4</td>
<td id="3368B65CFFD3E2BFFB5BF9B7FA94F91B" box="[1204,1370,1572,1717]" gridcol="4" gridrow="3" pageId="2" pageNumber="577">12.5 27.1 68.7 20.9 45.4 13.5 29.3</td>
</tr>
<tr id="70B9DF20FFD3E2BFFF70F95AFA94F8A6" box="[159,1370,1737,1800]" gridrow="4" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70F95AFE01F8A6" box="[159,463,1737,1800]" gridcol="0" gridrow="4" pageId="2" pageNumber="577">% Base+sec/main branch II Ext. main branch III</th>
<td id="3368B65CFFD3E2BFFE1DF95AFD6AF8A6" box="[498,676,1737,1800]" gridcol="1" gridrow="4" pageId="2" pageNumber="577">77.9 18.1 50.6</td>
<td id="3368B65CFFD3E2BFFD22F95AFCBDF8A6" box="[717,883,1737,1800]" gridcol="2" gridrow="4" pageId="2" pageNumber="577">? 18.1 51.7</td>
<td id="3368B65CFFD3E2BFFC48F95AFBA9F8A6" box="[935,1127,1737,1800]" gridcol="3" gridrow="4" pageId="2" pageNumber="577">72.4 17.9 44.3</td>
<td id="3368B65CFFD3E2BFFB5BF95AFA94F8A6" box="[1204,1370,1737,1800]" gridcol="4" gridrow="4" pageId="2" pageNumber="577">64.6 22.4 48.7</td>
</tr>
<tr id="70B9DF20FFD3E2BFFF70F88FFA94F82A" box="[159,1370,1820,1924]" gridrow="5" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70F88FFE01F82A" box="[159,463,1820,1924]" gridcol="0" gridrow="5" pageId="2" pageNumber="577">Base+sec. branch III % Base+sec/main branch III Posterior main branch IV</th>
<td id="3368B65CFFD3E2BFFE1DF88FFD6AF82A" box="[498,676,1820,1924]" gridcol="1" gridrow="5" pageId="2" pageNumber="577">13.2 36.9 72.9 21.2 59.2</td>
<td id="3368B65CFFD3E2BFFD22F88FFCBDF82A" box="[717,883,1820,1924]" gridcol="2" gridrow="5" pageId="2" pageNumber="577">13.5 38.6 74.6 20.8 59.4</td>
<td id="3368B65CFFD3E2BFFC48F88FFBA9F82A" box="[935,1127,1820,1924]" gridcol="3" gridrow="5" pageId="2" pageNumber="577">12.8 31.7 71.5 21.8 54.0</td>
<td id="3368B65CFFD3E2BFFB5BF88FFA94F82A" box="[1204,1370,1820,1924]" gridcol="4" gridrow="5" pageId="2" pageNumber="577">14.4 31.3 64.3 27.0 58.7</td>
</tr>
<tr id="70B9DF20FFD3E2BFFF70F80BFA94F879" box="[159,1370,1944,2007]" gridrow="6" pageId="2" pageNumber="577">
<th id="3368B65CFFD3E2BFFF70F80BFE01F879" box="[159,463,1944,2007]" gridcol="0" gridrow="6" pageId="2" pageNumber="577">Posterior base+sec. branch IV % Base+sec/main branch IV</th>
<td id="3368B65CFFD3E2BFFE1DF80BFD6AF879" box="[498,676,1944,2007]" gridcol="1" gridrow="6" pageId="2" pageNumber="577">14.9 41.6 70.3</td>
<td id="3368B65CFFD3E2BFFD22F80BFCBDF879" box="[717,883,1944,2007]" gridcol="2" gridrow="6" pageId="2" pageNumber="577">14.9 42.6 71.6</td>
<td id="3368B65CFFD3E2BFFC48F80BFBA9F879" box="[935,1127,1944,2007]" gridcol="3" gridrow="6" pageId="2" pageNumber="577">14.0 34.7 64.2</td>
<td id="3368B65CFFD3E2BFFB5BF80BFA94F879" box="[1204,1370,1944,2007]" gridcol="4" gridrow="6" pageId="2" pageNumber="577">15.5 33.7 57.4</td>
</tr>
</table>
</paragraph>
<caption id="9AF68DEAFFD21D43FF78FA48FC24F9BE" httpUri="https://zenodo.org/record/271910/files/figure.png" pageId="3" pageNumber="578" targetBox="[190,1395,198,1470]" targetPageId="3">
<paragraph id="CE36DD62FFD21D43FF78FA48FC24F9BE" blockId="3.[151,1436,1499,1552]" pageId="3" pageNumber="578">
<emphasis id="FCFD0170FFD21D43FF78FA48FE82FA5F" bold="true" box="[151,332,1499,1521]" pageId="3" pageNumber="578">FIGURE 2. AC</emphasis>
,
<taxonomicName id="0989A6E1FFD21D43FEB5FA48FD85FA5F" box="[346,587,1499,1521]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="tardigradum">
<emphasis id="FCFD0170FFD21D43FEB5FA48FD85FA5F" box="[346,587,1499,1521]" italics="true" pageId="3" pageNumber="578">Milnesium tardigradum</emphasis>
</taxonomicName>
(paratype, slide no. 5489). A, Buccal tube (the arrow indicates a thickening of the buccal tube wall). B, Claws of legs II. C, Claws of the hind legs. Scale bar = 10 µm.
</paragraph>
</caption>
<subSubSection id="86938EE9FFD21D43FF28F9A4FAB2F9FE" box="[199,1404,1591,1616]" pageId="3" pageNumber="578" type="etymology">
<paragraph id="CE36DD62FFD21D43FF28F9A4FAB2F9FE" blockId="3.[151,1436,1591,2012]" box="[199,1404,1591,1616]" pageId="3" pageNumber="578">
<emphasis id="FCFD0170FFD21D43FF28F9A4FE80F9FE" bold="true" box="[199,334,1591,1616]" pageId="3" pageNumber="578">Etymology.</emphasis>
The specific name
<taxonomicName id="0989A6E1FFD21D43FDC6F9ABFDA7F9E1" box="[553,617,1592,1615]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="vorax">
<emphasis id="FCFD0170FFD21D43FDC6F9ABFDA7F9E1" box="[553,617,1592,1615]" italics="true" pageId="3" pageNumber="578">vorax</emphasis>
</taxonomicName>
(
<taxonomicName id="0989A6E1FFD21D43FD96F9ABFD77F9E1" box="[633,697,1592,1615]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="vorax">
<emphasis id="FCFD0170FFD21D43FD96F9ABFD77F9E1" box="[633,697,1592,1615]" italics="true" pageId="3" pageNumber="578">vorax</emphasis>
</taxonomicName>
= voracious) refers to the remarkable width of the buccal tube.
</paragraph>
</subSubSection>
<subSubSection id="86938EE9FFD21D44FF28F9C8FCEAFF7A" lastPageId="4" lastPageNumber="579" pageId="3" pageNumber="578" type="diagnosis">
<paragraph id="CE36DD62FFD21D43FF28F9C8FCCFF915" blockId="3.[151,1436,1591,2012]" pageId="3" pageNumber="578">
<emphasis id="FCFD0170FFD21D43FF28F9C8FE1FF9DA" bold="true" box="[199,465,1627,1652]" pageId="3" pageNumber="578">Differential diagnosis.</emphasis>
Three species are known having smooth cuticle, six peribuccal lamellae and claws configuration [2-3]-[3-2]:
<taxonomicName id="0989A6E1FFD21D43FE23F913FDAFF939" box="[460,609,1663,1687]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="reductum">
<emphasis id="FCFD0170FFD21D43FE23F913FDAFF939" box="[460,609,1663,1687]" italics="true" pageId="3" pageNumber="578">M. reductum</emphasis>
</taxonomicName>
,
<taxonomicName id="0989A6E1FFD21D43FD97F913FCFCF939" box="[632,818,1663,1687]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="tardigradum">
<emphasis id="FCFD0170FFD21D43FD97F913FCFCF939" box="[632,818,1663,1687]" italics="true" pageId="3" pageNumber="578">M. tardigradum</emphasis>
</taxonomicName>
, and, according to
<bibRefCitation id="AA18A093FFD21D43FBC8F913FA80F939" author="Meyer" box="[1063,1358,1663,1688]" pageId="3" pageNumber="578" refString="Meyer, H. A &amp; Hinton, J. C (2012) Terrestrial Tardigrada of the Island of Barbados in the West Indies, with the description of Milnesium barbadosense sp. n. (Eutardigrada: Apochela: Milnesiidae). Caribbean Journal of Science, 46, 2 - 3, 194 - 202." type="journal article" year="2012">Meyer &amp; Hinton (2012)</bibRefCitation>
, some specimens of
<taxonomicName id="0989A6E1FFD21D43FEDEF937FD33F915" authority="Meyer &amp; Hinton, 2012" authorityName="Meyer &amp; Hinton" authorityYear="2012" box="[305,765,1699,1724]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="barbadosense">
<emphasis id="FCFD0170FFD21D43FEDEF937FE3CF915" box="[305,498,1699,1723]" italics="true" pageId="3" pageNumber="578">M. barbadosense</emphasis>
<bibRefCitation id="AA18A093FFD21D43FE16F937FD33F915" author="Meyer" box="[505,765,1699,1724]" pageId="3" pageNumber="578" refString="Meyer, H. A &amp; Hinton, J. C (2012) Terrestrial Tardigrada of the Island of Barbados in the West Indies, with the description of Milnesium barbadosense sp. n. (Eutardigrada: Apochela: Milnesiidae). Caribbean Journal of Science, 46, 2 - 3, 194 - 202." type="journal article" year="2012">Meyer &amp; Hinton, 2012</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="CE36DD62FFD21D43FF28F954FECDF8E5" blockId="3.[151,1436,1591,2012]" pageId="3" pageNumber="578">
<taxonomicName id="0989A6E1FFD21D43FF28F954FE4BF971" box="[199,389,1735,1759]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD21D43FF28F954FE4BF971" box="[199,389,1735,1759]" italics="true" pageId="3" pageNumber="578">Milnesium vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD21D43FE63F954FE28F94E" box="[396,486,1735,1760]" pageId="3" pageNumber="578" rank="species">
<emphasis id="FCFD0170FFD21D43FE63F954FE28F94E" bold="true" box="[396,486,1735,1760]" pageId="3" pageNumber="578">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="0989A6E1FFD21D43FD93F95BFCE0F971" box="[636,814,1735,1759]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="tardigradum">
<emphasis id="FCFD0170FFD21D43FD93F95BFCE0F971" box="[636,814,1735,1759]" italics="true" pageId="3" pageNumber="578">M. tardigradum</emphasis>
</taxonomicName>
by the shape of the buccal tube, which is clearly wider and without a thickened wall posterior to the stylet supports (see
<tableCitation id="830BE8D9FFD21D43FC9EF978FC0AF8AD" box="[881,964,1771,1796]" captionStart="TABLE 1" captionStartId="2.[151,239,1124,1147]" captionTargetBox="[159,1370,1206,2007]" captionTargetPageId="2" captionText="TABLE 1. Measurements in µm, pt index values relative to some structures, and percent ratio between the base + secondary and the main claw branches of Milnesium vorax sp. nov.." httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="3" pageNumber="578" tableUuid="9AF68DEAFFD31D42FF78FBF7FC64FB35">Table 1</tableCitation>
and
<figureCitation id="56B2C1E7FFD21D43FC15F978FB95F8AD" box="[1018,1115,1771,1796]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="3" pageNumber="578">Figs. 1A</figureCitation>
and
<figureCitation id="56B2C1E7FFD21D43FB7DF978FB78F8AD" box="[1170,1206,1771,1795]" captionStart="FIGURE 2. A C" captionStartId="3.[151,250,1499,1521]" captionTargetBox="[190,1395,198,1470]" captionTargetId="figure@3.[183,1404,193,1477]" captionTargetPageId="3" captionText="FIGURE 2. A C, Milnesium tardigradum (paratype, slide no. 5489). A, Buccal tube (the arrow indicates a thickening of the buccal tube wall). B, Claws of legs II. C, Claws of the hind legs. Scale bar = 10 µm." httpUri="https://zenodo.org/record/271910/files/figure.png" pageId="3" pageNumber="578">2A</figureCitation>
, arrow). In addition, in
<taxonomicName id="0989A6E1FFD21D43FF5AF883FED2F889" box="[181,284,1808,1831]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD21D43FF5AF883FED2F889" box="[181,284,1808,1831]" italics="true" pageId="3" pageNumber="578">M. vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD21D43FECBF89CFEB1F889" box="[292,383,1807,1832]" pageId="3" pageNumber="578" rank="species">
<emphasis id="FCFD0170FFD21D43FECBF89CFEB7F886" bold="true" box="[292,377,1807,1832]" pageId="3" pageNumber="578">sp. nov</emphasis>
.
</taxonomicNameLabel>
the percent ratio between the base+secondary and the main claw branches is slightly higher (
<tableCitation id="830BE8D9FFD21D43FF71F8A0FF3CF8E5" box="[158,242,1843,1868]" captionStart="TABLE 1" captionStartId="2.[151,239,1124,1147]" captionTargetBox="[159,1370,1206,2007]" captionTargetPageId="2" captionText="TABLE 1. Measurements in µm, pt index values relative to some structures, and percent ratio between the base + secondary and the main claw branches of Milnesium vorax sp. nov.." httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="3" pageNumber="578" tableUuid="9AF68DEAFFD31D42FF78FBF7FC64FB35">Table 1</tableCitation>
).
</paragraph>
<paragraph id="CE36DD62FFD21D44FF28F8C4FCEAFF7A" blockId="3.[151,1436,1591,2012]" lastBlockId="4.[151,1436,151,501]" lastPageId="4" lastPageNumber="579" pageId="3" pageNumber="578">
The new species differs from
<taxonomicName id="0989A6E1FFD21D43FDFAF8CBFD6AF8C1" box="[533,676,1879,1903]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="reductum">
<emphasis id="FCFD0170FFD21D43FDFAF8CBFD6AF8C1" box="[533,676,1879,1903]" italics="true" pageId="3" pageNumber="578">M. reductum</emphasis>
</taxonomicName>
by the shape of the buccal tube, which is clearly wider and without a thickened wall posterior to the stylet supports (see
<tableCitation id="830BE8D9FFD21D43FD15F8E8FC81F83D" box="[762,847,1915,1940]" captionStart="TABLE 1" captionStartId="2.[151,239,1124,1147]" captionTargetBox="[159,1370,1206,2007]" captionTargetPageId="2" captionText="TABLE 1. Measurements in µm, pt index values relative to some structures, and percent ratio between the base + secondary and the main claw branches of Milnesium vorax sp. nov.." httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="3" pageNumber="578" tableUuid="9AF68DEAFFD31D42FF78FBF7FC64FB35">Table 1</tableCitation>
and
<figureCitation id="56B2C1E7FFD21D43FC64F8E8FC23F83D" box="[907,1005,1915,1940]" captionStart="FIGURE 1. A D" captionStartId="1.[151,250,1840,1862]" captionTargetBox="[213,1369,629,1808]" captionTargetId="figure@1.[209,1378,623,1815]" captionTargetPageId="1" captionText="FIGURE 1. A D, Milnesium vorax sp. nov. (holotype). A, Buccal tube. B, Claws of legs I (arrow a indicates a lunule; arrow b indicates the long cuticular thickening). C, Claws of legs III (the arrow indicates the long cuticular thickening). D, Claws of the hind legs (the arrow indicates a lunule). Scale bar = 10 µm." httpUri="https://zenodo.org/record/271909/files/figure.png" pageId="3" pageNumber="578">Figs. 1A</figureCitation>
and
<figureCitation id="56B2C1E7FFD21D43FBC6F8E8FB81F83D" box="[1065,1103,1915,1939]" captionStart="FIGURE 3. A C" captionStartId="4.[151,250,1754,1776]" captionTargetBox="[184,1402,562,1724]" captionTargetId="figure@4.[177,1410,555,1732]" captionTargetPageId="4" captionText="FIGURE 3. A C, Milnesium reductum (paratype, slide no. 5102). A, Buccal tube (the arrow indicates a thickening of the buccal tube wall). B, Claws of legs III. C, Claws of the hind legs. Scale bar = 10 µm." httpUri="https://zenodo.org/record/271911/files/figure.png" pageId="3" pageNumber="578">3A</figureCitation>
: arrow); by having the stylet supports inserted on the buccal tube in a more cephalic position (
<emphasis id="FCFD0170FFD21D43FC86F833FCB1F819" box="[873,895,1952,1975]" italics="true" pageId="3" pageNumber="578">pt</emphasis>
=
<quantity id="09717087FFD21D43FC73F80CFBDDF816" box="[924,1043,1951,1976]" metricMagnitude="0" metricUnit="m" metricValue="1.57734" metricValueMax="1.57988" metricValueMin="1.5748" pageId="3" pageNumber="578" unit="in" value="62.1" valueMax="62.2" valueMin="62.0">6262.2 in</quantity>
<taxonomicName id="0989A6E1FFD21D43FBF5F833FBB0F819" box="[1050,1150,1952,1975]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD21D43FBF5F833FBB0F819" box="[1050,1150,1952,1975]" italics="true" pageId="3" pageNumber="578">M. vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD21D43FB6BF80CFB10F816" box="[1156,1246,1951,1976]" pageId="3" pageNumber="578" rank="species">
<emphasis id="FCFD0170FFD21D43FB6BF80CFB10F816" bold="true" box="[1156,1246,1951,1976]" pageId="3" pageNumber="578">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<quantity id="09717087FFD21D43FB04F80CFAB9F816" box="[1259,1399,1951,1976]" metricMagnitude="0" metricUnit="m" metricValue="1.71196" metricValueMax="1.76022" metricValueMin="1.6637" pageId="3" pageNumber="578" unit="in" value="67.4" valueMax="69.3" valueMin="65.5">
65.5
<emphasis id="FCFD0170FFD21D43FAF3F80CFAE4F816" bold="true" box="[1308,1322,1951,1976]" pageId="3" pageNumber="578"></emphasis>
69.3 in
</quantity>
<taxonomicName id="0989A6E1FFD21D43FA92F833FECEF875" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="578" phylum="Annelida" rank="species" species="reductum">
<emphasis id="FCFD0170FFD21D43FA92F833FECEF875" italics="true" pageId="3" pageNumber="578">M. reductum</emphasis>
</taxonomicName>
according to
<bibRefCitation id="AA18A093FFD21D43FE4CF857FD9CF872" author="Tumanov" box="[419,594,1987,2012]" pageId="3" pageNumber="578" refString="Tumanov, D. V. (2006) Five new species of the genus Milnesium (Tardigrada, Eutardigrada, Milnesiidae). Zootaxa, 1122, 1 - 23." type="journal article" year="2006">Tumanov 2006</bibRefCitation>
) (
<tableCitation id="830BE8D9FFD21D43FD84F850FD0CF875" box="[619,706,1987,2012]" captionStart="TABLE 1" captionStartId="2.[151,239,1124,1147]" captionTargetBox="[159,1370,1206,2007]" captionTargetPageId="2" captionText="TABLE 1. Measurements in µm, pt index values relative to some structures, and percent ratio between the base + secondary and the main claw branches of Milnesium vorax sp. nov.." httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="3" pageNumber="578" tableUuid="9AF68DEAFFD31D42FF78FBF7FC64FB35">Table 1</tableCitation>
); by having claw main branches with accessory points, higher values of the
<emphasis id="FCFD0170FFD51D44FEDFFF0AFE88FF1E" box="[304,326,153,176]" italics="true" pageId="4" pageNumber="579">pt</emphasis>
index relative to the base+secondary claws and, as a consequence, a higher percent ratio between the base+secondary and the main claw branches (
<tableCitation id="830BE8D9FFD51D44FD2FFF2FFCDDFF7A" box="[704,787,188,213]" captionStart="TABLE 1" captionStartId="2.[151,239,1124,1147]" captionTargetBox="[159,1370,1206,2007]" captionTargetPageId="2" captionText="TABLE 1. Measurements in µm, pt index values relative to some structures, and percent ratio between the base + secondary and the main claw branches of Milnesium vorax sp. nov.." httpUri="http://table.plazi.org/id/9AF68DEAFFD31D42FF78FBF7FC64FB35" pageId="4" pageNumber="579" tableUuid="9AF68DEAFFD31D42FF78FBF7FC64FB35">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86938EE9FFD51D44FF28FF73FB8DFE5A" pageId="4" pageNumber="579" type="reference_group">
<paragraph id="CE36DD62FFD51D44FF28FF73FB8DFE5A" blockId="4.[151,1436,151,501]" pageId="4" pageNumber="579">
In
<taxonomicName id="0989A6E1FFD51D44FF07FF72FE65FF56" box="[232,427,224,248]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="579" phylum="Annelida" rank="species" species="barbadosense">
<emphasis id="FCFD0170FFD51D44FF07FF72FE65FF56" box="[232,427,224,248]" italics="true" pageId="4" pageNumber="579">M. barbadosense</emphasis>
</taxonomicName>
the claw configuration is [3-3]-[3-3], but
<bibRefCitation id="AA18A093FFD51D44FC7DFF73FB68FF56" author="Meyer" box="[914,1190,223,248]" pageId="4" pageNumber="579" refString="Meyer, H. A &amp; Hinton, J. C (2012) Terrestrial Tardigrada of the Island of Barbados in the West Indies, with the description of Milnesium barbadosense sp. n. (Eutardigrada: Apochela: Milnesiidae). Caribbean Journal of Science, 46, 2 - 3, 194 - 202." type="journal article" year="2012">Meyer &amp; Hinton (2012)</bibRefCitation>
also attributed to the same species a few specimens, most less than 300 µm long and some larger, with claw configuration [2-3]-[3-2].
<taxonomicName id="0989A6E1FFD51D44FF78FEBAFF30FEEE" box="[151,254,297,320]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="579" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD51D44FF78FEBAFF30FEEE" box="[151,254,297,320]" italics="true" pageId="4" pageNumber="579">M. vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD51D44FEE9FEB4FEAFFEEE" box="[262,353,295,320]" pageId="4" pageNumber="579" rank="species">
<emphasis id="FCFD0170FFD51D44FEE9FEB4FEAFFEEE" bold="true" box="[262,353,295,320]" pageId="4" pageNumber="579">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from these specimens by having: eye spots present; higher
<emphasis id="FCFD0170FFD51D44FBB2FEBAFBBDFEEE" box="[1117,1139,297,320]" italics="true" pageId="4" pageNumber="579">pt</emphasis>
index value of the buccal tube width (about
<quantity id="09717087FFD51D44FE8CFEDFFE52FECB" box="[355,412,332,357]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="579" unit="in" value="60.0">60 in</quantity>
<taxonomicName id="0989A6E1FFD51D44FE4CFEDEFDC6FECA" box="[419,520,333,356]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="579" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD51D44FE4CFEDEFDC6FECA" box="[419,520,333,356]" italics="true" pageId="4" pageNumber="579">M. vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD51D44FDE0FEDFFDA6FECB" box="[527,616,332,357]" pageId="4" pageNumber="579" rank="species">
<emphasis id="FCFD0170FFD51D44FDE0FEDFFDA6FECB" bold="true" box="[527,616,332,357]" pageId="4" pageNumber="579">sp. nov.</emphasis>
</taxonomicNameLabel>
, about
<quantity id="09717087FFD51D44FD54FEDFFD3AFECB" box="[699,756,332,357]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="579" unit="in" value="50.0">50 in</quantity>
larger specimens of
<taxonomicName id="0989A6E1FFD51D44FC33FEDEFB53FECA" box="[988,1181,332,356]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="579" phylum="Annelida" rank="species" species="barbadosense">
<emphasis id="FCFD0170FFD51D44FC33FEDEFB53FECA" box="[988,1181,332,356]" italics="true" pageId="4" pageNumber="579">M. barbadosense</emphasis>
</taxonomicName>
according to
<bibRefCitation id="AA18A093FFD51D44FAD9FEDEFEFEFE26" author="Meyer" pageId="4" pageNumber="579" refString="Meyer, H. A &amp; Hinton, J. C (2012) Terrestrial Tardigrada of the Island of Barbados in the West Indies, with the description of Milnesium barbadosense sp. n. (Eutardigrada: Apochela: Milnesiidae). Caribbean Journal of Science, 46, 2 - 3, 194 - 202." type="journal article" year="2012">Meyer &amp; Hinton, 2012</bibRefCitation>
); stylet supports inserted on the buccal tube in a more cephalic position (
<emphasis id="FCFD0170FFD51D44FBB2FEE2FBBDFE26" box="[1117,1139,369,392]" italics="true" pageId="4" pageNumber="579">pt</emphasis>
=
<quantity id="09717087FFD51D44FB7FFEFCFB07FE26" box="[1168,1225,367,392]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" pageId="4" pageNumber="579" unit="in" value="62.0">62 in</quantity>
<taxonomicName id="0989A6E1FFD51D44FB3FFEE2FAFBFE26" box="[1232,1333,369,392]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="579" phylum="Annelida" rank="species" species="vorax" status="sp. nov.">
<emphasis id="FCFD0170FFD51D44FB3FFEE2FAFBFE26" box="[1232,1333,369,392]" italics="true" pageId="4" pageNumber="579">M. vorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7CEBC0BFFD51D44FAD3FEFCFA5BFE26" box="[1340,1429,367,392]" pageId="4" pageNumber="579" rank="species">
<emphasis id="FCFD0170FFD51D44FAD3FEFCFA5BFE26" bold="true" box="[1340,1429,367,392]" pageId="4" pageNumber="579">sp. nov.</emphasis>
</taxonomicNameLabel>
, at least
<quantity id="09717087FFD51D44FF00FE07FEE7FE03" box="[239,297,404,429]" metricMagnitude="0" metricUnit="m" metricValue="1.6764" pageId="4" pageNumber="579" unit="in" value="66.0">66 in</quantity>
<taxonomicName id="0989A6E1FFD51D44FEDEFE06FE3AFE02" box="[305,500,404,428]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="579" phylum="Annelida" rank="species" species="barbadosense">
<emphasis id="FCFD0170FFD51D44FEDEFE06FE3AFE02" box="[305,500,404,428]" italics="true" pageId="4" pageNumber="579">M. barbadosense</emphasis>
</taxonomicName>
(according to
<bibRefCitation id="AA18A093FFD51D44FD75FE06FC52FE02" author="Meyer" box="[666,924,404,429]" pageId="4" pageNumber="579" refString="Meyer, H. A &amp; Hinton, J. C (2012) Terrestrial Tardigrada of the Island of Barbados in the West Indies, with the description of Milnesium barbadosense sp. n. (Eutardigrada: Apochela: Milnesiidae). Caribbean Journal of Science, 46, 2 - 3, 194 - 202." type="journal article" year="2012">Meyer &amp; Hinton 2012</bibRefCitation>
); higher values of the
<emphasis id="FCFD0170FFD51D44FB74FE06FB7FFE02" box="[1179,1201,405,428]" italics="true" pageId="4" pageNumber="579">pt</emphasis>
index relative to the base+secondary claws and, as a consequence, higher percent ratio between the base+secondary and the main claw branches (the difference is clearer in the claws IIII, and very small in the claw IV).
</paragraph>
</subSubSection>
</treatment>
</document>