treatments-xml/data/46/18/BD/4618BD5BFFC3FFEDF7F024E120EDFD14.xml
2024-06-21 12:35:10 +02:00

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<document id="D0299E7B92A691C948D42241F7CE0ED6" ID-DOI="10.5281/zenodo.5275243" ID-ISSN="0253-116X" ID-Zenodo-Dep="5275243" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629968455998" checkinUser="marcus" docAuthor="Balkenohl, Michael" docDate="2020" docId="4618BD5BFFC3FFEDF7F024E120EDFD14" docLanguage="en" docName="LinzerbiolBeitr.2020.52.1.127-140.pdf" docOrigin="Linzer biologische Beiträge 52 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Salcedia occidentalis Balkenohl 2020, nov.sp." docType="treatment" docVersion="3" lastPageNumber="135" masterDocId="BA21C523FFC5FFE5F76726482134FF88" masterDocTitle="Salcedia FAIRMAIRE, 1899 from Africa with descriptions of the two new species S. bembereka nov. sp. and S. occidentalis nov. sp. (Coleoptera, Carabidae, Scaritinae, Salcediini)" masterLastPageNumber="140" masterPageNumber="127" pageNumber="133" updateTime="1699132443490" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F3C9766D066D9E15C0D6AFD6B487C53C">Salcedia FAIRMAIRE, 1899 from Africa with descriptions of the two new species S. bembereka nov. sp. and S. occidentalis nov. sp. (Coleoptera, Carabidae, Scaritinae, Salcediini)</mods:title>
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<mods:namePart id="EE05B07115BD71E19D8275DAD66F0173">Balkenohl, Michael</mods:namePart>
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<mods:title id="36FDB2A31DDAD84E09330269D99DF83C">Linzer biologische Beiträge</mods:title>
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<mods:date id="2FA6AAD3E9E7AA55254580A9198534ED">2020</mods:date>
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<mods:number id="7E82BBCEEC5D36C580527E2777F2B0B2">2020-07-31</mods:number>
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<mods:number id="E015306255FA2048F78A98667386D687">52</mods:number>
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<mods:identifier id="30478F57A53689BD806B33B78418A69E" type="ISSN">0253-116X</mods:identifier>
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<subSubSection id="86AB5FC6FFC3FFE3F7F024E12250FC9D" pageId="6" pageNumber="133" type="nomenclature">
<paragraph id="CE0E0C4DFFC3FFE3F7F024E1235CFD37" blockId="6.[151,616,681,703]" box="[151,616,681,703]" pageId="6" pageNumber="133">
<heading id="9546BB21FFC3FFE3F7F024E1235CFD37" box="[151,616,681,703]" fontSize="8" level="2" pageId="6" pageNumber="133" reason="2">
<emphasis id="FCC5D05FFFC3FFE3F7F024E1235CFD37" bold="true" box="[151,616,681,703]" pageId="6" pageNumber="133">
<taxonomicName id="09B177CEFFC3FFE3F7F024E12046FD37" authority="Balkenohl, 2020" authorityName="Balkenohl" authorityYear="2020" box="[151,370,681,703]" class="Insecta" family="Carabidae" genus="Salcedia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="FCC5D05FFFC3FFE3F7F024E12046FD37" bold="true" box="[151,370,681,703]" italics="true" pageId="6" pageNumber="133">Salcedia occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7F66D24FFC3FFE3F61F24E120F1FD36" box="[376,453,681,702]" pageId="6" pageNumber="133" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="568A10C8FFC3FFE3F6B524E12321FD36" box="[466,533,681,703]" captionStart="Figs 1, 2" captionStartId="11.[151,191,1468,1490]" captionTargetBox="[165,1028,334,1435]" captionTargetId="figure-58@11.[152,1051,313,1460]" captionTargetPageId="11" captionText="Figs 1, 2: Salcedia species, habitus, dorsal view, holotype. (1) S. bembereka nov.sp.; (2) S. occidentalis nov.sp." figureDoi="http://doi.org/10.5281/zenodo.5275246" httpUri="https://zenodo.org/record/5275246/files/figure.png" pageId="6" pageNumber="133">Figs 2</figureCitation>
, 4, 6, 8)
</emphasis>
</heading>
</paragraph>
<paragraph id="CE0E0C4DFFC3FFE3F7F024872250FC9D" blockId="6.[151,1059,719,855]" pageId="6" pageNumber="133">
T y p e m a t e r i a l:
<typeStatus id="110AB2EFFFC3FFE3F611248720FFFD6D" box="[374,459,719,741]" pageId="6" pageNumber="133" type="holotype">Holotype</typeStatus>
:
<specimenCount id="D8B7C7C4FFC3FFE3F6B0248720DDFD6C" box="[471,489,719,740]" pageId="6" pageNumber="133" type="male"></specimenCount>
, with labels and data: white, black printed, black framed &quot;N GUINEA-CONACRY NEAR MBOURIA N BALAKI 03.-
<date id="BA0F2A8DFFC3FFE3F5B224AF2208FD74" box="[725,828,743,764]" pageId="6" pageNumber="133" value="2004-04-07">04.07.2004</date>
A. KUDRNA JR. LGT.&quot; (CBB).
<typeStatus id="110AB2EFFFC3FFE3F66D24B72056FC9C" box="[266,354,767,788]" pageId="6" pageNumber="133" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="D8B7C7C4FFC3FFE3F60E24B720B2FC9C" box="[361,390,767,788]" pageId="6" pageNumber="133" type="male">1♂</specimenCount>
,
<specimenCount id="D8B7C7C4FFC3FFE3F6E824B72088FC9F" box="[399,444,767,791]" pageId="6" pageNumber="133" type="female">2♀♀</specimenCount>
, 2 specs., same data as
<typeStatus id="110AB2EFFFC3FFE3F5EF24B723E1FC9D" box="[648,725,767,789]" pageId="6" pageNumber="133" type="holotype">holotype</typeStatus>
(OLML, CBB).
</paragraph>
</subSubSection>
<subSubSection id="86AB5FC6FFC3FFEDF7F0256C20EDFD14" lastPageId="8" lastPageNumber="135" pageId="6" pageNumber="133" type="description">
<paragraph id="CE0E0C4DFFC3FFE3F7F0256C2516FCB1" blockId="6.[151,1059,719,855]" box="[151,1058,803,825]" pageId="6" pageNumber="133">E t y m o l o g y: The name refers to the most westerly occurrence of the genus in</paragraph>
<paragraph id="CE0E0C4DFFC3FFE3F7F0250921EBFCDF" blockId="6.[151,1059,719,855]" box="[151,223,833,855]" pageId="6" pageNumber="133">Africa.</paragraph>
<paragraph id="CE0E0C4DFFC3FFE3F7F0252F201FFB29" blockId="6.[151,1059,870,1185]" pageId="6" pageNumber="133">
D i a g n o s i s: A large sized species, with elongate outline of the elytra with maximum width behind middle and the pronotum with three additional carinae of which the inner one is small and the outer lateral carina is shortened anteriorly and posteriorly. The pseudohumerus is slightly obtuse angular, slightly bent down, distinctly dentate. The antennomeres are super-moniliform. Distinguished most clearly from the similar species
<emphasis id="FCC5D05FFFC3FFE3F7F025B12022FB86" box="[151,278,1017,1038]" italics="true" pageId="6" pageNumber="133">S. baroensis</emphasis>
BALKENOHL by the lateral margin of the pronotum with the more distinct and separated tubercles and the more distinct developed inner and outer lateral carinae. From the similar species
<taxonomicName id="09B177CEFFC3FFE3F6C7227C23B5FBC0" authority="ALLUAUD" authorityName="ALLUAUD" box="[416,641,1075,1097]" class="Insecta" family="Carabidae" genus="Salcedia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="FCC5D05FFFC3FFE3F6C7227C2320FBC0" box="[416,532,1075,1097]" italics="true" pageId="6" pageNumber="133">S. elongata</emphasis>
ALLUAUD
</taxonomicName>
, it differs by the broader head which is three quarters of the width of the pronotum and the elytron with its maximum width behind middle. Distinguished from all other species by the super-moniliform antennomeres.
</paragraph>
<paragraph id="CE0E0C4DFFC3FFE3F7F022F92058FB4E" blockId="6.[151,364,1201,1223]" box="[151,364,1201,1223]" pageId="6" pageNumber="133">D e s c r i p t i o n:</paragraph>
<paragraph id="CE0E0C4DFFC3FFE3F7F0229F20A7FB63" blockId="6.[151,403,1238,1260]" box="[151,403,1238,1260]" pageId="6" pageNumber="133">
Measurements in
<tableCitation id="833339F6FFC3FFE3F62C229E20BBFB63" box="[331,399,1238,1260]" captionStart="Tab" captionStartId="2.[148,189,469,494]" captionText="Tab. 2. Measurements and ratios of Salcedia species, updated table. N = number of specimens measured (see Material and Methods);  = arithmetic mean; &quot;term used&quot; refers to the shape of the elytra and represents as well the ratio length/width of both of the elytra." pageId="6" pageNumber="133">Tab. 2</tableCitation>
.
</paragraph>
<paragraph id="CE0E0C4DFFC3FFE3F7F022B322E7FAA7" blockId="6.[151,1058,1275,1327]" pageId="6" pageNumber="133">Colour and surface: Anthracite grey with slight piceous shine, areas between carinae and pits shiny; mandibles and legs piceous, and antennae hinnuleous, palpi leoninous.</paragraph>
<paragraph id="CE0E0C4DFFC3FFE2F7F02376231FFD87" blockId="6.[151,1059,1342,1599]" lastBlockId="7.[151,1059,211,527]" lastPageId="7" lastPageNumber="134" pageId="6" pageNumber="133">
Head: Three-quarters of pronotum width. Outline shaped like frustum of a pyramid. Clypeus wide, straight anteriorly, fused with clypeal wings, separated from supraantennal plates by distinct notches, with convex semicircular field at middle, separated from frons by moderately deep transverse furrow. Frons with two raised paramedian carinae, joining anteriorly, prolonged anteriorly into a central distinctly erected tubercle, with two small rounded glossy teeth bilaterally anterior to central tubercle, with two parallel carinae paralaterally near base; frons and clypeus separated from supraantennal and supraorbital plates by deep broad furrows, each furrow with deep slightly longitudinal pit between supraantennal plate and clypeus, with circular conspicuously deep pit at front-eye level; supraantennal and supraorbital plates acutely margined, margin raised, carina-like, supraantennal plates slightly vaulted. Base emarginated at middle, slightly obtuse angled laterally (angle 106-108°). Eyes convex, with transverse-pentagonal shape in lateral view, with slightly concave posterior margin, with small part just visible in dorsal view; genae slightly convex, slightly higher than eyes. Antenna with segments five to ten super-moniliform (L/
<geoCoordinate id="AB856A8AFFC2FFE2F46A272E227AFEF4" box="[781,846,358,380]" degrees="1.1" direction="west" orientation="longitude" pageId="7" pageNumber="134" precision="5555" value="-1.1">W 1.1</geoCoordinate>
), densely pubescent, segments two to four scarcely pubescent, scapus and pedicellus with longitudinal reticulation. Labrum straight anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Apical segment of maxillary palpomere moderately long. Mentum small, with obtuse tooth at middle, epilobes wide, projecting and distinctly angled anteriorly, margined anteriorly, surface coriacious.
</paragraph>
<paragraph id="CE0E0C4DFFC2FFE2F7F024572254FCFF" blockId="7.[151,1059,542,887]" pageId="7" pageNumber="134">Pronotum (Fig. 4): Outline rectangular, transverse, more than a quarter wider than long. Lateral margin slightly convex at middle, maximum width at end of second third, distinctly converging anteriorly and slightly posteriorly. Lateral margin distinctly crenulated, with ten tubercles, with distinct emargination at posterior angles. Base straight laterally, with wide notch, with decreasing keel at middle. Disc with two raised paramedian carinae parallel to median line, diverging posteriorly, with long median line ending in pits anteriorly and posteriorly, with four additional shorter carinae bilaterally, the paralateral ones joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, extended anteriorly as distinctly raised paralateral carina. With two short and less raised inner and outer lateral carinae, the inner one very small and isolated. All carinae sub-crenulate. Lateral margin and space between lateral margin and paralateral carina wing-like bent up, with six large transverse pits.</paragraph>
<paragraph id="CE0E0C4DFFC2FFE2F7F025CE2287FBE1" blockId="7.[151,1059,902,1129]" pageId="7" pageNumber="134">Elytron: Flattened in anterior two thirds (lateral view), moderately convex in frontal view. Elongate, margin straight in anterior third but diverging, maximum width behind middle. Pseudohumerus slightly obtuse angular, slightly bent down, distinctly dentate. Apex rounded, acutely and small denticulate at suture. Disc with interneur six subcrenulate, interneur two running to apex as slightly convex line, conspicuously raised, not reaching apex; interneur four running convex towards apex, decreasing before reaching basal area, not reaching base, not reaching apex. Interneur five and six with two rows of serial pits, pits clearly separated, merging transversally in apical sixth.</paragraph>
<paragraph id="CE0E0C4DFFC2FFE2F7F0223120F1FB07" blockId="7.[151,453,1145,1167]" box="[151,453,1145,1167]" pageId="7" pageNumber="134">Hind wings: Fully developed.</paragraph>
<paragraph id="CE0E0C4DFFC2FFE2F7F022D72068FAA1" blockId="7.[151,1059,1182,1321]" pageId="7" pageNumber="134">Lower surface: Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with two rows of pits, the lateral one interrupted at middle, lateral margin of elytron sub-crenulate. Metepisternum distinctly elongated, with circular pits. Last visible sternum with large pit at apex, lateral pits merged longitudinally with blunt irregular carinae in between.</paragraph>
<paragraph id="CE0E0C4DFFC2FFE2F7F023712081FA4C" blockId="7.[151,1059,1337,1476]" pageId="7" pageNumber="134">Legs: Profemora with dorsal surface isodiametrically reticulated. Protibia stout, with short, robust, moderately curved terminal spine, laterally with two moderately large and two small sized teeth, dorsally with one distinct and one indistinct carina. Movable spur short, length a quarter of first tarsalia. First tarsalia distinctly elongated, almost as long as tarsalia two to five together.</paragraph>
<paragraph id="CE0E0C4DFFC2FFE2F7F0239B2033F9AC" blockId="7.[151,1058,1491,1572]" pageId="7" pageNumber="134">External sexual dimorphism: The large pit at the apex of the last visible sternum of the abdomen is slightly larger and circular in females. In males it is longitudinally developed.</paragraph>
<paragraph id="CE0E0C4DFFCDFFEDF7F0269B20D8FE3F" blockId="8.[151,1059,211,439]" pageId="8" pageNumber="135">
Male genitalia (
<figureCitation id="568A10C8FFCDFFEDF65B269B2048FF61" box="[316,380,211,233]" captionStart="Figs 5, 6" captionStartId="12.[151,191,1372,1394]" captionTargetBox="[165,1001,255,1348]" captionTargetId="figure-78@12.[149,1055,225,1358]" captionTargetPageId="12" captionText="Figs 5, 6: Salcedia species, male genitalia, dorsal view of aedeagus (a), parameres (b, c) and lateral view (d). Black small sketches are symbolizing the apex in cross section. (5). S. bembereka nov.sp.; (6) S. occidentalis nov.sp." figureDoi="http://doi.org/10.5281/zenodo.5275248" httpUri="https://zenodo.org/record/5275248/files/figure.png" pageId="8" pageNumber="135">Fig. 6</figureCitation>
): Median lobe robust, in dorsal view regularly arcuate, distinctly arcuate in apical fifth, in lateral view continuously becoming smaller towards apex, with fine scattered pili in middle part, apex stick-like, in cross section broad-oval towards apex, directly at apex round and hollowed out convexly ventrally. Oroficium half as long as median lobe. Endophallus with group of microtrichia conspicuously densely arranged, with additional small group of finer and short trichia basally. Dorsal paramere elongated, slightly bisinuate, with elongated robust apophyses; ventral one broad triangular like; both parameres slightly distorted.
</paragraph>
<paragraph id="CE0E0C4DFFCDFFEDF7F0278E2316FD9F" blockId="8.[151,1059,454,535]" pageId="8" pageNumber="135">Female genitalia (Fig. 8): Coxostylus regularly broadened to base, distinctly curved, at end of basal third with one small seta, one strong and six slender nematiform setae laterally, SSO with one microtrichium.</paragraph>
<paragraph id="CE0E0C4DFFCDFFEDF7F0246E21E3FDD1" blockId="8.[151,1059,550,601]" pageId="8" pageNumber="135">Variation: The tubercles at the lateral margin of the pronotum vary in number from nine to ten.</paragraph>
<paragraph id="CE0E0C4DFFCDFFEDF7F0242120EDFD14" blockId="8.[151,1058,617,668]" pageId="8" pageNumber="135">
D i s t r i b u t i o n: The species occurs in Conacry, West of
<collectingCountry id="B6A64CDDFFCDFFEDF47A24212252FDF7" box="[797,870,617,639]" name="Guinea-Bissau" pageId="8" pageNumber="135">Guinea</collectingCountry>
close to the banks of the river Konkoure (Fig. 10).
</paragraph>
</subSubSection>
</treatment>
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