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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.46.104548" ID-Pensoft-Pub="1314-2615-46-21" ID-Pensoft-UUID="7C8C1294D4B05D749CC9DF10F2D7458D" ID-ZooBank="17340FB714C84903B1E48B209FE73C93" ModsDocID="1314-2615-46-21" checkinTime="1692896423125" checkinUser="pensoft" docAuthor="Curcic, Srecko, Vesovic, Nikola, Vrbica, Maja, Popovic, Slađana, Radovanovic, Zeljko, Curcic, Nina B., Yamashkin, Anatoliy A., Radovic, Dejan, Yamashkin, Stanislav A., Vranic, Sofija &amp; Rađa, Tonci" docDate="2023" docId="39CA5DDBE2655904B9964A2E1DE21EF0" docLanguage="en" docName="SubterBiol 46: 21-46" docOrigin="Subterranean Biology 46" docPubDate="2023-08-24" docSource="http://dx.doi.org/10.3897/subtbiol.46.104548" docTitle="Spelaeobates (Spelaeobates) coriniensis subsp. nonveilleri Curcic, Vesovic, Vrbica &amp; Rađa 2023, ssp. nov." docType="treatment" docUuid="D92C0413-A52D-45A8-AE12-759986CD2F8F" docUuidSource="ZooBank" docVersion="2" id="7C8C1294D4B05D749CC9DF10F2D7458D" lastPageNumber="21" masterDocId="7C8C1294D4B05D749CC9DF10F2D7458D" masterDocTitle="The surprising discovery of two new subterranean Leptodirini of the genus Spelaeobates Mueller, 1901 (Coleoptera, Leiodidae, Cholevinae) from Croatia after more than a century" masterLastPageNumber="46" masterPageNumber="21" pageNumber="21" updateTime="1692896725838" updateUser="ExternalLinkService">
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<mods:title id="A9EC32F1C746144480F6BF6213B4DD73">The surprising discovery of two new subterranean Leptodirini of the genus Spelaeobates Mueller, 1901 (Coleoptera, Leiodidae, Cholevinae) from Croatia after more than a century</mods:title>
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<mods:namePart id="8F209812C81118E6C29DB47AB2AF2D15">Curcic, Srecko</mods:namePart>
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<mods:affiliation id="B0DF02E2C13798EBC946768936E92D93">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:nameIdentifier id="14A0277D998D20FBC2C4B60913160495" type="email">srecko@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="0C0B584BCF9F2A0EA6504C0734E6C0BA">Vesovic, Nikola</mods:namePart>
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<mods:namePart id="D3247B90E1059B43FA74178AB878B5ED">Vrbica, Maja</mods:namePart>
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<mods:namePart id="39AF4F8310BED415727141BBE01A535D">Popovic, Slađana</mods:namePart>
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<mods:affiliation id="0366154A4D8836A795474671A40B222B">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="BBF300DBE727C6C74C7DBF68AEAA1A9D">Radovanovic, Zeljko</mods:namePart>
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<mods:namePart id="D8A57E7797AAE58408B5CFD22D67395B">Curcic, Nina B.</mods:namePart>
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<mods:affiliation id="9C5CF10614810CB1AC4518BB2E03C4EA">University of Belgrade, Innovation Center of the Faculty of Technology and Metallurgy, Karnegijeva 4, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="1FF30A2DEDA50C11B0F3392B540F4F85">Yamashkin, Anatoliy A.</mods:namePart>
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<mods:affiliation id="5D0582EF9A549D5BAD23590C143CACBB">Geographical Institute &quot; Jovan Cvijic &quot;, Serbian Academy of Sciences and Arts, Đure Jaksica 9, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="0CA6E596786C74D5F47C3E477D111082">Radovic, Dejan</mods:namePart>
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<mods:namePart id="BB5041380DE15699E351BCB77C3D3548">Vranic, Sofija</mods:namePart>
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<mods:namePart id="9BC09B2950120AA21E4EFD43C3EDFE7C">Rađa, Tonci</mods:namePart>
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<mods:date id="41931B28A240AF1917248E1B619556C8">2023</mods:date>
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<taxonomicName id="4751380796BEB3F2E778291011E20122" LSID="https://zoobank.org/D92C0413-A52D-45A8-AE12-759986CD2F8F" authority="Ćurčić, Vesović, Vrbica &amp; Rađa" authorityName="Curcic, Vesovic, Vrbica &amp; Rađa" authorityYear="2023" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) coriniensis subsp. nonveilleri" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="coriniensis" status="ssp. nov." subGenus="Spelaeobates" subSpecies="nonveilleri">
Spelaeobates (Spelaeobates) coriniensis nonveilleri 
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, 
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, Vrbica &amp; 
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<paragraph id="6B1B26223FA65E1732AB5D5FA76B02FA" pageId="0" pageNumber="21">
<figureCitation id="1F655574C52F25B0219747A4C9BFD7E6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">Figs 4</figureCitation>
<figureCitation id="8D5017C050FF8C21A9D8D4F435EA4807" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">, 5</figureCitation>
<figureCitation id="7192388E9AA047FC1C7B65E81E6ED794" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bright-field images of certain morphological traits of Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov. from the Jamurka (Rnjakusa II) Pit, village of Popovici, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia (A, B) and S. (S.) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia (C-E) A, C PT female, spermatheca, lateral view B PT female, gonostyli, dorsal view D HT male, abdominal sternite IX (urite) E PT female, abdominal ventrite VIII. Scale bars: 0.05 mm (A, C); 0.10 mm (B, D, E)." figureDoi="10.3897/subtbiol.46.104548.figure6" httpUri="https://binary.pensoft.net/fig/896585" pageId="0" pageNumber="21">, 6</figureCitation>
</paragraph>
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<paragraph id="1DE0D644583D724E9B7DD67D60818045" pageId="0" pageNumber="21">Type material.</paragraph>
<paragraph id="B7C78D9676E2407AB170EEEF3A9BFDD6" pageId="0" pageNumber="21">
<emphasis id="CA241786FEECAA5BEE209E3AC5D2C827" bold="true" italics="true" pageId="0" pageNumber="21">Holotype</emphasis>
: male (SSM) labeled as follows: &quot;CROATIA, NORTHERN DALMATIA: city of 
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, island of Murter, settlement of Tisno, village of Jezera, Jezeranka Pit, 42 m a.s.l., 
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, 
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, 2.VI.2019, TR&quot; (white label, printed) / &quot;Holotypus 
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ssp. nov. 
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, 
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, Vrbica &amp; 
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det. 2022&quot; (red label, printed).
</paragraph>
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<emphasis id="E368CFBDFC9F55C89235F6109065EC15" bold="true" italics="true" pageId="0" pageNumber="21">Paratypes</emphasis>
(10 specimens). The same data as for HT [three males and five females, IZFB]; two males (IZFB) labeled as follows: &quot;CROATIA, NORTHERN DALMATIA: city of 
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, village of Banjevci, 
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Pit, 149 m a.s.l., 
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, 
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, 5.XI.2019, TR&quot;. All paratypes are labeled with white, printed locality labels and with red printed labels &quot;Paratypus 
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ssp. nov. 
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, 
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, Vrbica &amp; 
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det. 2022&quot; (Fig. 
<figureCitation id="A0A2BF4F33597B050A2A91AA4D879EDC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4</figureCitation>
).
</paragraph>
<caption id="9AB98D0FB0FA2938379B5328CBF03E04" doi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21" start="Figure 4" startId="F4">
<paragraph id="5637C261FB9FCA5C76C1763C2952A719" pageId="0" pageNumber="21">
<emphasis id="0C27185F48469F093A96D2B69339CF2E" bold="true" pageId="0" pageNumber="21">Figure 4.</emphasis>
SEM images of the morphological structures of PT male (
<emphasis id="84350A7033227FB443A254F335E144AC" bold="true" pageId="0" pageNumber="21">A-G, J-L</emphasis>
) and PT female (
<emphasis id="DD0FBBA809E9BEE77E4C89ED1CA04BDF" bold="true" pageId="0" pageNumber="21">H, I</emphasis>
) of 
<taxonomicName id="99F729E4E0BC2D9513B6512ABF13B48B" authorityName="Curcic, Vesovic, Vrbica &amp; Rađa" authorityYear="2023" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) coriniensis subsp. nonveilleri" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="coriniensis" subGenus="Spelaeobates" subSpecies="nonveilleri">Spelaeobates (Spelaeobates) coriniensis nonveilleri</taxonomicName>
ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of 
<normalizedToken id="F45545F42B6D77080137E719E3244241" originalValue="Šibenik">Sibenik</normalizedToken>
, northern Dalmatia, Croatia 
<emphasis id="C5AE81E17AA5FB12AAF0211239E7E00A" bold="true" pageId="0" pageNumber="21">A</emphasis>
habitus, dorsal view 
<emphasis id="7CA5CBBAFC78329F1332DBA8A8AFF29C" bold="true" pageId="0" pageNumber="21">B</emphasis>
habitus, lateral view 
<emphasis id="2525DDB68F5BFAD2E930A03ECC5CE36D" bold="true" pageId="0" pageNumber="21">C</emphasis>
head, dorsal view 
<emphasis id="46E2FE5977F5E2504CBBD46DE38B8E15" bold="true" pageId="0" pageNumber="21">D</emphasis>
microsculpture of head, dorsal view 
<emphasis id="DC9044856ECCBF69770241C90454C875" bold="true" pageId="0" pageNumber="21">E</emphasis>
right antenna, dorsal view 
<emphasis id="7AAD5B4E3FCBB618E5845CC2A7189A79" bold="true" pageId="0" pageNumber="21">F</emphasis>
pronotum, dorsal view 
<emphasis id="ED25983244BEFEAE77EF5CEC9F708188" bold="true" pageId="0" pageNumber="21">G</emphasis>
microsculpture of pronotum, dorsal view 
<emphasis id="6A672D1152368714C57EEBACFDB54E41" bold="true" pageId="0" pageNumber="21">H</emphasis>
mesoventral carina, lateral view 
<emphasis id="08A8508086401BD120B913876FCEB243" bold="true" pageId="0" pageNumber="21">I</emphasis>
mesoventrite, ventral view 
<emphasis id="3850304D08E58138FAC567831C39C30F" bold="true" pageId="0" pageNumber="21">J</emphasis>
scutellum, dorsal view 
<emphasis id="A4D9634F366FC86E99960F5F9E838E4C" bold="true" pageId="0" pageNumber="21">K</emphasis>
elytra, dorsal view 
<emphasis id="03A658E6DBC537335EB08296C0975AFC" bold="true" pageId="0" pageNumber="21">L</emphasis>
microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (
<emphasis id="0686F5AC25824C8E2E574168C8A25A5F" bold="true" pageId="0" pageNumber="21">A, B</emphasis>
); 0.5 mm (
<emphasis id="DDF02656A285C7D4E149C864B77A5AA0" bold="true" pageId="0" pageNumber="21">E, K</emphasis>
); 0.2 mm (
<emphasis id="753FB2B643131AA50D6F502A9FB98489" bold="true" pageId="0" pageNumber="21">C, F, H, I</emphasis>
); 0.1 mm (
<emphasis id="D95D045C8DE959DADC21A86651F64CCA" bold="true" pageId="0" pageNumber="21">J</emphasis>
); 0.05 mm (
<emphasis id="08B3E5585CB2947C16DCA234F6E488A1" bold="true" pageId="0" pageNumber="21">D, G, L</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C15574FE51DC733D0D930B71D42D04EB" pageId="0" pageNumber="21" type="etymology">
<paragraph id="C0ADB1D2E1B626147BF731A8655315E6" pageId="0" pageNumber="21">Etymology.</paragraph>
<paragraph id="4E2B00DD3947F0E91D5E40A625FE5534" pageId="0" pageNumber="21">
<taxonomicName id="D8B60C20855B1B7B312695D7E7622ADE" authorityName="Curcic, Vesovic, Vrbica &amp; Rađa" authorityYear="2023" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) coriniensis subsp. nonveilleri" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="coriniensis" subGenus="Spelaeobates" subSpecies="nonveilleri">Spelaeobates (Spelaeobates) coriniensis nonveilleri</taxonomicName>
ssp. nov. is named after late Prof. Dr Guido Nonveiller, a famous Serbian and Croatian biospeleologist and an excellent connoisseur of the subterranean beetle fauna of the Balkans.
</paragraph>
</subSubSection>
<subSubSection id="40033DD50044502289AAFB67DAF3C6BF" pageId="0" pageNumber="21" type="diagnosis">
<paragraph id="BAD952B68201DCFCD7A725E3BD0E18F6" pageId="0" pageNumber="21">Diagnosis.</paragraph>
<paragraph id="AA915396BB9B6723E0716076D1D4AEA8" pageId="0" pageNumber="21">
The new subspecies is morphologically closest to the nominotypic subspecies 
<taxonomicName id="E0502820EA5120AE5FA2F6DB76927DB4" authority="coriniensis" lsidName="S. (S.) coriniensis subsp. coriniensis" pageId="0" pageNumber="21" rank="subSpecies" species="coriniensis" subGenus="S." subSpecies="coriniensis">S. (S.) coriniensis coriniensis</taxonomicName>
ssp. nov., with which we compared it.
</paragraph>
<paragraph id="7315494B3EC1A4499429339EB7F3F004" pageId="0" pageNumber="21">
<taxonomicName id="F38BCECCF0A1EE7753141E824A3A0CC5" authorityName="Curcic, Vesovic, Vrbica &amp; Rađa" authorityYear="2023" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (S.) coriniensis subsp. nonveilleri" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="coriniensis" subGenus="S." subSpecies="nonveilleri">Spelaeobates (S.) coriniensis nonveilleri</taxonomicName>
ssp. nov. differs from 
<taxonomicName id="3263A723178E5C840BF8EDCFA4FAE4EF" authority="coriniensis" lsidName="S. (S.) coriniensis subsp. coriniensis" pageId="0" pageNumber="21" rank="subSpecies" species="coriniensis" subGenus="S." subSpecies="coriniensis">S. (S.) coriniensis coriniensis</taxonomicName>
ssp. nov. with respect to head shape (more elongate and narrower vs. shorter and wider), A6L/A3L (R 0.81-0.88 vs. R 0.88-1.00), shape of certain antennomeres (antennomeres I-V and IX-XI more elongate in first subpecies, whereas antennomeres VI-VIII more elongate in second subspecies), shape of pronotum (more elongate, widest before level of first third, lateral margins strongly convex anteriorly, with right hind angles vs. less elongate, widest at level of first third, lateral margins rounded anteriorly, with obtuse hind angles), PL/PW (R 1.24-1.38 vs. R 1.21-1.28), shape of process between mesocoxae on mesoventrite (sub-parallel vs. gradually narrowing apically), shape of elytra (more narrowed apically vs. less narrowed apically), shape of basal bulb in dorsal view (broadened distally vs. narrow, sub-parallel), and shape of spermatheca (less constricted medially vs. more constricted medially) (Table 
<tableCitation id="7CDF05D58B13683F57875C66F59AC703" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Linear measurements and morphometric ratios in Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov., S. (S.) coriniensis nonveilleri ssp. nov., and S. (S.) novaki. Values outside parentheses are mean values, while those inside parentheses are ranges." httpUri="http://table.plazi.org/id/A94F7FBCE48202F29A888BF18D2FCAD6" pageId="0" pageNumber="21" tableUuid="A94F7FBCE48202F29A888BF18D2FCAD6">1</tableCitation>
, Figs 
<figureCitation id="820F05CB0AF4EE122D8F473251EEC942" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis sp. nov. from the Jamurka (Rnjakusa II) Pit, village of Popovici, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure2" httpUri="https://binary.pensoft.net/fig/896581" pageId="0" pageNumber="21">2</figureCitation>
- 
<figureCitation id="46105F8FF77EFD27F886757151A2A8CD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bright-field images of certain morphological traits of Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov. from the Jamurka (Rnjakusa II) Pit, village of Popovici, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia (A, B) and S. (S.) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia (C-E) A, C PT female, spermatheca, lateral view B PT female, gonostyli, dorsal view D HT male, abdominal sternite IX (urite) E PT female, abdominal ventrite VIII. Scale bars: 0.05 mm (A, C); 0.10 mm (B, D, E)." figureDoi="10.3897/subtbiol.46.104548.figure6" httpUri="https://binary.pensoft.net/fig/896585" pageId="0" pageNumber="21">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8A44D1114887F0F1985A138D3E723D1D" pageId="0" pageNumber="21" type="description">
<paragraph id="94DEE035D55132EBCB600572CD833277" pageId="0" pageNumber="21">Description.</paragraph>
<paragraph id="72FC99A9CC941C3295B6028E88A5AAB9" pageId="0" pageNumber="21">Small-sized leptodirine. TL M 2.46 mm (2.42 mm in males, 2.52 mm in females), R 2.34-2.59 mm (2.34-2.52 mm in males, 2.47-2.59 mm in females).</paragraph>
<paragraph id="78895C1206278E75F7DFE4A3F5D59419" pageId="0" pageNumber="21">
<emphasis id="1F59BFF000937D3E6C37A585E265535C" bold="true" italics="true" pageId="0" pageNumber="21">Habitus</emphasis>
: Body shape leptodiroid (Fig. 
<figureCitation id="BF970DD66D5CBE34204B3CB7625AB2C8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, B</figureCitation>
), colour yellowish.
</paragraph>
<paragraph id="76FDFC130C9D229904A1DFC6D4EA6CB4" pageId="0" pageNumber="21">
<emphasis id="48499820D1C5FCD0ECE9BBB69BF0DF06" bold="true" italics="true" pageId="0" pageNumber="21">Integument</emphasis>
: Shiny, microsculptured both dorsally and ventrally (Fig. 
<figureCitation id="67B1ACFA5E0860BF789EDC53E452AADB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4C, D, F-L</figureCitation>
). Sparsely distributed deep punctures present on head, while densely distributed, fine and separated on both pronotum and elytra (Fig. 
<figureCitation id="DD42B0E316718F63EFDBC2AF36E5CE3C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4C, D, F, G, K, L</figureCitation>
). Entire body dorsally covered with yellow pubescence of short length (erect on head, while recumbent on both pronotum and elytra) (Fig. 
<figureCitation id="CA47CCDDCCE7366CA7F90D230D6AA001" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, B</figureCitation>
).
</paragraph>
<paragraph id="CAAC7E8655678219CF72A0755AB83C3F" pageId="0" pageNumber="21">
<emphasis id="BF55EACE539493422E6648931FE42606" bold="true" italics="true" pageId="0" pageNumber="21">Head</emphasis>
: More than one and a half times as long as wide (HL/HW M 1.57, R 1.53-1.61), with no differences in shape between males and females, anophthalmous, occipital carina in the shape of a curved concave line (Fig. 
<figureCitation id="8EC570E5A167C8583E63219ADD376AE4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, C</figureCitation>
). Head widest in first quarter or between first quarter and third. Frons roundly impressed between antennal insertions. Labrum transverse, with a few long setae. First maxillary palpomere of similar length and width, shorter than second maxillary palpomere. Maxillary palpomeres II and III of similar length (M3L/M2L M 0.97, R 0.85-1.09). Penultimate maxillary palpomere widened apically. Last maxillary palpomere short, thin, gradually narrowing apically. Antennae inserted in basal quarter of head, slender, narrow proximally (except for first two antennomeres, which are widened), slightly widened distally, longer in males, AL M 1.86 mm, R 1.77-1.94 mm (1.86-1.94 mm in males, 1.77-1.85 mm in females), reaching end of elytra in males (Fig. 
<figureCitation id="19449247C11B5ACBBC013CFDAD001126" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, B, E</figureCitation>
). Antennomeres I and II short and wide, second of which slightly longer and narrower. Following four antennomeres more slender and slightly longer than antennomere II. Antennomere III longer than adjacent antennomeres (A3L/A2L M 1.34, R 1.23-1.45; A3L/A4L M 1.21, R 1.07-1.31). Antennomeres VII, IX, and X quite dilated distally. Antennomere VIII relatively short and narrow, shorter and narrower than anatennomeres VII, IX, X, and XI. Ultimate antennomere thin, widened sub-distally, then narrowing apically, narrower than preceding (A11W/A10W M 0.84, R 0.67-1.00). Antennomere I shortest, while antennomeres IX and XI longest. Other ratios of length of certain antennomeres: A6L/A3L M 0.86, R 0.81-0.88; A8L/A3L M 0.75, R 0.65-0.81; A11L/A8L M 2.03, R 1.77-2.25.
</paragraph>
<paragraph id="1109C637684928A1FA1F92F312ACE24F" pageId="0" pageNumber="21">
<emphasis id="F1D8DEA73C566A3C6D5224B72F2B08E0" bold="true" italics="true" pageId="0" pageNumber="21">Prothorax</emphasis>
: Pronotum bell-shaped, elongate, longer than wide (PL/PW M 1.30, R 1.24-1.38; M 1.29, R 1.24-1.33 in males; M 1.31, R 1.28-1.38 in females), widest between first fourth and third, wider (HW/PW M 0.91, R 0.87-0.97) and shorter than head (PL/HL M 0.91, R 0.89-0.96) (Fig. 
<figureCitation id="34BCA16D7BA0578C93E606C338B09553" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, F</figureCitation>
). Lateral margins strongly convex anteriorly, then narrowing towards posterior end, markedly concave posteriorly. Pronotal base almost straight, slightly shorter than elytral base. PB/AM M 0.89, R 0.83-0.96. Anterior margin convex medially. Lateral margins and pronotal base rimmed. Fore pronotal angles weakly expressed, rounded, obtuse. Hind pronotal angles well-expressed, right, not protruding backwards. Pronotal disc moderately convex (Fig. 
<figureCitation id="890E405F2DD414DEDA6B05EF87BA96FA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4B</figureCitation>
).
</paragraph>
<paragraph id="9B9711C3AE65A4BB6DE98AFDDF63162C" pageId="0" pageNumber="21">
<emphasis id="0BAA6FD317F3FBA4C0C9B53E6AB433CB" bold="true" italics="true" pageId="0" pageNumber="21">Mesothorax</emphasis>
: Mesoventral carina very low, barely noticeable, with a few setae (Fig. 
<figureCitation id="E9A668D0305267BF8A7598C079C7AFD0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4H</figureCitation>
). No tooth, anterior and posterior margins observed. Mesoventrite with a long, sub-parallel process between mesocoxae (Fig. 
<figureCitation id="300903AE656B5F98F1F4044F1241BCDA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4I</figureCitation>
). Scutellum large, sub-triangular (Fig. 
<figureCitation id="E5E538137D42987F74F37F0A24A3A975" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4K, J</figureCitation>
).
</paragraph>
<paragraph id="4495208313ED9FCCEF766547842FAD51" pageId="0" pageNumber="21">
<emphasis id="88537E8B6A6AE47ED155C8177665756C" bold="true" italics="true" pageId="0" pageNumber="21">Metathorax</emphasis>
: Metaventrite with no carina.
</paragraph>
<paragraph id="60C92C824A9953F2539AF2991BC308E5" pageId="0" pageNumber="21">
<emphasis id="815E522F03B91F82C1F8A324BB4825E4" bold="true" italics="true" pageId="0" pageNumber="21">Elytra</emphasis>
: Broad, ovoid, almost of same width in males and females (EL/EW M 1.63, R 1.55-1.76 in males; M 1.66, R 1.55-1.73 in females), markedly wider than pronotum (EW/PW M 2.28, R 2.18-2.36) (Fig. 
<figureCitation id="F108A121C542B7ACB80C8A3DD1878110" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, K</figureCitation>
). Maximum width a little before middle. Lateral margins arcuate. Marginal furrows not visible from above. Shoulders barely noticeable, obtuse, covered by hind pronotal angles. Elytral disc markedly convex, steeply declining basally and gently declining apically in lateral view (Fig. 
<figureCitation id="04454E57AC38F92EC03EC717591CC6E8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4B</figureCitation>
). Parasutural stria absent. Elytral apex slightly attenuated, rounded. Pygidium covered by elytra.
</paragraph>
<paragraph id="54D5E16BF0FD8B8962C0F6722B412054" pageId="0" pageNumber="21">
<emphasis id="A4C5B4E0E7FEC2AD46D061C1FE0B6627" bold="true" italics="true" pageId="0" pageNumber="21">Legs</emphasis>
: Elongate and thin (Fig. 
<figureCitation id="CC0EE4E6857098E06E1D78DB9CEA3EB4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the morphological structures of PT male (A-G, J-L) and PT female (H, I) of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D microsculpture of head, dorsal view E right antenna, dorsal view F pronotum, dorsal view G microsculpture of pronotum, dorsal view H mesoventral carina, lateral view I mesoventrite, ventral view J scutellum, dorsal view K elytra, dorsal view L microsculpture of elytra, dorsal view. Scale bars: 1.0 mm (A, B); 0.5 mm (E, K); 0.2 mm (C, F, H, I); 0.1 mm (J); 0.05 mm (D, G, L)." figureDoi="10.3897/subtbiol.46.104548.figure4" httpUri="https://binary.pensoft.net/fig/896583" pageId="0" pageNumber="21">4A, B</figureCitation>
). Femora broadened basally, constricted in distal half. Tibiae slender, gently curved, gradually widening distally. Each protibia with a very fine comb over entire apical third of outer margin. Fore tarsi four-segmented in both sexes, only first protarsomere in males slightly dilated (P1W/P2W M 1.20, R 1.00-1.50). Tarsal claws thin, elongate, curved, pointed apically.
</paragraph>
<paragraph id="FA763B5401055347B2C7E568F2D8AE08" pageId="0" pageNumber="21">
<emphasis id="932590C68C0116C0E7A3854C0D3BD328" bold="true" italics="true" pageId="0" pageNumber="21">Male genitalia</emphasis>
: Aedeagus elongate, thin, small, well chitinised, almost the same as in the nominotypic subspecies (Fig. 
<figureCitation id="94621836A036F6589719B39BB55587B4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A, B</figureCitation>
). Median lobe in dorsal view straight, gradually narrowing apically, with a sharp apex, barely longer than parameres (Fig. 
<figureCitation id="F9DC80AE564C6D9AD648A8F3E185DC16" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A</figureCitation>
). Median lobe in lateral view quite flattened, curved basally, almost straight proximally, narrowing apically (Fig. 
<figureCitation id="9A54A5C1A7CFF448917D2C43FCAEBC42" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5B</figureCitation>
). Basal bulb small, broadened distally and bilobed in dorsal view (Fig. 
<figureCitation id="E477F257DCC37AC19DF170C0E30227B3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A</figureCitation>
), elongate and broadened basally in lateral view (Fig. 
<figureCitation id="0FC7E09A0DB7D1B44B9761A8161EF850" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5B</figureCitation>
). Tegmen wide from above (Fig. 
<figureCitation id="52C40EB53150FE42DEE429339374054A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A</figureCitation>
), in the shape of a ring around basal bulb (Fig. 
<figureCitation id="C1F5EFB71EB6F9AD98CBF08C2B28FD78" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5B</figureCitation>
). Parameres elongate, thin, arcuate, sub-apically curved exteriorly, each with a moderately broadened rounded apex in dorsal view (Fig. 
<figureCitation id="FF131BAA32FF265D6651B38B947BF391" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A</figureCitation>
), while straight, sub-parallel in lateral view (Fig. 
<figureCitation id="C5D1CB4F37E88F0B5B63E02A3151D5B7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5B</figureCitation>
). Each paramera carrying four apical close-set setae, three of which longer, while one shorter (Fig. 
<figureCitation id="D2127D00E266F8462EC0BEF46E6FF132" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A</figureCitation>
). No copulatory piece observed within inner sac (Fig. 
<figureCitation id="917B19A2BFF6E7BF072F4EDDF26E3259" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bright-field images of the male genitalia of HT of Spelaeobates (Spelaeobates) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21">5A, B</figureCitation>
).
</paragraph>
<caption id="9ADF14152417BA955790D85910FCB0A9" doi="10.3897/subtbiol.46.104548.figure5" httpUri="https://binary.pensoft.net/fig/896584" pageId="0" pageNumber="21" start="Figure 5" startId="F5">
<paragraph id="90B831BEB19F08060C495451599307F5" pageId="0" pageNumber="21">
<emphasis id="0FCC90520C883D6414AC3A27FFD34029" bold="true" pageId="0" pageNumber="21">Figure 5.</emphasis>
Bright-field images of the male genitalia of HT of 
<taxonomicName id="0C6631AB51D18469D9E6427C117FD949" authorityName="Curcic, Vesovic, Vrbica &amp; Rađa" authorityYear="2023" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) coriniensis subsp. nonveilleri" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="coriniensis" subGenus="Spelaeobates" subSpecies="nonveilleri">Spelaeobates (Spelaeobates) coriniensis nonveilleri</taxonomicName>
ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of 
<normalizedToken id="CF0DFFBD68B319468476BC449112C857" originalValue="Šibenik">Sibenik</normalizedToken>
, northern Dalmatia, Croatia 
<emphasis id="48DCC8F9557BB9715CB91415B7C9FEDB" bold="true" pageId="0" pageNumber="21">A</emphasis>
aedeagus, dorsal view 
<emphasis id="F11D03F346DF79CDDA15F13C9F1203C7" bold="true" pageId="0" pageNumber="21">B</emphasis>
aedeagus, lateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="B176FF35BD0B00ECB5087E451BDE67E6" pageId="0" pageNumber="21">
<emphasis id="E743A0478CE7159B488B38C9514033DA" bold="true" italics="true" pageId="0" pageNumber="21">Female genitalia</emphasis>
: Spermatheca small, chitinised, curved, somewhat constricted medially, spherical both basally and apically (Fig. 
<figureCitation id="4EE6177E95FDAA11B7EE89B44ED1D065" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bright-field images of certain morphological traits of Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov. from the Jamurka (Rnjakusa II) Pit, village of Popovici, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia (A, B) and S. (S.) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia (C-E) A, C PT female, spermatheca, lateral view B PT female, gonostyli, dorsal view D HT male, abdominal sternite IX (urite) E PT female, abdominal ventrite VIII. Scale bars: 0.05 mm (A, C); 0.10 mm (B, D, E)." figureDoi="10.3897/subtbiol.46.104548.figure6" httpUri="https://binary.pensoft.net/fig/896585" pageId="0" pageNumber="21">6C</figureCitation>
). Gonostyli short, straight, moderately broadened, gradually narrowing distally, pointed apically. Each gonostylus with one long apical seta.
</paragraph>
<caption id="96D27E03275B3D63B061AA48B3C7860E" doi="10.3897/subtbiol.46.104548.figure6" httpUri="https://binary.pensoft.net/fig/896585" pageId="0" pageNumber="21" start="Figure 6" startId="F6">
<paragraph id="6949B5ECEE06DD1247923B1A22A4BFB8" pageId="0" pageNumber="21">
<emphasis id="F2C2433B67C5556980DB95D32B504E05" bold="true" pageId="0" pageNumber="21">Figure 6.</emphasis>
Bright-field images of certain morphological traits of 
<taxonomicName id="5369E830B4489509E3F1C0D5C017B808" authority="coriniensis" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) coriniensis subsp. coriniensis" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="coriniensis" subGenus="Spelaeobates" subSpecies="coriniensis">Spelaeobates (Spelaeobates) coriniensis coriniensis</taxonomicName>
ssp. nov. from the Jamurka (
<normalizedToken id="161F1F5C013C9F0C69AB70A4DD4E5D0F" originalValue="Rnjakuša">Rnjakusa</normalizedToken>
II) Pit, village of 
<normalizedToken id="B3BB3E326148C595F5E33411667C8E2A" originalValue="Popovići">Popovici</normalizedToken>
, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia (
<emphasis id="69795FFC2E3820147460E0487C3C23F2" bold="true" pageId="0" pageNumber="21">A, B</emphasis>
) and 
<taxonomicName id="EF028B000F429B90C9C2384EB26BE1C9" lsidName="S. (S.) coriniensis subsp. nonveilleri" pageId="0" pageNumber="21" rank="subSpecies" species="coriniensis" subGenus="S." subSpecies="nonveilleri">S. (S.) coriniensis nonveilleri</taxonomicName>
ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of 
<normalizedToken id="F66D4B9563ECFA66051EAE6B9DC410C7" originalValue="Šibenik">Sibenik</normalizedToken>
, northern Dalmatia, Croatia (
<emphasis id="70836D779562765B06BAF8EE57BBF7EC" bold="true" pageId="0" pageNumber="21">C-E</emphasis>
) 
<emphasis id="5F7E1C82FEFB176CE1096D92A21DC5A6" bold="true" pageId="0" pageNumber="21">A, C</emphasis>
PT female, spermatheca, lateral view 
<emphasis id="7FB65885AB0D17F34786658A6D9199A5" bold="true" pageId="0" pageNumber="21">B</emphasis>
PT female, gonostyli, dorsal view 
<emphasis id="CD4EBA752550BE1B526517587ABE8C38" bold="true" pageId="0" pageNumber="21">D</emphasis>
HT male, abdominal sternite IX (urite) 
<emphasis id="2FC7FC7CB76F0E4896BFCCE0E3F18D9D" bold="true" pageId="0" pageNumber="21">E</emphasis>
PT female, abdominal ventrite VIII. Scale bars: 0.05 mm (
<emphasis id="A2E1821CF4E752B6063222C1862B28A0" bold="true" pageId="0" pageNumber="21">A, C</emphasis>
); 0.10 mm (
<emphasis id="929D73A501CF8FEF7DF2F548527CAA51" bold="true" pageId="0" pageNumber="21">B, D, E</emphasis>
).
</paragraph>
</caption>
<paragraph id="4D7CE0E1C45AEE5ED65A927BA29393C9" pageId="0" pageNumber="21">
<emphasis id="0E204016992504BE86687DF657B6125D" bold="true" italics="true" pageId="0" pageNumber="21">Male abdominal sternite IX (urite)</emphasis>
: Small, narrowing apically, sub-triangular (Fig. 
<figureCitation id="7047D1C94424AAC90246C33F76FDE287" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bright-field images of certain morphological traits of Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov. from the Jamurka (Rnjakusa II) Pit, village of Popovici, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia (A, B) and S. (S.) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia (C-E) A, C PT female, spermatheca, lateral view B PT female, gonostyli, dorsal view D HT male, abdominal sternite IX (urite) E PT female, abdominal ventrite VIII. Scale bars: 0.05 mm (A, C); 0.10 mm (B, D, E)." figureDoi="10.3897/subtbiol.46.104548.figure6" httpUri="https://binary.pensoft.net/fig/896585" pageId="0" pageNumber="21">6D</figureCitation>
).
</paragraph>
<paragraph id="24BA93ECD996521543D960DC052EE05D" pageId="0" pageNumber="21">
<emphasis id="6BD5F27EC489C766A496B1FFBA15B6FB" bold="true" italics="true" pageId="0" pageNumber="21">Female abdominal ventrite VIII</emphasis>
: Small, transverse, with no anterior process, hairy, especially posteriorly (Fig. 
<figureCitation id="EEAA3473C6AA491269337710FF303CBF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bright-field images of certain morphological traits of Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov. from the Jamurka (Rnjakusa II) Pit, village of Popovici, settlement of Gornji Karin, town of Benkovac, northern Dalmatia, Croatia (A, B) and S. (S.) coriniensis nonveilleri ssp. nov. from the Jezeranka Pit, village of Jezera, settlement of Tisno, island of Murter, city of Sibenik, northern Dalmatia, Croatia (C-E) A, C PT female, spermatheca, lateral view B PT female, gonostyli, dorsal view D HT male, abdominal sternite IX (urite) E PT female, abdominal ventrite VIII. Scale bars: 0.05 mm (A, C); 0.10 mm (B, D, E)." figureDoi="10.3897/subtbiol.46.104548.figure6" httpUri="https://binary.pensoft.net/fig/896585" pageId="0" pageNumber="21">6E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9D4CFF29CE4800977F283031827B8D9E" pageId="0" pageNumber="21" type="intrasubspecific variability">
<paragraph id="02CAC0F21648FF578D9737CF1C7A445A" pageId="0" pageNumber="21">Intrasubspecific variability.</paragraph>
<paragraph id="CF76C8105FE9187F1401E6F61477C7A1" pageId="0" pageNumber="21">
Some degree of intrasubspecific variability was noted in the new subspecies. It refers to the differences between the two known populations (one from the Jezeranka Pit, and the other from the 
<normalizedToken id="FBF00030612BCFEF6ADE499C01BFFECC" originalValue="Šušnjevača">Susnjevaca</normalizedToken>
Pit). The following differences were observed between the individuals of the two populations mentioned: (i) head is widest in the first quarter in individuals from the population from the Jezeranka Pit vs. head is widest between the first quarter and third in specimens from the 
<normalizedToken id="E941441A05FB2885678760857B198DCF" originalValue="Šušnjevača">Susnjevaca</normalizedToken>
Pit; (ii) antennomeres III, V, and IX, maxillary palpomere I, and elytra are more elongate in individuals from the Jezeranka Pit; (iii) antennomeres VI-VIII, X, and XI are more elongate in individuals from the 
<normalizedToken id="BFAF4937C0C683D3FC8D53FB7C89044B" originalValue="Šušnjevača">Susnjevaca</normalizedToken>
Pit; (iv) maxillary palpomere III is longer than maxillary palpomere II in individuals from the 
<normalizedToken id="14F9F4D48D4276298371B4534C24C11F" originalValue="Šušnjevača">Susnjevaca</normalizedToken>
Pit vs. maxillary palpomere III is shorter than maxillary palpomere II in individuals from the Jezeranka Pit. However, the identical shape of the aedeagus indicates that individuals from both populations belong to the same subspecies.
</paragraph>
</subSubSection>
<subSubSection id="2AF923B3F1ACCDA09B8E40C0422C0AAC" pageId="0" pageNumber="21" type="sexual dimorphism">
<paragraph id="265AD87E9E7D4DA1005CE7B931C2383E" pageId="0" pageNumber="21">Sexual dimorphism.</paragraph>
<paragraph id="6C03F00C94027E00C6CB573C4421ACFE" pageId="0" pageNumber="21">Some degree of sexual dimorphism was noted in this new subspecies. Namely, it was found that: (i) the females are slightly longer than the males; (ii) the antennae of the males are longer than those of the females; (iii) antennomeres VIII-X are more elongate in the males than in the females; (iv) first protarsomere is broader in the males than in the females.</paragraph>
</subSubSection>
<subSubSection id="105BA0D2EB5DBC4BC569D9E4E9EEA3B6" pageId="0" pageNumber="21" type="type locality">
<paragraph id="73A55D004FF718F41E681B35B3F1FD44" pageId="0" pageNumber="21">Type locality.</paragraph>
<paragraph id="BC321035302007FE3DBCFFF8D7DA3D3A" pageId="0" pageNumber="21">
Jezeranka Pit, village of Jezera, close to the settlement of Tisno and the city of 
<normalizedToken id="F2B4F477B2A414648E09433EE834E0A9" originalValue="Šibenik">Sibenik</normalizedToken>
, northern Dalmatia, Croatia.
</paragraph>
</subSubSection>
<subSubSection id="8E1F3519D3A0E7115340A615F48C2B92" pageId="0" pageNumber="21" type="distribution">
<paragraph id="4C2BB548648143F98A8609CD556D8ADA" pageId="0" pageNumber="21">Geographic distribution.</paragraph>
<paragraph id="5B60B7176EA581AA1437D3A8B05EF9CA" pageId="0" pageNumber="21">
This new subspecies is currently known from only two localities in the vicinity of the city of 
<normalizedToken id="A77B835997910A44C0A88EDF4DF96A0B" originalValue="Šibenik">Sibenik</normalizedToken>
, northern Dalmatia, Croatia - the Jezeranka Pit (type locality) in the village of Jezera, near the settlement of Tisno, and the 
<normalizedToken id="62A6771A274F2467090F4C6F8525F658" originalValue="Šušnjevača">Susnjevaca</normalizedToken>
Pit in the village of Banjevci. The first site is on the island (Murter), while the second is on the mainland. At the same time, this is the second official finding of a species of the genus 
<taxonomicName id="74E1447773A79767415BE21F5AF46E75" authorityName="J.Muller" authorityYear="1901" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="998D96199B075535EF213F705E5D662B" italics="true" pageId="0" pageNumber="21">Spelaeobates</emphasis>
</taxonomicName>
on the mainland. It is likely that the new subspecies also lives at other insular and non-insular subterranean sites in the surrounding area in northern Dalmatia.
</paragraph>
</subSubSection>
<subSubSection id="751DFACD0DF549D876F55626A7223BDB" pageId="0" pageNumber="21" type="bionomy and habitat">
<paragraph id="B1CDB391E209AA2EAA5A5C5DCBD98288" pageId="0" pageNumber="21">Bionomy and habitat.</paragraph>
<paragraph id="11AF94EE248348F2B174500E1AB75043" pageId="0" pageNumber="21">
Specimens of 
<taxonomicName id="968F73EEA2520ADD4847EA634FE913F2" lsidName="S. (S.) coriniensis subsp. nonveilleri" pageId="0" pageNumber="21" rank="subSpecies" species="coriniensis" subGenus="S." subSpecies="nonveilleri">S. (S.) coriniensis nonveilleri</taxonomicName>
ssp. nov. were collected manually from the walls and floor in the innermost parts of the Jezeranka and 
<normalizedToken id="099AA37E641352534F4265C9B6E272C0" originalValue="Šušnjevača">Susnjevaca</normalizedToken>
Pits, in places that were in complete darkness, with a high degree of humidity and the presence of trickling water.
</paragraph>
</subSubSection>
</treatment>
</document>