429 lines
58 KiB
XML
429 lines
58 KiB
XML
<document id="DD8D9E8D1593A4E7123D0C95E8D4022A" ID-DOI="10.11646/zootaxa.3734.5.1" ID-GBIF-Dataset="51673b52-5717-4920-846b-f1c560357734" ID-ISSN="1175-5326" ID-Zenodo-Dep="219026" ID-ZooBank="2C4411BF-233F-48F2-BA6F-423A9F8EE8B1" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460696948925" checkinUser="plazi" docAuthor="Vagalinski, Boyan, Lazányi, Eszter & Golovatch, Sergei" docDate="2013" docId="323D6308FFBEF308FF2FF8CC595FFCAE" docLanguage="en" docName="zt03734p520.pdf" docOrigin="Zootaxa 3734 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaphyllum (Parancistrum) curvifolii Verhoeff 1898" docType="treatment" docVersion="10" lastPageNumber="506" masterDocId="CE041B70FFBCF30DFFB8FFFF5865FFBB" masterDocTitle="Redescription of the subgenus Parancistrum Verhoeff, 1943, an eastern Mediterranean lineage of the millipede genus Megaphyllum Verhoeff, 1894 (Diplopoda: Julida: Julidae: Brachyiulini)" masterLastPageNumber="520" masterPageNumber="501" pageNumber="503" updateTime="1698353915693" updateUser="plazi">
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<mods:title id="AAA4F080C4238B1A951180CCCFD8FA78">Redescription of the subgenus Parancistrum Verhoeff, 1943, an eastern Mediterranean lineage of the millipede genus Megaphyllum Verhoeff, 1894 (Diplopoda: Julida: Julidae: Brachyiulini)</mods:title>
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<mods:namePart id="4575E6A9CFAEA410DB92EFCA84EECB1D">Vagalinski, Boyan</mods:namePart>
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<mods:namePart id="BA4E38A18DD1CE561081DD3ABAB8DC0F">Lazányi, Eszter</mods:namePart>
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<mods:roleTerm id="1C3819B928075A36AA0767C274DB05C9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A636015C9FC3F71128E153C661C334B5">Golovatch, Sergei</mods:namePart>
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<mods:typeOfResource id="F82477A6E4D26EFE2BDDD01EF221F3FC">text</mods:typeOfResource>
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<mods:title id="C14B23B60823A6BBB1B2E4D1242EDDB3">Zootaxa</mods:title>
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<mods:part id="FAE01821041AC5A2B8D8617DD04E1D2D">
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<mods:date id="5A80A11B64F031CA14033F60E23C1931">2013</mods:date>
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<mods:detail id="AC0BB496A26150B2C9766DB8700C61A0" type="volume">
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<mods:number id="39366C680B056166180AB78AFC59C351">3734</mods:number>
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<mods:identifier id="3B21EA55F4F4ACDA837FD335C536D912" type="DOI">10.11646/zootaxa.3734.5.1</mods:identifier>
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<mods:identifier id="5DD9E43B41963C0778FD0F97A8D8EB9B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="323D6308FFBEF308FF2FF8CC595FFCAE" ID-DOI="http://doi.org/10.5281/zenodo.5672394" ID-GBIF-Taxon="119581919" ID-Zenodo-Dep="5672394" LSID="urn:lsid:plazi:treatment:323D6308FFBEF308FF2FF8CC595FFCAE" httpUri="http://treatment.plazi.org/id/323D6308FFBEF308FF2FF8CC595FFCAE" lastPageId="5" lastPageNumber="506" pageId="2" pageNumber="503">
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<subSubSection id="F28E8195FFBEF30FFF2FF8CC5A28F868" pageId="2" pageNumber="503" type="nomenclature">
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<paragraph id="BA2BD21EFFBEF30FFF2FF8CC5B20F8F5" blockId="2.[151,837,1843,1903]" box="[151,837,1843,1870]" pageId="2" pageNumber="503">
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<heading id="E1636572FFBEF30FFF2FF8CC5B20F8F5" bold="true" box="[151,837,1843,1870]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
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<taxonomicName id="7D94A99DFFBEF30FFF2FF8CC5B20F8F5" ID-CoL="8KR83" authority="Verhoeff, 1898" authorityName="Verhoeff" authorityYear="1898" box="[151,837,1843,1870]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="curvifolii" subGenus="Parancistrum">
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<emphasis id="88E00E0CFFBEF30FFF2FF8CC5B20F8F5" bold="true" box="[151,837,1843,1870]" pageId="2" pageNumber="503">
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<emphasis id="88E00E0CFFBEF30FFF2FF8CC5A13F8F6" bold="true" box="[151,630,1843,1869]" italics="true" pageId="2" pageNumber="503">Megaphyllum (Parancistrum) curvifolii</emphasis>
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(Verhoeff, 1898)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="BA2BD21EFFBEF30FFF2FF8A9589DF8D4" blockId="2.[151,837,1843,1903]" box="[151,248,1878,1903]" pageId="2" pageNumber="503">
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<figureCitation id="22AFCE9BFFBEF30FFF2FF8A9589DF8D4" box="[151,248,1878,1903]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="2" pageNumber="503">Figs 1–9</figureCitation>
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</paragraph>
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<paragraph id="BA2BD21EFFBEF30FFF2FF8615A28F868" blockId="2.[151,717,1950,2034]" pageId="2" pageNumber="503">
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<taxonomicName id="7D94A99DFFBEF30FFF2FF8615A22F80F" authority="Verhoeff, 1898: 300" authorityName="Verhoeff" authorityPageNumber="300" authorityYear="1898" box="[151,583,1950,1972]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="curvifolii">
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<emphasis id="88E00E0CFFBEF30FFF2FF8615911F80F" box="[151,372,1950,1972]" italics="true" pageId="2" pageNumber="503">Brachyiulus curvifolii</emphasis>
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Verhoeff, 1898: 300
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</taxonomicName>
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–301, fig. 13.
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<taxonomicName id="7D94A99DFFBEF30FFF2FF8425A2CF868" authority="Verhoeff 1923: 131" authorityName="Verhoeff" authorityPageNumber="131" authorityYear="1923" box="[151,585,1981,2003]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="curvifolii">
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<emphasis id="88E00E0CFFBEF30FFF2FF8425911F868" box="[151,372,1981,2003]" italics="true" pageId="2" pageNumber="503">Brachyiulus curvifolii</emphasis>
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: Verhoeff 1923: 131
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F28E8195FFBEF30EFF2FF8235A3DFEB1" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="description">
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||
<paragraph id="BA2BD21EFFBEF30FFF2FF8235A3EF849" blockId="2.[151,717,1950,2034]" box="[151,603,2012,2034]" pageId="2" pageNumber="503">
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<taxonomicName id="7D94A99DFFBEF30FFF2FF8235A32F849" authority="Attems 1927: 224" authorityName="Attems" authorityPageNumber="224" authorityYear="1927" box="[151,599,2012,2034]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="curvifolii">
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<emphasis id="88E00E0CFFBEF30FFF2FF82359F4F849" box="[151,401,2012,2034]" italics="true" pageId="2" pageNumber="503">Chromatoiulus curvifolii</emphasis>
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: Attems 1927: 224
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="BA2BD21EFFBFF30EFF2FFF675A3EFF50" blockId="3.[151,843,152,266]" pageId="3" pageNumber="504">
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<taxonomicName id="7D94A99DFFBFF30EFF2FFF675A8DFF15" authority="Attems 1940: 302" authorityName="Attems" authorityPageNumber="302" authorityYear="1940" box="[151,744,152,174]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curvifolii" subGenus="Phauloiulus">
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<emphasis id="88E00E0CFFBFF30EFF2FFF675A47FF15" box="[151,546,152,174]" italics="true" pageId="3" pageNumber="504">Chromatoiulus (Phauloiulus) curvifolii</emphasis>
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: Attems 1940: 302
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</taxonomicName>
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.
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<taxonomicName id="7D94A99DFFBFF30EFF2FFF495A2BFF77" authority="Verhoeff, 1943: 231" authorityName="Verhoeff" authorityPageNumber="231" authorityYear="1943" box="[151,590,182,204]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="adanensis">
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<emphasis id="88E00E0CFFBFF30EFF2FFF495919FF77" box="[151,380,182,204]" italics="true" pageId="3" pageNumber="504">Brachyiulus adanensis</emphasis>
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Verhoeff, 1943: 231
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</taxonomicName>
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–232, figs 16–19,
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<emphasis id="88E00E0CFFBFF30EFCBDFF495B2EFF77" bold="true" box="[773,843,182,204]" pageId="3" pageNumber="504">syn. n.</emphasis>
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<taxonomicName id="7D94A99DFFBFF30EFF2FFF2A5A32FF50" authority="Enghoff 2006: 182" authorityName="Enghoff" authorityPageNumber="182" authorityYear="2006" box="[151,599,213,235]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="adanense">
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<emphasis id="88E00E0CFFBFF30EFF2FFF2A59E2FF50" box="[151,391,213,235]" italics="true" pageId="3" pageNumber="504">Megaphyllum adanense</emphasis>
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: Enghoff 2006: 182
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="BA2BD21EFFBFF30EFF2FFF0B5A3DFEB1" blockId="3.[151,843,152,266]" box="[151,600,244,266]" pageId="3" pageNumber="504">
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||
<taxonomicName id="7D94A99DFFBFF30EFF2FFF0B5A31FEB1" authority="Enghoff 2006: 183" authorityName="Enghoff" authorityPageNumber="183" authorityYear="2006" box="[151,596,244,266]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curvifolii">
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||
<emphasis id="88E00E0CFFBFF30EFF2FFF0B59E0FEB1" box="[151,389,244,266]" italics="true" pageId="3" pageNumber="504">Megaphyllum curvifolii</emphasis>
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: Enghoff 2006: 183
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="F28E8195FFBFF30EFF2FFEC95BBBFDEA" pageId="3" pageNumber="504" type="materials_examined">
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<paragraph id="BA2BD21EFFBFF30EFF2FFEC95BBBFDEA" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">
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||
<emphasis id="88E00E0CFFBFF30EFF2FFEC959E7FEF4" bold="true" box="[151,386,310,335]" pageId="3" pageNumber="504">Material examined.</emphasis>
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MNKB:
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<taxonomicName id="7D94A99DFFBFF30EFE56FEC95A80FEF5" box="[494,741,310,334]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curvifolii">
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<emphasis id="88E00E0CFFBFF30EFE56FEC95A80FEF5" box="[494,741,310,334]" italics="true" pageId="3" pageNumber="504">Brachyiulus curvifolii</emphasis>
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</taxonomicName>
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||
, ZMB 4379,
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<specimenCount id="AC921997FFBFF30EFC38FEC85BCEFEF5" box="[896,939,311,335]" pageId="3" pageNumber="504" type="generic">1 ♂</specimenCount>
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||
, 1 ♀
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<typeStatus id="652F6CBCFFBFF30EFC55FEC85C37FEF4" box="[1005,1106,311,335]" pageId="3" pageNumber="504" type="syntype">syntypes</typeStatus>
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, Verh.-slide 1199: gonopods, 1st, 2nd leg-pairs
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<typeStatus id="652F6CBCFFBFF30EFEDDFEA159F0FECD" box="[357,405,350,374]" pageId="3" pageNumber="504">type</typeStatus>
|
||
slide preparation,
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<collectingCountry id="C283928EFFBFF30EFD38FEA25AB4FECD" box="[640,721,349,374]" name="Turkey" pageId="3" pageNumber="504">Turkey</collectingCountry>
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||
, “Cilicien” [south-central
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||
<collectingCountry id="C283928EFFBFF30EFBAAFEA25C0DFECD" box="[1042,1128,349,374]" name="Turkey" pageId="3" pageNumber="504">Turkey</collectingCountry>
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||
], M. Holtz leg. NHMW:
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<taxonomicName id="7D94A99DFFBFF30EFF2FFE7E5AD0FE22" authority="Inv. No." authorityName="Inv. No." box="[151,693,385,409]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curvifolii" subGenus="Chromatoiulus">
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||
<emphasis id="88E00E0CFFBFF30EFF2FFE7E5A29FE22" box="[151,588,385,409]" italics="true" pageId="3" pageNumber="504">Brachyiulus (Chromatoiulus) curvifolii</emphasis>
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||
, Inv. No.
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||
</taxonomicName>
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||
3103,
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<specimenCount id="AC921997FFBFF30EFCB9FE7D5B4CFE22" box="[769,809,386,410]" pageId="3" pageNumber="504" type="generic">1 ♂</specimenCount>
|
||
, gonopods as slide preparation, 1 ♀, probably
|
||
<typeStatus id="652F6CBCFFBFF30EFA8BFE7D5DFDFE21" box="[1331,1432,386,410]" pageId="3" pageNumber="504" type="syntype">syntypes</typeStatus>
|
||
,
|
||
<collectingCountry id="C283928EFFBFF30EFF2FFE5A588DFE05" box="[151,232,421,446]" name="Turkey" pageId="3" pageNumber="504">Turkey</collectingCountry>
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||
, Cilicien. ZSM:
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<taxonomicName id="7D94A99DFFBFF30EFE1FFE595A5DFE06" box="[423,568,421,445]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="adanense">
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||
<emphasis id="88E00E0CFFBFF30EFE1FFE595A5DFE06" box="[423,568,421,445]" italics="true" pageId="3" pageNumber="504">M. adanense</emphasis>
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||
</taxonomicName>
|
||
,
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||
<specimenCount id="AC921997FFBFF30EFDF0FE595A16FE06" box="[584,627,422,446]" pageId="3" pageNumber="504" type="generic">1 ♂</specimenCount>
|
||
, questioned
|
||
<typeStatus id="652F6CBCFFBFF30EFCBCFE595B52FE05" box="[772,823,422,446]" pageId="3" pageNumber="504">type</typeStatus>
|
||
, Reg. Nr. A20060652, Asia,
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||
<collectingCountry id="C283928EFFBFF30EFB3DFE5A5CB3FE05" box="[1157,1238,421,446]" name="Turkey" pageId="3" pageNumber="504">Turkey</collectingCountry>
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||
, Adana; Reg. Nr. A
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<date id="CE2AF4DEFFBFF30EFF12FE33597FFE5E" box="[170,282,460,485]" pageId="3" pageNumber="504" value="1622-04-20">20041622</date>
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||
slide preparation: gonopods, gnathochilarium, antenna, 1st, 2nd, 3rd leg-pairs and some other legs; Reg. Nr. A
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||
<date id="CE2AF4DEFFBFF30EFF60FE0F592DFDB2" box="[216,328,496,521]" pageId="3" pageNumber="504" value="1623-04-20">20041623</date>
|
||
slide preparation: same body parts. MHNG:
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<taxonomicName id="7D94A99DFFBFF30EFCE1FE0F5C14FDB3" box="[857,1137,496,520]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curvifolii">
|
||
<emphasis id="88E00E0CFFBFF30EFCE1FE0F5C14FDB3" box="[857,1137,496,520]" italics="true" pageId="3" pageNumber="504">Chromatoiulus curvifolii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="AC921997FFBFF30EFB3AFE0F5CCAFDB3" box="[1154,1199,496,520]" pageId="3" pageNumber="504" type="generic">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="AC921997FFBFF30EFB78FE0F5D44FDB2" box="[1216,1313,496,521]" pageId="3" pageNumber="504" type="generic">1 subad.</specimenCount>
|
||
1 ♀, Asie Mineure [=
|
||
<collectingCountry id="C283928EFFBFF30EFE9AFDEB59C2FD96" box="[290,423,532,557]" name="Turkey" pageId="3" pageNumber="504">Asia Minor</collectingCountry>
|
||
] (probably ex. coll. J. Carl). ZMUC:
|
||
<taxonomicName id="7D94A99DFFBFF30EFCE2FDEA5B89FD97" box="[858,1004,532,556]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="adanense">
|
||
<emphasis id="88E00E0CFFBFF30EFCE2FDEA5B89FD97" box="[858,1004,532,556]" italics="true" pageId="3" pageNumber="504">M. adanense</emphasis>
|
||
</taxonomicName>
|
||
, 4 ♂♂,
|
||
<collectingCountry id="C283928EFFBFF30EFBF4FDEB5CF8FD96" box="[1100,1181,532,557]" name="Turkey" pageId="3" pageNumber="504">Turkey</collectingCountry>
|
||
, vil. Antalya, Irmasàn Geçidi,
|
||
<quantity id="7D6C7FFBFFBFF30EFF57FDC759E8FDEB" box="[239,397,568,593]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.52" metricValueMin="1.43" pageId="3" pageNumber="504" unit="m" value="1475.0" valueMax="1520.0" valueMin="1430.0">1430–1520 m</quantity>
|
||
,
|
||
<date id="CE2AF4DEFFBFF30EFE20FDC75A72FDEB" box="[408,535,568,592]" pageId="3" pageNumber="504" value="1982-04-28">28.IV.1982</date>
|
||
, A. Vigna Taglianti leg., H. Enghoff det.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F28E8195FFBFF308FF7FFDA358A0FF43" lastPageId="5" lastPageNumber="506" pageId="3" pageNumber="504" type="diagnosis">
|
||
<paragraph id="BA2BD21EFFBFF30EFF7FFDA35978FCBE" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">
|
||
<emphasis id="88E00E0CFFBFF30EFF7FFDA35924FDCE" bold="true" box="[199,321,604,629]" pageId="3" pageNumber="504">Diagnosis.</emphasis>
|
||
Differs from
|
||
<taxonomicName id="7D94A99DFFBFF30EFE61FDA25A06FDCF" box="[473,611,605,628]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="serratum">
|
||
<emphasis id="88E00E0CFFBFF30EFE61FDA25A06FDCF" box="[473,611,605,628]" italics="true" pageId="3" pageNumber="504">M. serratum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7D94A99DFFBFF30EFD20FDA25B7AFDCF" box="[664,799,605,628]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="turcicum">
|
||
<emphasis id="88E00E0CFFBFF30EFD20FDA25B7AFDCF" box="[664,799,605,628]" italics="true" pageId="3" pageNumber="504">M. turcicum</emphasis>
|
||
</taxonomicName>
|
||
by the smooth, rather than serrated, anterior opisthomere process; from
|
||
<taxonomicName id="7D94A99DFFBFF30EFEFAFD7E5995FD23" box="[322,496,640,664]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="tenenbaumi">
|
||
<emphasis id="88E00E0CFFBFF30EFEFAFD7E5995FD23" box="[322,496,640,664]" italics="true" pageId="3" pageNumber="504">M. tenenbaumi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7D94A99DFFBFF30EFD89FD7E5AA3FD23" box="[561,710,641,664]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="arcuatum" status="sp. nov.">
|
||
<emphasis id="88E00E0CFFBFF30EFD89FD7E5AA3FD23" box="[561,710,641,664]" italics="true" pageId="3" pageNumber="504">M. arcuatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="88E00E0CFFBFF30EFD6BFD7F5B72FD22" bold="true" box="[723,791,640,665]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="93D3B377FFBFF30EFD6BFD7F5B72FD22" box="[723,791,640,665]" pageId="3" pageNumber="504" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the apically gradually convex, rather than incised, promere; and from
|
||
<taxonomicName id="7D94A99DFFBFF30EFECFFD5A5A33FD07" box="[375,598,676,700]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="genezarethanum">
|
||
<emphasis id="88E00E0CFFBFF30EFECFFD5A5A33FD07" box="[375,598,676,700]" italics="true" pageId="3" pageNumber="504">M. genezarethanum</emphasis>
|
||
</taxonomicName>
|
||
by the posterior opisthomere process (
|
||
<emphasis id="88E00E0CFFBFF30EFBA0FD5A5C51FD07" bold="true" box="[1048,1076,677,700]" italics="true" pageId="3" pageNumber="504">pp</emphasis>
|
||
) being subequal to, rather than shorter than, solenomere (
|
||
<emphasis id="88E00E0CFFBFF30EFE6DFD375985FD64" bold="true" box="[469,480,712,735]" italics="true" pageId="3" pageNumber="504">s</emphasis>
|
||
), and by the smooth anterior solenomere protrusion (
|
||
<emphasis id="88E00E0CFFBFF30EFBC3FD375CC7FD64" bold="true" box="[1147,1186,712,735]" italics="true" pageId="3" pageNumber="504">asp</emphasis>
|
||
) devoid of plumose outgrowths.
|
||
</paragraph>
|
||
<paragraph id="BA2BD21EFFBFF30EFF7FFCEF5CB0FC03" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">
|
||
<emphasis id="88E00E0CFFBFF30EFF7FFCEF5911FC92" bold="true" box="[199,372,784,809]" pageId="3" pageNumber="504">Redescription.</emphasis>
|
||
Measurements:
|
||
<typeStatus id="652F6CBCFFBFF30EFD95FCEE5AE1FC92" box="[557,644,785,809]" pageId="3" pageNumber="504" type="syntype">syntype</typeStatus>
|
||
(MNKB) ♂ in stadium XI, body ring number 58+1+T, body length
|
||
<quantity id="7D6C7FFBFFBFF30EFA38FCEF58A0FCF7" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="504" unit="mm" value="27.0">27 mm</quantity>
|
||
, height
|
||
<quantity id="7D6C7FFBFFBFF30EFEA5FCCA5913FCF7" box="[285,374,821,845]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="504" unit="mm" value="2.1">2.1 mm</quantity>
|
||
;
|
||
<typeStatus id="652F6CBCFFBFF30EFE39FCCA59BDFCF6" box="[385,472,821,845]" pageId="3" pageNumber="504" type="syntype">syntype</typeStatus>
|
||
♀ (MNKB) in stadium XI, body ring number 57+1+T, body length
|
||
<quantity id="7D6C7FFBFFBFF30EFB68FCCB5D44FCF7" box="[1232,1313,820,845]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="504" unit="mm" value="35.0">35 mm</quantity>
|
||
, height
|
||
<quantity id="7D6C7FFBFFBFF30EFAC1FCCA58A3FCCB" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="504" unit="mm" value="2.3">2.3 mm</quantity>
|
||
; probable
|
||
<typeStatus id="652F6CBCFFBFF30EFE83FCA659F7FCCA" box="[315,402,857,881]" pageId="3" pageNumber="504" type="syntype">syntype</typeStatus>
|
||
♂ (NHMW) in stadium XII, body ring number 53+1+T, length
|
||
<quantity id="7D6C7FFBFFBFF30EFBDBFCA75CACFCCB" box="[1123,1225,856,881]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="3" pageNumber="504" unit="mm" value="35.1">35.1 mm</quantity>
|
||
, height
|
||
<quantity id="7D6C7FFBFFBFF30EFA9BFCA75D19FCCB" box="[1315,1404,856,880]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="504" unit="mm" value="2.1">2.1 mm</quantity>
|
||
; ♀ (NHMW) in stadium XII, body ring number 51+1+T, length
|
||
<quantity id="7D6C7FFBFFBFF30EFCE9FC835BDCFC2F" box="[849,953,892,917]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="3" pageNumber="504" unit="mm" value="35.6">35.6 mm</quantity>
|
||
, height
|
||
<quantity id="7D6C7FFBFFBFF30EFBADFC835C15FC2F" box="[1045,1136,892,917]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="504" unit="mm" value="2.6">2.6 mm</quantity>
|
||
; non-type ♂♂ (ZMUC) in stadia X–XII, body ring number (47+3+T)–(55+1+T), length
|
||
<quantity id="7D6C7FFBFFBFF30EFCF1FC5F5B8BFC03" box="[841,1006,928,953]" metricMagnitude="-2" metricUnit="m" metricValue="3.375" metricValueMax="4.08" metricValueMin="2.67" pageId="3" pageNumber="504" unit="mm" value="33.75" valueMax="40.8" valueMin="26.7">26.7–40.8 mm</quantity>
|
||
, height
|
||
<quantity id="7D6C7FFBFFBFF30EFBFFFC5F5CB4FC03" box="[1095,1233,928,952]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="3" pageNumber="504" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7–2.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA2BD21EFFBFF30EFF7FFC3B5D38FBBA" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">Coloration: Somewhat faded, overall brownish, with a narrow, blackish, mid-dorsal stripe and broad, short, dark, transverse stripes on prozona extending from ozopore level up to some distance from mid-dorsal stripe.</paragraph>
|
||
<paragraph id="BA2BD21EFFBFF30EFF7FFBF35D0EFAD2" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">
|
||
External structures: 20 labral setae. Gnathochilarium typically julid, promentum relatively large, nearly 2 times longer than broad. Collum smooth, with 3–4 short scratches near posterolateral corner. Prozona with numerous short, parallel scratches; metazona densely striated, with 10 striae in a square with sides equal to metazonal length just below ozopore level; posterior margin with one moderately dense row (ca 2/3 to equal to metazonal length) of setae. Ozopores placed just behind sutures between pro- and metazona, suture itself being straight in anterior body rings, slightly sinuous in posterior ones. Tarsi of midbody legs slightly shorter than tibiae. Anal valves with moderately dense, very long pilosity; no distinct rows of short setae along anal valves’ caudal margins. Pre-anal process long, straight, shorter and stouter in females; with a long, thin, somewhat down-turned, hyaline tip, slightly surpassed by the longest setae on anal valves. Subanal scale triangular, pointed, strongly protruding caudally behind anal valves’ contour by almost half of its length in males (
|
||
<figureCitation id="22AFCE9BFFBFF30EFCCDFAAF5BD2FAD3" box="[885,951,1360,1385]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Fig. 1</figureCitation>
|
||
); much shorter and blunter in females.
|
||
</paragraph>
|
||
<paragraph id="BA2BD21EFFBFF30EFF7FFA8B5AA7F9A6" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">
|
||
Male sexual characters: Mandibular stipites enlarged, protruding mostly anteriad, forming a distinct rounded anterior corner. Leg-pair 1 typical, parallel hooks with somewhat diverging tips; tarsal remnants absent. Pleurotergum 7 ventrally with rather short, rounded lobes at border between pro- and metazona, protruding mostly mesad behind opisthomeres (
|
||
<figureCitation id="22AFCE9BFFBFF30EFE5DFA1F5A2BFA43" box="[485,590,1504,1529]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Figs 2–3</figureCitation>
|
||
). Walking legs with pads on femur, postfemur and tibia; postfemoral and tibial ones undulate, femoral one small, flattened.
|
||
</paragraph>
|
||
<paragraph id="BA2BD21EFFBFF30EFF7FF9D75B92F83E" blockId="3.[151,1437,310,2033]" pageId="3" pageNumber="504">
|
||
Gonopods (
|
||
<figureCitation id="22AFCE9BFFBFF30EFEF1F9D759AEF9FB" box="[329,459,1576,1601]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Figs 2, 4–7</figureCitation>
|
||
): Promere (
|
||
<figureCitation id="22AFCE9BFFBFF30EFDF4F9D75AEBF9FA" box="[588,654,1576,1601]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Fig. 5</figureCitation>
|
||
and
|
||
<emphasis id="88E00E0CFFBFF30EFD7CF9D75AB0F984" bold="true" box="[708,725,1576,1599]" italics="true" pageId="3" pageNumber="504">P</emphasis>
|
||
in
|
||
<figureCitation id="22AFCE9BFFBFF30EFD40F9D75BE7F9FB" box="[760,898,1576,1601]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Figs 2 and 4</figureCitation>
|
||
) more than 3 times higher than broad, gradually narrowing distally, with a broadly rounded tip; a prominent median ridge (
|
||
<emphasis id="88E00E0CFFBFF30EFC6CF9B25BBAF9DF" bold="true" box="[980,991,1613,1636]" italics="true" pageId="3" pageNumber="504">r</emphasis>
|
||
) and a deep, narrow, median groove (
|
||
<emphasis id="88E00E0CFFBFF30EFA3CF9B25DF7F9DF" bold="true" box="[1412,1426,1613,1636]" italics="true" pageId="3" pageNumber="504">g</emphasis>
|
||
) on caudal face. Flagellum (
|
||
<emphasis id="88E00E0CFFBFF30EFE73F99059B9F933" bold="true" box="[459,476,1647,1672]" italics="true" pageId="3" pageNumber="504">fl</emphasis>
|
||
in
|
||
<figureCitation id="22AFCE9BFFBFF30EFDB9F98F5A26F932" box="[513,579,1648,1673]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Fig. 5</figureCitation>
|
||
) slightly longer than promere, thin in distal half. Opisthomere (
|
||
<figureCitation id="22AFCE9BFFBFF30EFAABF98F5DF1F932" box="[1299,1428,1648,1673]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Figs 4, 6–7</figureCitation>
|
||
) strongly elongated, basally curved posteriad, bearing two processes: a large, flat posterior process (
|
||
<emphasis id="88E00E0CFFBFF30EFB57F96A5D6EF917" bold="true" box="[1263,1291,1685,1708]" italics="true" pageId="3" pageNumber="504">pp</emphasis>
|
||
) subequal in length to solenomere (
|
||
<emphasis id="88E00E0CFFBFF30EFE2CF94759FAF974" bold="true" box="[404,415,1720,1743]" italics="true" pageId="3" pageNumber="504">s</emphasis>
|
||
) and bearing several blunt teeth at its apical margin; and a spiniform anterior process (
|
||
<emphasis id="88E00E0CFFBFF30EFACEF9475DF7F974" bold="true" box="[1398,1426,1720,1743]" italics="true" pageId="3" pageNumber="504">ap</emphasis>
|
||
) deviating mesad. Solenomere simple, tubular, with a short anterior protrusion (
|
||
<emphasis id="88E00E0CFFBFF30EFBA7F9225C23F94F" bold="true" box="[1055,1094,1757,1780]" italics="true" pageId="3" pageNumber="504">asp</emphasis>
|
||
). Several spine-like filaments (
|
||
<emphasis id="88E00E0CFFBFF30EFF18F90058D1F8A3" bold="true" box="[160,180,1791,1816]" italics="true" pageId="3" pageNumber="504">sf</emphasis>
|
||
) present basally along flagellum channel (
|
||
<emphasis id="88E00E0CFFBFF30EFD21F9005ACAF8A3" bold="true" box="[665,687,1791,1816]" italics="true" pageId="3" pageNumber="504">fc</emphasis>
|
||
), these looking more setiform apically. Gonopods
|
||
<emphasis id="88E00E0CFFBFF30EFB48F8FE5D52F8A3" box="[1264,1335,1793,1816]" italics="true" pageId="3" pageNumber="504">in situ</emphasis>
|
||
(
|
||
<figureCitation id="22AFCE9BFFBFF30EFAFFF8FF5DE8F8A3" box="[1351,1421,1792,1817]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Fig. 2</figureCitation>
|
||
): conspicuously protruding outside gonopod sinus, their tips reaching up to coxae 11 or 12; promere slightly outreaching solenomere; both promeres tightly connected to one another by transverse muscles and by a small chitinous knot between their bases, promere-opisthomere connection weaker.
|
||
</paragraph>
|
||
<paragraph id="BA2BD21EFFBFF308FF7FF86F58A0FF43" blockId="3.[151,1437,310,2033]" lastBlockId="5.[151,1436,151,789]" lastPageId="5" lastPageNumber="506" pageId="3" pageNumber="504">
|
||
Female sexual characters: First two leg-pairs slightly enlarged. Vulvae (
|
||
<figureCitation id="22AFCE9BFFBFF30EFB9BF86F5C0EF813" box="[1059,1131,1936,1961]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="3" pageNumber="504">Fig. 8</figureCitation>
|
||
) about as high as broad, cylindrical, somewhat tapering apically; bursa almost completely symmetrical, with a distinct apical margin, a soft median groove and an ellipsoid opening (
|
||
<emphasis id="88E00E0CFFBFF30EFD3AF8275AF5F854" bold="true" box="[642,656,2008,2031]" italics="true" pageId="3" pageNumber="504">o</emphasis>
|
||
) not exceeding the margin; operculum (
|
||
<emphasis id="88E00E0CFFBFF30EFBD4F8275CECF854" bold="true" box="[1132,1161,2008,2031]" italics="true" pageId="3" pageNumber="504">op</emphasis>
|
||
) subequal to bursa, its hyaline protrusions (
|
||
<emphasis id="88E00E0CFFB9F308FEC6FF6859C7FF0B" bold="true" box="[382,418,151,176]" italics="true" pageId="5" pageNumber="506">opl</emphasis>
|
||
) barely exceeding bursal ones (
|
||
<emphasis id="88E00E0CFFB9F308FCBDFF685B7EFF0B" bold="true" box="[773,795,151,176]" italics="true" pageId="5" pageNumber="506">bl</emphasis>
|
||
). Inner structures of vulva consisting of a large, globular, posterior ampulla (
|
||
<emphasis id="88E00E0CFFB9F308FED4FF4359EDFF68" bold="true" box="[364,392,188,211]" italics="true" pageId="5" pageNumber="506">pa</emphasis>
|
||
) joining a finger-shaped apodematic tube (
|
||
<emphasis id="88E00E0CFFB9F308FCD1FF435B1AFF68" bold="true" box="[873,895,188,211]" italics="true" pageId="5" pageNumber="506">at</emphasis>
|
||
) through a thin, spirally folded, connecting tube (
|
||
<emphasis id="88E00E0CFFB9F308FF18FF1F58D1FF4C" bold="true" box="[160,180,224,247]" italics="true" pageId="5" pageNumber="506">ct</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="EEEB8296FFB8F309FF2FF9265D30F84B" httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="4" pageNumber="505" targetBox="[227,1360,189,1728]" targetPageId="4">
|
||
<paragraph id="BA2BD21EFFB8F309FF2FF9265D30F84B" blockId="4.[151,1436,1752,2032]" pageId="4" pageNumber="505">
|
||
<emphasis id="88E00E0CFFB8F309FF2FF9265921F94B" bold="true" box="[151,324,1753,1776]" pageId="4" pageNumber="505">FIGURES 1–9.</emphasis>
|
||
<taxonomicName id="7D94A99DFFB8F309FEF4F9275A2CF94B" box="[332,585,1752,1776]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="curvifolii">
|
||
<emphasis id="88E00E0CFFB8F309FEF4F9275A2CF94B" box="[332,585,1752,1776]" italics="true" pageId="4" pageNumber="505">Megaphyllum curvifolii</emphasis>
|
||
</taxonomicName>
|
||
males (Figs 1–7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “Cilicien”, south-central Turkey:
|
||
<emphasis id="88E00E0CFFB8F309FDF4F9065A0FF8AB" bold="true" box="[588,618,1785,1808]" pageId="4" pageNumber="505">(1)</emphasis>
|
||
telson;
|
||
<emphasis id="88E00E0CFFB8F309FD72F9065A8DF8AB" bold="true" box="[714,744,1785,1808]" pageId="4" pageNumber="505">(2)</emphasis>
|
||
pleurotergum 7 and gonopods
|
||
<emphasis id="88E00E0CFFB8F309FBF7F9065CF3F8AB" box="[1103,1174,1785,1808]" italics="true" pageId="4" pageNumber="505">in situ</emphasis>
|
||
, lateral view;
|
||
<emphasis id="88E00E0CFFB8F309FA86F9065D38F8AB" bold="true" box="[1342,1373,1785,1808]" pageId="4" pageNumber="505">(3)</emphasis>
|
||
right pleurotergal protrusion, ventral view;
|
||
<emphasis id="88E00E0CFFB8F309FD8AF8E65A35F88B" bold="true" box="[562,592,1817,1840]" pageId="4" pageNumber="505">(4)</emphasis>
|
||
right gonopods, postero-mesal view;
|
||
<emphasis id="88E00E0CFFB8F309FC54F8E65C6EF88B" bold="true" box="[1004,1035,1817,1840]" pageId="4" pageNumber="505">(5)</emphasis>
|
||
connected promeres, posterior view;
|
||
<emphasis id="88E00E0CFFB8F309FF2FF8C658D0F8EB" bold="true" box="[151,181,1849,1872]" pageId="4" pageNumber="505">(6)</emphasis>
|
||
distal half of left opisthomere, mesal view;
|
||
<emphasis id="88E00E0CFFB8F309FD26F8C65AD8F8EB" bold="true" box="[670,701,1849,1872]" pageId="4" pageNumber="505">(7)</emphasis>
|
||
left opisthomere, antero-apical view;
|
||
<emphasis id="88E00E0CFFB8F309FBE7F8C65C18F8EB" bold="true" box="[1119,1149,1849,1872]" pageId="4" pageNumber="505">(8)</emphasis>
|
||
left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis);
|
||
<emphasis id="88E00E0CFFB8F309FB76F8A65C88F8CB" bold="true" box="[1230,1261,1881,1904]" pageId="4" pageNumber="505">(9)</emphasis>
|
||
penis, posterior view. Abbreviations:
|
||
<emphasis id="88E00E0CFFB8F309FE3AF88759F8F82B" bold="true" box="[386,413,1912,1936]" italics="true" pageId="4" pageNumber="505">ap</emphasis>
|
||
: anterior process,
|
||
<emphasis id="88E00E0CFFB8F309FDD3F8875AF5F82B" bold="true" box="[619,656,1912,1936]" italics="true" pageId="4" pageNumber="505">asp</emphasis>
|
||
: anterior protrusion of the solenomere,
|
||
<emphasis id="88E00E0CFFB8F309FBF4F8875C05F82B" bold="true" box="[1100,1120,1912,1936]" italics="true" pageId="4" pageNumber="505">at</emphasis>
|
||
: apodematic tube,
|
||
<emphasis id="88E00E0CFFB8F309FA8DF8875D2CF82B" bold="true" box="[1333,1353,1912,1936]" italics="true" pageId="4" pageNumber="505">bl</emphasis>
|
||
: bursal lamellar protrusions,
|
||
<emphasis id="88E00E0CFFB8F309FE3FF86759FFF80B" bold="true" box="[391,410,1944,1968]" italics="true" pageId="4" pageNumber="505">ct</emphasis>
|
||
: connecting tube,
|
||
<emphasis id="88E00E0CFFB8F309FDCAF8675AE2F80B" bold="true" box="[626,647,1944,1968]" italics="true" pageId="4" pageNumber="505">fc</emphasis>
|
||
: flagellum channel,
|
||
<emphasis id="88E00E0CFFB8F309FCCDF8675BE0F80B" bold="true" box="[885,901,1944,1968]" italics="true" pageId="4" pageNumber="505">fl</emphasis>
|
||
: flagellum,
|
||
<emphasis id="88E00E0CFFB8F309FBACF8675C44F80B" bold="true" box="[1044,1057,1944,1968]" italics="true" pageId="4" pageNumber="505">g</emphasis>
|
||
: median groove,
|
||
<emphasis id="88E00E0CFFB8F309FB56F8675C9EF80B" bold="true" box="[1262,1275,1944,1968]" italics="true" pageId="4" pageNumber="505">o</emphasis>
|
||
: opening,
|
||
<emphasis id="88E00E0CFFB8F309FAC2F8675DF1F80B" bold="true" box="[1402,1428,1944,1968]" italics="true" pageId="4" pageNumber="505">op</emphasis>
|
||
: operculum,
|
||
<emphasis id="88E00E0CFFB8F309FEAFF847595CF86B" bold="true" box="[279,313,1976,2000]" italics="true" pageId="4" pageNumber="505">opl</emphasis>
|
||
: operculum’s lamellar protrusions,
|
||
<emphasis id="88E00E0CFFB8F309FD02F8475AAFF86B" bold="true" box="[698,714,1976,2000]" italics="true" pageId="4" pageNumber="505">P</emphasis>
|
||
: promere,
|
||
<emphasis id="88E00E0CFFB8F309FCF9F8475B3EF86B" bold="true" box="[833,859,1976,2000]" italics="true" pageId="4" pageNumber="505">pa</emphasis>
|
||
: posterior ampulla,
|
||
<emphasis id="88E00E0CFFB8F309FB8EF8475C37F86B" bold="true" box="[1078,1106,1976,2000]" italics="true" pageId="4" pageNumber="505">ph</emphasis>
|
||
: posterior hump,
|
||
<emphasis id="88E00E0CFFB8F309FAABF8475D4BF86B" bold="true" box="[1299,1326,1976,2000]" italics="true" pageId="4" pageNumber="505">pp</emphasis>
|
||
: posterior process,
|
||
<emphasis id="88E00E0CFFB8F309FF4CF827589BF84B" bold="true" box="[244,254,2008,2032]" italics="true" pageId="4" pageNumber="505">r</emphasis>
|
||
: median ridge,
|
||
<emphasis id="88E00E0CFFB8F309FE1EF82759D5F84B" bold="true" box="[422,432,2008,2032]" italics="true" pageId="4" pageNumber="505">s</emphasis>
|
||
: solenomere,
|
||
<emphasis id="88E00E0CFFB8F309FDFEF8275A3CF84B" bold="true" box="[582,601,2008,2032]" italics="true" pageId="4" pageNumber="505">sf</emphasis>
|
||
: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="F28E8195FFB9F308FF7FFEFB5B05FE16" pageId="5" pageNumber="506" type="distribution">
|
||
<paragraph id="BA2BD21EFFB9F308FF7FFEFB5B05FE16" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="506">
|
||
<emphasis id="88E00E0CFFB9F308FF7FFEFB5905FEA6" bold="true" box="[199,352,260,285]" pageId="5" pageNumber="506">Distribution.</emphasis>
|
||
<collectingCountry id="C283928EFFB9F308FEDFFEFB59DEFEA6" box="[359,443,260,285]" name="Turkey" pageId="5" pageNumber="506">Turkey</collectingCountry>
|
||
: Cilicien (
|
||
<typeStatus id="652F6CBCFFB9F308FD94FEFB5A3BFEA7" box="[556,606,260,284]" pageId="5" pageNumber="506">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="7D94A99DFFB9F308FD66FEFA5B0CFEA7" box="[734,873,260,284]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="curvifolii">
|
||
<emphasis id="88E00E0CFFB9F308FD66FEFA5B0CFEA7" box="[734,873,260,284]" italics="true" pageId="5" pageNumber="506">M. curvifolii</emphasis>
|
||
</taxonomicName>
|
||
) (Verhoeff 1898) [between Mersin and Tarsus (cf. Hoffman & Lohmander1964)]; Adana, near the bay of Iskenderun (
|
||
<typeStatus id="652F6CBCFFB9F308FC35FED75BDAFEFB" box="[909,959,296,320]" pageId="5" pageNumber="506">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="7D94A99DFFB9F308FBF8FED65CB5FEFB" box="[1088,1232,296,320]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="adanense">
|
||
<emphasis id="88E00E0CFFB9F308FBF8FED65CB5FEFB" box="[1088,1232,296,320]" italics="true" pageId="5" pageNumber="506">M. adanense</emphasis>
|
||
</taxonomicName>
|
||
) (Verhoeff 1943); vil. Mersin, between Tarsus and Gülek, environs of Tasobagi; Mersin, Taurus, environs of Çamalan; vil. Mersin, environs of Çamliyayla; vil. Konya, environs of Mus; vil. Antalya, Irmasan Geçidi; vil. Antalya, Topraktepe; vil. Antalya, Korküteli (localities for
|
||
<taxonomicName id="7D94A99DFFB9F308FDB5FE6A5AF9FE17" box="[525,668,404,428]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="adanense">
|
||
<emphasis id="88E00E0CFFB9F308FDB5FE6A5AF9FE17" box="[525,668,404,428]" italics="true" pageId="5" pageNumber="506">M. adanense</emphasis>
|
||
</taxonomicName>
|
||
) (Enghoff 2006).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F28E8195FFB9F308FF7FFE48595FFCAE" pageId="5" pageNumber="506" type="discussion">
|
||
<paragraph id="BA2BD21EFFB9F308FF7FFE485A26FE4F" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="506">
|
||
<emphasis id="88E00E0CFFB9F308FF7FFE48595EFE6B" bold="true" box="[199,315,439,464]" pageId="5" pageNumber="506">Remarks.</emphasis>
|
||
Studying the
|
||
<typeStatus id="652F6CBCFFB9F308FE61FE475A6CFE6B" box="[473,521,440,464]" pageId="5" pageNumber="506">type</typeStatus>
|
||
material leaves no doubt that Verhoeff failed to recognize
|
||
<taxonomicName id="7D94A99DFFB9F308FB1AFE465D4AFE6B" box="[1186,1327,440,464]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="curvifolii">
|
||
<emphasis id="88E00E0CFFB9F308FB1AFE465D4AFE6B" box="[1186,1327,440,464]" italics="true" pageId="5" pageNumber="506">M. curvifolii</emphasis>
|
||
</taxonomicName>
|
||
when he, 45 years later, described
|
||
<taxonomicName id="7D94A99DFFB9F308FE15FE225A59FE4F" box="[429,572,476,500]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="adanense">
|
||
<emphasis id="88E00E0CFFB9F308FE15FE225A59FE4F" box="[429,572,476,500]" italics="true" pageId="5" pageNumber="506">M. adanense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA2BD21EFFB9F308FF7FFDFF595FFCAE" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="506">
|
||
Penes were only investigated in the non-type males. The only successfully dissected and drawn penis (
|
||
<figureCitation id="22AFCE9BFFB9F308FAF7FE005DF1FDA3" box="[1359,1428,511,536]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,269,1753,1776]" captionTargetBox="[227,1360,189,1728]" captionTargetId="figure@4.[227,1361,173,1728]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Megaphyllum curvifolii males (Figs 1 – 7; 9) from Irmasàn Geçidi, Turkey and syntype female (Fig. 8) from “ Cilicien ”, south-central Turkey: (1) telson; (2) pleurotergum 7 and gonopods in situ, lateral view; (3) right pleurotergal protrusion, ventral view; (4) right gonopods, postero-mesal view; (5) connected promeres, posterior view; (6) distal half of left opisthomere, mesal view; (7) left opisthomere, antero-apical view; (8) left vulva, mesal, slightly anterior view (receptaculum seen by transparency, but drawn with solid lines for better emphasis); (9) penis, posterior view. Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, o: opening, op: operculum, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like and setiform filaments. Scale bars of the drawings: 0.2 mm." httpUri="https://zenodo.org/record/219028/files/figure.png" pageId="5" pageNumber="506">Fig. 9</figureCitation>
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) was very soft, transparent and apparently hollow. It was elongated, straight, with two short parallel lobes and two short triangular lamellae—one was probably broken off during dissection. This rather unusual (compared to other examined species of the tribe Brachyiulini) shape may be the result either of a malformation or a normal alteration concerning the period immediately before or after copulation. Another dissected male seemed to have a very similar penis to the one observed in
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<taxonomicName id="7D94A99DFFB9F308FDF6FD4A5ABFFD77" box="[590,730,693,716]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="turcicum">
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<emphasis id="88E00E0CFFB9F308FDF6FD4A5ABFFD77" box="[590,730,693,716]" italics="true" pageId="5" pageNumber="506">M. turcicum</emphasis>
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</taxonomicName>
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(
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<figureCitation id="22AFCE9BFFB9F308FD56FD4B5BC2FD76" box="[750,935,692,717]" captionStart="FIGURES 35 – 40" captionStartId="12.[151,269,1625,1648]" captionTargetBox="[189,1400,208,1575]" captionTargetId="figure@12.[178,1409,195,1600]" captionTargetPageId="12" captionText="FIGURES 35 – 40. Megaphyllum turcicum male (35 – 39) and female (40) from near Aladağ, Turkey: (35) and (36) penis, caudal and lateral views, respectively; (37) right gonopods, mesal view; (38) left promere, posterior view; (39) left opisthomere, anterior, slightly mesal view; (40) right vulva, posterior, slightly mesal view (receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, bl: bursal lamellar protrusions, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, lh: lateral hump, mh: mesal hump, opl: operculum’s lamellar protrusions, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like filaments. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/219032/files/figure.png" pageId="5" pageNumber="506">Figs 35 and 36</figureCitation>
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), but it was accidentally lost before being depicted. However, we cannot say anything definitive regarding the penis of
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<taxonomicName id="7D94A99DFFB9F308FBFDFD265CBCFD4B" box="[1093,1241,728,752]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="curvifolii">
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<emphasis id="88E00E0CFFB9F308FBFDFD265CBCFD4B" box="[1093,1241,728,752]" italics="true" pageId="5" pageNumber="506">M. curvifolii</emphasis>
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</taxonomicName>
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pending further investigations.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |