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<document ID-DOI="http://doi.org/10.5281/zenodo.5522359" ID-GBIF-Dataset="c482bd26-6128-4a6f-8707-b44b82e77f10" ID-GBIF-Taxon="159223491" ID-Zenodo-Dep="5522359" checkinTime="1566393397064" checkinUser="jeremy" docAuthor="Leonora K. Gloyd" docDate="1980" docId="254D87B7FFC6FFD79F91FB65FE4AFB06" docLanguage="en" docName="Gloyd - 1980 - The taxonomic status of the genera Superlestes ABBYYpdf.pdf.imf" docOrigin="Occasional Papers of the Museum of Zoology, University of Michigan 694" docStyle="DocumentStyle{}" docTitle="Superlestes Williamson 1921" docType="treatment" docVersion="10" lastPageId="2" lastPageNumber="3" masterDocId="D974FFCFFFC6FFD59E3DFFB4FF88FFE1" masterDocTitle="The Taxonomic status of the genera Superlestes and Cyptolestes Williamson, 1921. (Odonata: Lestidae)." masterLastPageNumber="4" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1632327108397" updateUser="ExternalLinkService">
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<mods:title>The Taxonomic status of the genera Superlestes and Cyptolestes Williamson, 1921. (Odonata: Lestidae).</mods:title>
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<mods:namePart>Leonora K. Gloyd</mods:namePart>
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<mods:date>1980</mods:date>
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph blockId="0.[307,1299,1179,1325]" box="[428,1166,1233,1269]" pageId="0" pageNumber="1">
<heading allCaps="true" box="[428,1166,1233,1269]" centered="true" fontSize="12" level="2" pageId="0" pageNumber="1" reason="4">
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[428,722,1233,1269]" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[428,722,1233,1269]" italics="true" pageId="0" pageNumber="1">SUPERLESTES</emphasis>
</taxonomicName>
AND
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[865,1166,1233,1269]" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[865,1166,1233,1269]" italics="true" pageId="0" pageNumber="1">CYPTOLESTES</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="0.[307,1299,1179,1325]" box="[315,1298,1287,1323]" pageId="0" pageNumber="1">
<heading allCaps="true" box="[315,1298,1287,1323]" centered="true" fontSize="12" level="2" pageId="0" pageNumber="1" reason="5">
WILLIAMSON, 1921. (
<taxonomicName authorityName="Fabricius" authorityYear="1793" box="[797,1030,1287,1323]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">ODONATA</taxonomicName>
:
<taxonomicName authorityName="Calvert" authorityYear="1901" box="[1061,1281,1287,1323]" class="Insecta" family="Lestidae" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">LESTIDAE</taxonomicName>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[579,1009,1389,1425]" pageId="0" pageNumber="1" type="description">
<paragraph blockId="0.[579,1017,1388,1425]" box="[579,1009,1389,1425]" pageId="0" pageNumber="1">
By
<docAuthor box="[650,1009,1389,1425]" pageId="0" pageNumber="1">
Leonora K.
<emphasis box="[900,1009,1396,1424]" italics="true" pageId="0" pageNumber="1">Gloyd</emphasis>
</docAuthor>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph blockId="0.[151,1451,1541,1999]" pageId="0" pageNumber="1">
In 1943 when searching for a genus in which to place a very elegant and large specimen of a new lestine species from
<collectingCountry box="[1078,1212,1595,1631]" name="Mexico" pageId="0" pageNumber="1">Mexico</collectingCountry>
, I first compared it with Williamsons male of
<taxonomicName authorityName="Hagen in Selys" authorityYear="1862" box="[845,1210,1647,1683]" class="Insecta" family="Lestidae" genus="Superlestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis box="[845,1040,1647,1683]" italics="true" pageId="0" pageNumber="1">Superlestes</emphasis>
exoletus
</taxonomicName>
(Selys) and with the
<typeStatus box="[340,496,1699,1735]" pageId="0" pageNumber="1">holotype</typeStatus>
and
<typeStatus box="[616,773,1699,1735]" pageId="0" pageNumber="1">paratype</typeStatus>
males of
<taxonomicName authority="Williamson" authorityName="Williamson" authorityYear="1921" box="[991,1443,1699,1735]" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tuberalatus">
<emphasis box="[991,1443,1699,1735]" italics="true" pageId="0" pageNumber="1">Cyptolestes tuberculatus</emphasis>
</taxonomicName>
Williamson before checking it against males of
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[1093,1445,1750,1786]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grandis">
<emphasis box="[1093,1287,1750,1786]" italics="true" pageId="0" pageNumber="1">Archilestes</emphasis>
grandis
</taxonomicName>
(Rambur). The specimens of these four species were all very similar in general appearance, but were readily distinguished by color patterns and their abdominal appendages. The new species
<taxonomicName authorityName="Gloyd" authorityYear="1944" box="[1188,1303,1908,1944]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="regalis">
<emphasis box="[1188,1303,1908,1944]" italics="true" pageId="0" pageNumber="1">regalis</emphasis>
</taxonomicName>
seemed best placed in the genus
<taxonomicName authority="(Gloyd, 1944)" authorityName="Gloyd" authorityYear="1944" baseAuthorityName="Gloyd" baseAuthorityYear="1944" box="[623,1097,1960,1996]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[623,817,1960,1996]" italics="true" pageId="0" pageNumber="1">Archilestes</emphasis>
(
<bibRefCitation author="Gloyd, Leonora K." box="[860,1087,1960,1996]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="0" pageNumber="1" pagination="1 - 4" part="483" refId="ref942" refString="Gloyd, Leonora K. 1944. A new species of Archilestes from Mexico (Odonata: Lestinae). Occ. Pap. Mus. Zool. Univ. Mich. 483: 1 - 4." title="A new species of Archilestes from Mexico (Odonata: Lestinae)" type="journal article" year="1944">Gloyd, 1944</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1" type="discussion">
<paragraph blockId="0.[150,1448,2043,2345]" lastBlockId="1.[243,1549,355,925]" lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1">
The monotypic genera
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[623,818,2046,2082]" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[623,818,2046,2082]" italics="true" pageId="0" pageNumber="1">Superlestes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[920,1118,2046,2082]" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[920,1118,2046,2082]" italics="true" pageId="0" pageNumber="1">Cyptolestes</emphasis>
</taxonomicName>
described by Mr. Williamson in 1921 were distinguished from
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1026,1133,2098,2134]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[1026,1133,2098,2134]" italics="true" pageId="0" pageNumber="1">Lestes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Selys" authorityYear="1862" box="[1250,1442,2098,2134]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[1250,1442,2098,2134]" italics="true" pageId="0" pageNumber="1">Archilestes</emphasis>
</taxonomicName>
primarily by a detailed study of the venation of one male of the former and three males and two females of the latter. In view of the venational variation of certain supposedly diagnostic generic characters noted in a series of specimens of several species of
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1183,1290,2306,2342]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis box="[1183,1290,2306,2342]" italics="true" pageId="0" pageNumber="1">Lestes</emphasis>
</taxonomicName>
(
<bibRefCitation author="Gloyd, Leonora K." journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1" pagination="1 - 4" part="483" refId="ref942" refString="Gloyd, Leonora K. 1944. A new species of Archilestes from Mexico (Odonata: Lestinae). Occ. Pap. Mus. Zool. Univ. Mich. 483: 1 - 4." title="A new species of Archilestes from Mexico (Odonata: Lestinae)" type="journal article" year="1944">Gloyd 1944: 8</bibRefCitation>
), data on only four males and two females would hardly seem adequate to indicate how constant or reliable the differences in the area of wings distal to the nodus are. Although venational differences have been and still are used for the primary diagnoses of genera, we now know that some venational characters often employed as criteria may vary within a species as well as between species within a genus, or may be of specific value only. All this is well exemplified within the genus
<emphasis box="[443,550,728,764]" italics="true" pageId="1" pageNumber="2">
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[443,539,728,764]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Argia</taxonomicName>
.
</emphasis>
To give generic rank to species on venational data alone can give a false impression of the degree of evolutionary separation. Other characters may show a very close relationship indicative of generic unity.
</paragraph>
<paragraph blockId="1.[243,1548,965,1425]" pageId="1" pageNumber="2">
A comparison of the abdominal appendages of the males of
<taxonomicName authorityName="Hagen in Selys" authorityYear="1862" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis box="[248,338,1021,1057]" italics="true" pageId="1" pageNumber="2">Superlestes</emphasis>
exoletus
</taxonomicName>
(Selys),
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[683,1245,1021,1057]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="grandis">
<emphasis box="[683,877,1021,1057]" italics="true" pageId="1" pageNumber="2">Archilestes</emphasis>
grandis (Rambur)
</taxonomicName>
, and
<taxonomicName authorityName="Gloyd" authorityYear="1944" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="regalis">
<emphasis box="[1365,1543,1021,1057]" italics="true" pageId="1" pageNumber="2">A. regalis</emphasis>
Gloyd
</taxonomicName>
shows a striking resemblance (see
<bibRefCitation author="Williamson, E. B." box="[1075,1395,1075,1111]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="1" pageNumber="2" pagination="1 - 9" part="96" refId="ref980" refString="Williamson, E. B. 1921. Two new Neotropical genera of Lestinae (Odonata). Occ. Pap. Mus. Zool. Univ. Mich. 96: 1 - 9." title="Two new Neotropical genera of Lestinae (Odonata)" type="journal article" year="1921">Williamson 1921</bibRefCitation>
, Pl.
<collectionCode box="[1502,1535,1075,1111]" pageId="1" pageNumber="2">II</collectionCode>
, figs. 8-9 and
<bibRefCitation author="Gloyd, Leonora K." box="[495,710,1127,1163]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="1" pageNumber="2" pagination="1 - 4" part="483" refId="ref942" refString="Gloyd, Leonora K. 1944. A new species of Archilestes from Mexico (Odonata: Lestinae). Occ. Pap. Mus. Zool. Univ. Mich. 483: 1 - 4." title="A new species of Archilestes from Mexico (Odonata: Lestinae)" type="journal article" year="1944">Gloyd 1944</bibRefCitation>
, Pl.
<collectionCode box="[807,823,1127,1163]" pageId="1" pageNumber="2">I</collectionCode>
, figs. 7-9). The appendages of
<taxonomicName authority="Williamson" authorityName="Williamson" authorityYear="1921" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tuberalatus">
<emphasis box="[246,586,1179,1215]" italics="true" pageId="1" pageNumber="2">
<emphasis box="[1428,1548,1127,1163]" italics="true" pageId="1" pageNumber="2">Cyptolestes</emphasis>
tuberculatus
</emphasis>
</taxonomicName>
Williamson (1921, Pl.
<collectionCode box="[1057,1090,1179,1215]" pageId="1" pageNumber="2">II</collectionCode>
, figs. 10-11) differ considerably from those of the above three species but the difference is not greater than that found among species of
<emphasis box="[1151,1267,1283,1319]" italics="true" pageId="1" pageNumber="2">
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1151,1256,1283,1319]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Lestes</taxonomicName>
.
</emphasis>
The hamules (see
<collectionCode box="[326,371,1335,1371]" pageId="1" pageNumber="2">PL</collectionCode>
<collectionCode box="[395,411,1335,1371]" pageId="1" pageNumber="2">I</collectionCode>
, fig. 7 for
<taxonomicName authorityName="Gloyd" authorityYear="1944" box="[621,972,1335,1371]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="regalis">
<emphasis box="[621,736,1335,1371]" italics="true" pageId="1" pageNumber="2">regalis</emphasis>
<bibRefCitation author="Gloyd, Leonora K." box="[758,972,1335,1371]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="1" pageNumber="2" pagination="1 - 4" part="483" refId="ref942" refString="Gloyd, Leonora K. 1944. A new species of Archilestes from Mexico (Odonata: Lestinae). Occ. Pap. Mus. Zool. Univ. Mich. 483: 1 - 4." title="A new species of Archilestes from Mexico (Odonata: Lestinae)" type="journal article" year="1944">Gloyd 1944</bibRefCitation>
</taxonomicName>
), however, are very similar in all four species as well as in
<taxonomicName authorityName="McLachlan" authorityYear="1895" box="[780,1436,1387,1423]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="californica">
<emphasis box="[780,1183,1387,1423]" italics="true" pageId="1" pageNumber="2">Archilestes californica</emphasis>
MacLachlan.
</taxonomicName>
</paragraph>
<paragraph blockId="1.[238,1542,1464,1924]" pageId="1" pageNumber="2">
In the females of
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[616,813,1468,1504]" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[616,813,1468,1504]" italics="true" pageId="1" pageNumber="2">Superlestes</emphasis>
</taxonomicName>
exoletus,
<taxonomicName authority="Williamson" authorityName="Williamson" authorityYear="1921" box="[1010,1454,1468,1504]" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tuberalatus">
<emphasis box="[1010,1454,1468,1504]" italics="true" pageId="1" pageNumber="2">Cyptolestes tuberculatus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="McLachlan" authorityYear="1895" box="[243,651,1520,1556]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="californica">
<emphasis box="[243,651,1520,1556]" italics="true" pageId="1" pageNumber="2">Archilestes californica</emphasis>
</taxonomicName>
and grandis, the valves of the ovipositor are robust and the teeth on the lower margin of each are large, stout and few in number, unlike any I have observed in species of
<emphasis box="[1326,1441,1624,1660]" italics="true" pageId="1" pageNumber="2">
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1326,1430,1624,1660]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Lestes</taxonomicName>
.
</emphasis>
The female of
<emphasis box="[432,614,1676,1712]" italics="true" pageId="1" pageNumber="2">
<taxonomicName authorityName="Gloyd" authorityYear="1944" box="[432,603,1676,1712]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="regalis">A. regalis</taxonomicName>
,
</emphasis>
collected by George H. and Alice F. Beatty and as yet undescribed, also has large teeth as wide at base as high on the lower margin of each valve of the ovipositor according to inform ation kindly provided by Mr. Beatty (in litt.
<date box="[950,1193,1832,1868]" pageId="1" pageNumber="2" value="1980-05-15">May 15, 1980</date>
). Thus the females of these five species are similar in this respect.
</paragraph>
<paragraph blockId="1.[238,1542,1962,2316]" pageId="1" pageNumber="2">
In the species of
<emphasis box="[610,1043,1966,2002]" italics="true" pageId="1" pageNumber="2">
<taxonomicName authorityName="Selys" authorityYear="1862" box="[610,803,1966,2002]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Archilestes</taxonomicName>
,
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[836,1031,1966,2002]" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Superlestes</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[1156,1354,1966,2002]" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyptolestes</taxonomicName>
, the proportions of the quadrangle of the wings (
<bibRefCitation author="Williamson, E. B." box="[1018,1334,2018,2054]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="1" pageNumber="2" pagination="1 - 9" part="96" refId="ref980" refString="Williamson, E. B. 1921. Two new Neotropical genera of Lestinae (Odonata). Occ. Pap. Mus. Zool. Univ. Mich. 96: 1 - 9." title="Two new Neotropical genera of Lestinae (Odonata)" type="journal article" year="1921">Williamson 1921</bibRefCitation>
, Pl.
<collectionCode box="[1432,1445,2018,2054]" pageId="1" pageNumber="2">I</collectionCode>
and
<bibRefCitation author="Gloyd, Leonora K." box="[239,454,2070,2106]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="1" pageNumber="2" pagination="1 - 4" part="483" refId="ref942" refString="Gloyd, Leonora K. 1944. A new species of Archilestes from Mexico (Odonata: Lestinae). Occ. Pap. Mus. Zool. Univ. Mich. 483: 1 - 4." title="A new species of Archilestes from Mexico (Odonata: Lestinae)" type="journal article" year="1944">Gloyd 1944</bibRefCitation>
, Pl.
<collectionCode box="[553,569,2070,2106]" pageId="1" pageNumber="2">I</collectionCode>
), the metallic coloration, the structural characters of both male and females noted above, and their large size all indicate a compact group. Accordingly,
<taxonomicName authorityName="Williamson" authorityYear="1921" box="[807,1319,2175,2211]" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[807,1003,2175,2211]" italics="true" pageId="1" pageNumber="2">Superlestes</emphasis>
Williamson 1921
</taxonomicName>
and
<taxonomicName authorityName="Williamson" authorityYear="1921" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[238,329,2228,2264]" italics="true" pageId="1" pageNumber="2">Cyptolestes</emphasis>
Williamson 1921
</taxonomicName>
are hereby designated
<emphasis box="[1091,1351,2228,2264]" italics="true" pageId="1" pageNumber="2">new synonyms</emphasis>
of
<taxonomicName authorityName="Selys" authorityYear="1862" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="1" pageNumber="2">Archilestes</emphasis>
Selys 1862
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[150,1448,330,1255]" pageId="2" pageNumber="3">
Of the five species of
<emphasis box="[598,1008,334,370]" italics="true" pageId="2" pageNumber="3">
<taxonomicName authorityName="Selys" authorityYear="1862" box="[598,790,334,370]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Archilestes</taxonomicName>
,
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[821,1008,334,370]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="grandis">A. grandis</taxonomicName>
</emphasis>
has the most extensive range of distribution and a record of recent expansion. For many years, in the United States it was known only from the Pacific Coast states of Washington and California, the southwestern states of Arizona, Utah, and east to Texas, Oklahoma and Kansas. The first record east of the Mississippi River was in 1927 when it was collected in Ohio by Mr. E. B.
<bibRefCitation author="Williamson, E. B." box="[411,760,645,681]" journalOrPublisher="Ent. News" pageId="2" pageNumber="3" pagination="63 - 64" part="43" refId="ref1016" refString="Williamson, E. B. 1931. Archilestes grandis (Odonata: Agrionidae). Ent. News, 43: 63 - 64." title="Archilestes grandis (Odonata: Agrionidae)" type="journal article" year="1931">Williamson (1931)</bibRefCitation>
and it has now extended its range to states bordering the Atlantic coast, and to South Dakota. In Mexico it is known from Baja California and in the states of Chihuahua, Guanajuato, Guerrero, Jalisco, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Tabasco, and Veracruz. In Central America it is known from Costa Rica, Guatemala, Honduras, and Panama. In South America it has been reported only from Colombia and Venezuela.
<taxonomicName authorityName="McLachlan" authorityYear="1895" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="californica">
<emphasis italics="true" pageId="2" pageNumber="3">A. California</emphasis>
</taxonomicName>
occurs only in the Pacific Coast states and south into Baja California. The other three species are each recorded in the literature from one locality only:
<taxonomicName authorityName="Gloyd" authorityYear="1944" box="[713,825,1118,1150]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="regalis">regalis</taxonomicName>
from San Luis Potosi, Mexico;
<taxonomicName authority="Williamson" authorityName="Williamson" authorityYear="1921" box="[153,374,1170,1202]" class="Insecta" family="Lestidae" genus="Cyptolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tuberalatus">
<emphasis box="[153,374,1170,1202]" italics="true" pageId="2" pageNumber="3">tuberculatus</emphasis>
</taxonomicName>
from Carabobo, Venezuela; and
<taxonomicName authorityName="Hagen in Selys" authorityYear="1862" box="[1056,1200,1170,1202]" class="Insecta" family="Lestidae" genus="Superlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis box="[1056,1200,1170,1202]" italics="true" pageId="2" pageNumber="3">exoletus</emphasis>
</taxonomicName>
from Santa Catarina, Brazil.
</paragraph>
</subSubSection>
</treatment>
</document>