treatments-xml/data/1D/36/87/1D3687F8A77FA679FF62FA23FBB07166.xml

<document id="CB26002D37BF2595F1873B377E3C8CE0" ID-CLB-Dataset="22701" ID-DOI="10.3897/zookeys.7.117" ID-GBIF-Dataset="7a12aaad-a568-4920-91cf-af3e9407eadc" ID-ISSN="1313–2970" ID-Zenodo-Dep="576436" ID-ZooBank="67B4D2EC-2C6D-4226-847D-0BA2B2777AE3" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1587986763134" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2009" docId="1D3687F8A77FA679FF62FA23FBB07166" docLanguage="en" docName="ZK_article_1970.pdf" docOrigin="ZooKeys 7 (7)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus similis Golovatch, Geoffroy, Mauries &amp; VandenSpiegel 2009, sp. n." docType="treatment" docUuid="9CE9BBFF-66DF-42BC-B4E8-C44E2C71F972" docUuidSource="ZooBank" docVersion="7" lastPageNumber="29" masterDocId="E10FFF80A766A664FFEEFF9FFF8E742C" masterDocTitle="Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="26" updateTime="1693525343643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="0A298A242E0335A92F2BE7D361572F89">Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="769CA288E501AAC2F03E728AF1749385">Golovatch, Sergei</mods:namePart>
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<mods:affiliation id="E2A990E2454FBAD51788345DDF89D211">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="8656DC94A0B761332930CF63E4EFE219">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:affiliation id="646837854358A1E6046E2ED642D1C192">Muséum national d’Histoire naturelle, Département Ecologie &amp; Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, 4, avenue du Petit Château, F- 91800 Brunoy, France</mods:affiliation>
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<mods:affiliation id="B381335DAC1594A8C544319AD2582826">Muséum national d’Histoire naturelle, Département Systématique et Evolution, USM 602, Section Arthropodes Case Postale n ° 53, 61 rue Buffon F- 75231 Paris, France</mods:affiliation>
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<mods:namePart id="6DDD6A8ECC9282B700D37204EACC7AA2">VandenSpiegel, Didier</mods:namePart>
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<mods:affiliation id="2AC23F5FCC178014886336AE11FA44EC">Musée Royal de l’Afrique centrale, B- 3080 Tervuren, Belgium</mods:affiliation>
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<emphasis id="A7EBEAFCA77FA67DFF62FA23FEC671FB" box="[140,328,1468,1495]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Eutrichodesmus</emphasis>
similis
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Golovatch, Geoffroy, Mauriès &amp; VandenSpiegel
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, 
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<figureCitation id="0DA42A6BA77FA67DFF62F99CFE867232" box="[140,264,1539,1566]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStart-2="Figure 20" captionStart-3="Figure 21" captionStartId-0="27.[140,212,1161,1183]" captionStartId-1="28.[140,212,1198,1220]" captionStartId-2="29.[140,212,1150,1172]" captionStartId-3="30.[140,212,934,956]" captionTargetBox-0="[140,1108,165,1142]" captionTargetBox-1="[140,1108,165,1179]" captionTargetBox-2="[140,1108,165,1130]" captionTargetBox-3="[195,1054,171,908]" captionTargetId-0="figure@27.[140,1108,165,1142]" captionTargetId-1="figure@28.[140,1108,165,1179]" captionTargetId-2="figure@29.[140,1108,165,1131]" captionTargetId-3="figure@30.[188,1059,165,915]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionTargetPageId-3="30" captionText-0="Figure 18. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, anterior part of body, lateral view; B, midbody segments, lateral view; C, posterior part of body, lateral view; D, anterior part of body, dorsal view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A-E, 0.5 mm &amp; F, 0.2 mm." captionText-1="Figure 19. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, anterior part of body, ventral view; B, segments 5-9, ventral view; C, posterior part of body, ventral view; D, cross-section of segment 4, caudal view; E, texture of prozonite, limbus and bisegmented seta; F, metatergal cerotegument with a bisegmented seta. − Scale bars: A, D, 0.5 mm, B, 1.0 mm, C, 0.02, E, 0.01 mm &amp; F, 0.005 mm." captionText-2="Figure 20. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, paratergum with ozopore, lateral view; B, leg; C, gonopods, ventral view; D, gonopod apex, ventral view. − Scale bars: A-C, 0.1 mm &amp; D, 0.05 mm." captionText-3="Figure 21. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, left gonopod, mesal view; B &amp; C, distal half of right gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3787354" figureDoi-1="http://doi.org/10.5281/zenodo.3787356" figureDoi-2="http://doi.org/10.5281/zenodo.3769088" figureDoi-3="http://doi.org/10.5281/zenodo.3787358" httpUri-0="https://zenodo.org/record/3787354/files/figure.png" httpUri-1="https://zenodo.org/record/3787356/files/figure.png" httpUri-2="https://zenodo.org/record/3769088/files/figure.png" httpUri-3="https://zenodo.org/record/3787358/files/figure.png" pageId="25" pageNumber="26">Figs 18-21</figureCitation>
.
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<emphasis id="A7EBEAFCA77FA67DFF62F9D5FEB2724F" bold="true" box="[140,316,1609,1636]" pageId="25" pageNumber="26">Type material.</emphasis>
<materialsCitation id="25F73CB3A77FA67EFEA9F9D5FB9E7022" ID-GBIF-Occurrence="2610433314" collectingDate="2007-05-18" collectionCode="IZAS, MNHN, ZMUM, ZMUC, SEM" collectorName="F. Brehier" country="China" lastPageId="26" lastPageNumber="27" location="Gui Dong 2 Cave" municipality="Mulun Nature Reserve" pageId="25" pageNumber="26" specimenCode="CHI-GX07-18/20, JC 313" specimenCount="11" specimenCount-female="3" specimenCount-juvenile="6" specimenCount-male="5" stateProvince="Guangxi Prov." typeStatus="holotype">
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<location id="90406035A77FA67DFC9BF9D5FBC17248" LSID="urn:lsid:plazi:treatment:1D3687F8A77FA679FF62FA23FBB07166:90406035A77FA67DFC9BF9D5FBC17248" box="[885,1103,1610,1636]" country="China" municipality="Mulun Nature Reserve" name="Gui Dong 2 Cave" pageId="25" pageNumber="26" stateProvince="Guangxi Prov.">Gui Dong 2 Cave</location>
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18.V. 
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, leg. 
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(
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(
<collectionCode id="F38EAE2BA77FA67DFCD4F9F2FCF372AB" box="[826,893,1645,1671]" collectionName="China, Beijing, Chinese Academy of Sciences, Institute of Zoology" pageId="25" pageNumber="26">IZAS</collectionCode>
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: 
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<specimenCount id="8399FD67A77CA67EFF62FC6BFF317022" box="[140,191,1012,1038]" pageId="26" pageNumber="27" type="female">2 ♀♀</specimenCount>
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<specimenCount id="8399FD67A77CA67EFF35FC6BFE947021" box="[219,282,1011,1038]" pageId="26" pageNumber="27" type="generic">2 juv.</specimenCount>
(
<collectionCode id="F38EAE2BA77CA67EFEC6FC6BFE077022" box="[296,393,1012,1038]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
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), 
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, 
<specimenCount id="8399FD67A77CA67EFDDCFC6BFDD97022" box="[562,599,1012,1038]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
(
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), 
<specimenCount id="8399FD67A77CA67EFD0DFC6BFC857023" box="[739,779,1012,1039]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
(
<collectionCode id="F38EAE2BA77CA67EFCF0FC6BFCF57022" box="[798,891,1012,1038]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="27">ZMUC</collectionCode>
), 
<specimenCount id="8399FD67A77CA67EFC60FC6BFC387023" box="[910,950,1012,1039]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
(
<collectionCode id="F38EAE2BA77CA67EFC27FC6BFB867022" box="[969,1032,1012,1038]" pageId="26" pageNumber="27">SEM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="25F73CB3A77CA67EFBF5FC6BFDE47078" ID-GBIF-Occurrence="2610433322" collectingDate="2007-05-22" collectionCode="SCAU" collectorName="L. Deharveng &amp; A. Bedos" country="China" location="Shen Long Dong Cave" municipality="Mulun Nature Reserve" pageId="26" pageNumber="27" specimenCode="CHI-GX07-22/01" specimenCount="8" specimenCount-female="3" specimenCount-juvenile="6" specimenCount-male="1" stateProvince="Dong Cave">
same locality, 
<location id="90406035A77CA67EFF00FB88FE71701D" LSID="urn:lsid:plazi:treatment:1D3687F8A77FA679FF62FA23FBB07166:90406035A77CA67EFF00FB88FE71701D" box="[238,511,1047,1073]" country="China" name="Shen Long Dong Cave" pageId="26" pageNumber="27" stateProvince="Dong Cave">Shen Long Dong Cave</location>
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<collectingDate id="F165E9C6A77CA67EFDE1FB88FD07701D" box="[527,649,1047,1073]" pageId="26" pageNumber="27" value="2007-05-22">
<date id="E121102EA77CA67EFDE1FB88FD07701D" box="[527,649,1047,1073]" pageId="26" pageNumber="27" value="2007-05-22">
22.V. 
<date id="E121102EA77CA67EFDA5FB88FD07701D" box="[587,649,1047,1073]" pageId="26" pageNumber="27" value="2007" year="2007">2007</date>
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, leg. 
<collectorName id="386A5338A77CA67EFD22FB88FCE5701D" box="[716,875,1047,1073]" pageId="26" pageNumber="27">L. Deharveng</collectorName>
&amp; 
<collectorName id="386A5338A77CA67EFC74FB88FB8D701D" box="[922,1027,1047,1073]" pageId="26" pageNumber="27">A. Bedos</collectorName>
(
<specimenCode id="C5399E95A77CA67EFBFDFB88FEAD7078" pageId="26" pageNumber="27">CHI-GX07-22/01</specimenCode>
); 
<specimenCount id="8399FD67A77CA67EFED6FBA5FEEE707A" box="[312,352,1082,1110]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
, 
<specimenCount id="8399FD67A77CA67EFE9CFBA5FE287078" box="[370,422,1082,1108]" pageId="26" pageNumber="27" type="female">3 ♀♀</specimenCount>
, 
<specimenCount id="8399FD67A77CA67EFE2DFBA5FD8C7078" box="[451,514,1082,1108]" pageId="26" pageNumber="27" type="generic">4 juv.</specimenCount>
(
<collectionCode id="F38EAE2BA77CA67EFDFFFBA5FDEC7078" box="[529,610,1082,1108]" collectionName="SCAU" pageId="26" pageNumber="27">SCAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="C1E06666A77CA67EFF62FCFDFF747794" ID-DOI="http://doi.org/10.5281/zenodo.3787352" ID-Zenodo-Dep="3787352" httpUri="https://zenodo.org/record/3787352/files/figure.png" pageId="26" pageNumber="27" startId="26.[140,212,866,888]" targetBox="[147,1102,185,842]" targetPageId="26">
<paragraph id="952036EEA77CA67EFF62FCFDFF747794" blockId="26.[140,1108,866,952]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF62FCFDFF707754" bold="true" box="[140,254,866,888]" pageId="26" pageNumber="27">
<figureCitation id="0DA42A6BA77CA67EFF62FCFDFF767754" box="[140,248,866,888]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStartId-0="8.[140,212,1558,1580]" captionStartId-1="9.[140,212,1590,1612]" captionStartId-2="10.[140,212,1590,1612]" captionStartId-3="11.[140,212,1622,1644]" captionStartId-4="13.[140,212,1558,1580]" captionStartId-5="14.[140,210,1558,1580]" captionStartId-6="15.[140,212,965,987]" captionTargetBox-0="[140,1108,506,1539]" captionTargetBox-1="[140,1108,407,1570]" captionTargetBox-2="[140,1108,826,1570]" captionTargetBox-3="[200,1045,807,1593]" captionTargetBox-4="[140,1108,385,1538]" captionTargetBox-5="[140,1108,384,1538]" captionTargetBox-6="[140,1108,165,945]" captionTargetId-0="figure@8.[140,1108,505,1539]" captionTargetId-1="figure@9.[140,1108,407,1570]" captionTargetId-2="figure@10.[140,1108,825,1570]" captionTargetId-3="figure@11.[194,1054,799,1602]" captionTargetId-4="figure@13.[140,1108,384,1538]" captionTargetId-5="figure@14.[140,1108,384,1538]" captionTargetId-6="figure@15.[140,1108,165,946]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionText-0="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." captionText-1="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." captionText-2="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." captionText-3="Figure 4. Eutrichodesmus basalis sp. n., ♂ paratype; A, leg 9, caudal view; B &amp; C, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm." captionText-4="Figure 5. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, habitus, lateral view; B, midbody segments, lateral view; C, posterior part of body, lateral view; D, midbody segments, ventral view; E, posterior part of body, ventral view; F, telson, ventral view. − Scale bars: A, 1.0 mm, B, C, E, 0.2 mm, D, 0.5 mm &amp; F, 0.1 mm." captionText-5="Figure 6. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, anterior part of body, frontoventral view; B, cross-section of a midbody segment, caudal view; C, enlarged paraterga with ozopores, lateral view; D, paratergum with porostele, caudal view; E, porostele; F, texture of prozonite and limbus; G, cerotegument with a bisegmented seta. − Scale bars: A, B, 0.2 mm, C, D, 0.1 mm, E, G, 0.01 mm &amp; F, 0.05 mm." captionText-6="Figure 7. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, body segments 6 and 7, ventral view; B, gonopods, ventral view; C, same, ventral view; D, gonopod apices. − Scale bars: A, 0.5 mm, B, 0.2 mm, C, 0.05 mm &amp; D, 0.02 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3787336" figureDoi-1="http://doi.org/10.5281/zenodo.3769071" figureDoi-2="http://doi.org/10.5281/zenodo.3769073" figureDoi-3="http://doi.org/10.5281/zenodo.3769075" figureDoi-4="http://doi.org/10.5281/zenodo.3769077" figureDoi-5="http://doi.org/10.5281/zenodo.3769079" figureDoi-6="http://doi.org/10.5281/zenodo.3769081" httpUri-0="https://zenodo.org/record/3787336/files/figure.png" httpUri-1="https://zenodo.org/record/3769071/files/figure.png" httpUri-2="https://zenodo.org/record/3769073/files/figure.png" httpUri-3="https://zenodo.org/record/3769075/files/figure.png" httpUri-4="https://zenodo.org/record/3769077/files/figure.png" httpUri-5="https://zenodo.org/record/3769079/files/figure.png" httpUri-6="https://zenodo.org/record/3769081/files/figure.png" pageId="26" pageNumber="27">Figure 17</figureCitation>
.
</emphasis>
<taxonomicName id="529F4D6DA77CA67EFEE9FCFDFE4A775B" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[263,452,866,887]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="latus" status="sp. n.">
<emphasis id="A7EBEAFCA77CA67EFEE9FCFDFE4A775B" box="[263,452,866,887]" italics="true" pageId="26" pageNumber="27">Eutrichodesmus latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77CA67EFE22FCFCFD8C7754" box="[460,514,867,888]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A77CA67EFDE1FCFCFD907755" box="[527,542,867,889]" pageId="26" pageNumber="27">♂</specimenCount>
<typeStatus id="4A24884CA77CA67EFDC4FCFCFDF77754" box="[554,633,867,888]" pageId="26" pageNumber="27">paratype</typeStatus>
from Yan Wu Dong Cave; 
<emphasis id="A7EBEAFCA77CA67EFC66FCFDFC19775B" bold="true" box="[904,919,866,887]" pageId="26" pageNumber="27">A</emphasis>
, leg 9; 
<emphasis id="A7EBEAFCA77CA67EFC0DFCFDFBA97754" bold="true" box="[995,1063,866,888]" pageId="26" pageNumber="27">B &amp; C</emphasis>
, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 
<quantity id="52679B0BA77CA67EFC3AFC1DFBAF77B4" box="[980,1057,898,920]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="mm" value="0.1">0.1 mm</quantity>
; B &amp; C, 
<quantity id="52679B0BA77CA67EFF47FC3DFF787794" box="[169,246,930,952]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
</caption>
<subSubSection id="DD856565A77CA67EFF57FBC2FC67705B" box="[185,1001,1117,1144]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="952036EEA77CA67EFF57FBC2FC67705B" blockId="26.[140,1108,1011,1672]" box="[185,1001,1117,1144]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FBC2FE89705B" bold="true" box="[185,263,1117,1143]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FBC2FE8C705B" box="[185,258,1117,1143]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Name</emphasis>
.
</emphasis>
To emphasize the obvious similarities with the preceding species.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A77CA67EFF57FB1FFD5070B6" box="[185,734,1152,1178]" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="952036EEA77CA67EFF57FB1FFD5070B6" blockId="26.[140,1108,1011,1672]" box="[185,734,1152,1178]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FB1FFEB670B6" bold="true" box="[185,312,1152,1178]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FB1FFEBD70B6" box="[185,307,1152,1178]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Diagnosis</emphasis>
.
</emphasis>
See diagnosis of the previous species.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A77CA67EFF57FB3CFC5A71BD" pageId="26" pageNumber="27" type="description">
<paragraph id="952036EEA77CA67EFF57FB3CFC797128" blockId="26.[140,1108,1011,1672]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FB3CFEC17091" bold="true" box="[185,335,1187,1213]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FB3CFEC47091" box="[185,330,1187,1213]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Description</emphasis>
.
</emphasis>
Length of adults of both sexes 
<emphasis id="A7EBEAFCA77CA67EFD49FB3BFD307091" box="[679,702,1188,1213]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
<quantity id="52679B0BA77CA67EFD2BFB3BFCCE7092" box="[709,832,1188,1214]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="26" pageNumber="27" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11-13 mm</quantity>
, width 3.0- 
<quantity id="52679B0BA77CA67EFC2FFB3BFB937092" box="[961,1053,1188,1214]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="26" pageNumber="27" unit="mm" value="3.3">3.3 mm</quantity>
; 
<specimenCount id="8399FD67A77CA67EFBC9FB3BFBC27093" box="[1063,1100,1188,1215]" pageId="26" pageNumber="27">♂♂</specimenCount>
usually a little smaller than 
<specimenCount id="8399FD67A77CA67EFE2AFB58FE6C70CD" box="[452,482,1223,1249]" pageId="26" pageNumber="27" type="female">♀♀</specimenCount>
. 
<typeStatus id="4A24884CA77CA67EFE11FB58FDE470CD" box="[511,618,1223,1249]" pageId="26" pageNumber="27">Holotype</typeStatus>
<emphasis id="A7EBEAFCA77CA67EFD9FFB58FD0670CC" box="[625,648,1223,1248]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
<quantity id="52679B0BA77CA67EFD7EFB58FD6D70CD" box="[656,739,1223,1249]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="mm" value="12.0">12 mm</quantity>
long and 3.0 mm wide. Coloration uniformly pallid or light yellowish, only seldom marbled light-brownish.
</paragraph>
<paragraph id="952036EEA77CA67EFF57FA92FC737166" blockId="26.[140,1108,1011,1672]" pageId="26" pageNumber="27">
All characters virtually as in 
<taxonomicName id="529F4D6DA77CA67EFE16FA91FDC4710B" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[504,586,1293,1319]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="latus" status="sp. n.">
<emphasis id="A7EBEAFCA77CA67EFE16FA91FDC4710B" box="[504,586,1293,1319]" italics="true" pageId="26" pageNumber="27">E. latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77CA67EFDBFFA91FD007104" box="[593,654,1294,1320]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="0DA42A6BA77CA67EFD73FA92FD61710B" box="[669,751,1293,1319]" captionStart="Figure 18" captionStartId="27.[140,212,1161,1183]" captionTargetBox="[140,1108,165,1142]" captionTargetId="figure@27.[140,1108,165,1142]" captionTargetPageId="27" captionText="Figure 18. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, anterior part of body, lateral view; B, midbody segments, lateral view; C, posterior part of body, lateral view; D, anterior part of body, dorsal view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A-E, 0.5 mm &amp; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787354" httpUri="https://zenodo.org/record/3787354/files/figure.png" pageId="26" pageNumber="27">Figs 18</figureCitation>
A-C, E, F; 19B-F; 20A, B), but collum with a row of very conspicuous teeth at front margin (
<figureCitation id="0DA42A6BA77CA67EFCAAFAAFFC3E7166" box="[836,944,1328,1354]" captionStart="Figure 18" captionStartId="27.[140,212,1161,1183]" captionTargetBox="[140,1108,165,1142]" captionTargetId="figure@27.[140,1108,165,1142]" captionTargetPageId="27" captionText="Figure 18. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, anterior part of body, lateral view; B, midbody segments, lateral view; C, posterior part of body, lateral view; D, anterior part of body, dorsal view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A-E, 0.5 mm &amp; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787354" httpUri="https://zenodo.org/record/3787354/files/figure.png" pageId="26" pageNumber="27">Figs 18D</figureCitation>
; 
<figureCitation id="0DA42A6BA77CA67EFC53FAAFFC7E7166" box="[957,1008,1328,1354]" captionStart="Figure 19" captionStartId="28.[140,212,1198,1220]" captionTargetBox="[140,1108,165,1179]" captionTargetId="figure@28.[140,1108,165,1179]" captionTargetPageId="28" captionText="Figure 19. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, anterior part of body, ventral view; B, segments 5-9, ventral view; C, posterior part of body, ventral view; D, cross-section of segment 4, caudal view; E, texture of prozonite, limbus and bisegmented seta; F, metatergal cerotegument with a bisegmented seta. − Scale bars: A, D, 0.5 mm, B, 1.0 mm, C, 0.02, E, 0.01 mm &amp; F, 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.3787356" httpUri="https://zenodo.org/record/3787356/files/figure.png" pageId="26" pageNumber="27">19A</figureCitation>
).
</paragraph>
<paragraph id="952036EEA77CA67EFF57FACCFC5A71BD" blockId="26.[140,1108,1011,1672]" pageId="26" pageNumber="27">
Gonopods (
<figureCitation id="0DA42A6BA77CA67EFEAFFACCFE5A7142" box="[321,468,1363,1390]" captionStart="Figure 20" captionStartId="29.[140,212,1150,1172]" captionTargetBox="[140,1108,165,1130]" captionTargetId="figure@29.[140,1108,165,1131]" captionTargetPageId="29" captionText="Figure 20. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, paratergum with ozopore, lateral view; B, leg; C, gonopods, ventral view; D, gonopod apex, ventral view. − Scale bars: A-C, 0.1 mm &amp; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769088" httpUri="https://zenodo.org/record/3769088/files/figure.png" pageId="26" pageNumber="27">Figs 20C, D</figureCitation>
; 
<figureCitation id="0DA42A6BA77CA67EFE0CFACBFD8E7142" box="[482,512,1364,1390]" captionStart="Figure 21" captionStartId="30.[140,212,934,956]" captionTargetBox="[195,1054,171,908]" captionTargetId="figure@30.[188,1059,165,915]" captionTargetPageId="30" captionText="Figure 21. Eutrichodesmus similis sp. n., ♂ paratype from Gui Dong 2 Cave; A, left gonopod, mesal view; B &amp; C, distal half of right gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787358" httpUri="https://zenodo.org/record/3787358/files/figure.png" pageId="26" pageNumber="27">21</figureCitation>
) slightly more complex than in 
<taxonomicName id="529F4D6DA77CA67EFC9BFACBFC477141" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[885,969,1363,1389]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="latus" status="sp. n.">
<emphasis id="A7EBEAFCA77CA67EFC9BFACBFC477141" box="[885,969,1363,1389]" italics="true" pageId="26" pageNumber="27">E. latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77CA67EFC3CFACBFB9F7142" box="[978,1041,1364,1390]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
Telopodite distally somewhat expanded and folded, with a hairpad subapically.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A77CA679FF57FA05FBB07166" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27" type="discussion">
<paragraph id="952036EEA77CA67EFF57FA05FBDD72AB" blockId="26.[140,1108,1011,1672]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FA05FEA47198" bold="true" box="[185,298,1434,1460]" pageId="26" pageNumber="27">
<emphasis id="A7EBEAFCA77CA67EFF57FA05FEAB7198" box="[185,293,1434,1460]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Remarks</emphasis>
.
</emphasis>
Together with 
<taxonomicName id="529F4D6DA77CA67EFE37FA04FDA27198" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[473,556,1434,1460]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="latus" status="sp. n.">
<emphasis id="A7EBEAFCA77CA67EFE37FA04FDA27198" box="[473,556,1434,1460]" italics="true" pageId="26" pageNumber="27">E. latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77CA67EFDDAFA05FDFA7198" box="[564,628,1434,1460]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
, this remarkable species partly bridges the gap between “doratodesmoid” 
<taxonomicName id="529F4D6DA77CA67EFE0BFA22FD1871FB" authorityName="Cook" authorityYear="1895" box="[485,662,1469,1495]" class="Diplopoda" family="Haplodesmidae" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">Haplodesmidae</taxonomicName>
and the large tropical family 
<taxonomicName id="529F4D6DA77CA67EFC30FA22FEF571D6" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" class="Diplopoda" family="Cryptodesmidae" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">
Cryptodesmidae Karsch, 
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<quantity id="52679B0BA77CA67EFEAEFA7FFEF571D6" box="[320,379,1504,1530]" metricMagnitude="1" metricUnit="m" metricValue="4.77266" pageId="26" pageNumber="27" unit="in" value="1879.0">
<date id="E121102EA77CA67EFEAEFA7FFEF571D6" box="[320,379,1504,1530]" pageId="26" pageNumber="27" value="1879">1879</date>
</quantity>
in
</quantity>
</taxonomicName>
showing the very broad and, at most, only slightly declivous paraterga so characteristic of cryptodesmids, including those occurring in East and Southeast Asia (
<bibRefCitation id="F10E4B1FA77CA67EFEAAF9B8FE7B726D" author="Hoffman" box="[324,501,1575,1601]" firstAuthor="Hoffman" pageId="26" pageNumber="27" refId="ref19420" refString="Hoffman RL (1980) Classification of the Diplopoda. Musee d'histoire naturelle: Geneve, 237 pp. (for 1979)." type="book" year="1980">
Hoffman 
<date id="E121102EA77CA67EFE56F9B8FE7B726D" box="[440,501,1575,1601]" pageId="26" pageNumber="27" value="1980">1980</date>
</bibRefCitation>
; 
<bibRefCitation id="F10E4B1FA77CA67EFDEAF9B8FD32726D" author="Simonsen" box="[516,700,1575,1601]" firstAuthor="Simonsen" pageId="26" pageNumber="27" refId="ref20144" refString="Simonsen A (1990) Phylogeny and biogeography of the millipede order Polydesmida, with special emphasis on the suborder Polydesmidea. Museum of Zoology, University of Bergen, 114 p." type="book" year="1990">
Simonsen 
<date id="E121102EA77CA67EFD93F9B8FD32726D" box="[637,700,1575,1601]" pageId="26" pageNumber="27" value="1990">1990</date>
</bibRefCitation>
). However, the metaterga in 
<taxonomicName id="529F4D6DA77CA67EFBFEF9B8FE827248" authorityName="Karsch" authorityYear="1879" class="Diplopoda" family="Cryptodesmidae" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">Cryptodesmidae</taxonomicName>
are often densely setose, the collum is always enlarged, flabellate and covering the head from above to an even greater extent than it does in 
<taxonomicName id="529F4D6DA77CA67EFC4FF9F2FBC172AB" authorityName="Silvestri" authorityYear="1896" box="[929,1103,1645,1671]" class="Diplopoda" family="Pyrgodesmidae" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">Pyrgodesmidae</taxonomicName>
,
</paragraph>
<caption id="C1E06666A77DA67FFF62FB16FE0270D3" ID-DOI="http://doi.org/10.5281/zenodo.3787354" ID-Zenodo-Dep="3787354" httpUri="https://zenodo.org/record/3787354/files/figure.png" pageId="27" pageNumber="28" startId="27.[140,212,1161,1183]" targetBox="[140,1108,165,1142]" targetPageId="27">
<paragraph id="952036EEA77DA67FFF62FB16FE0270D3" blockId="27.[140,1109,1161,1279]" pageId="27" pageNumber="28">
<emphasis id="A7EBEAFCA77DA67FFF62FB16FF7270B3" bold="true" box="[140,252,1161,1183]" pageId="27" pageNumber="28">
<figureCitation id="0DA42A6BA77DA67FFF62FB16FF7970B3" box="[140,247,1161,1183]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStartId-0="8.[140,212,1558,1580]" captionStartId-1="9.[140,212,1590,1612]" captionStartId-2="10.[140,212,1590,1612]" captionStartId-3="11.[140,212,1622,1644]" captionStartId-4="13.[140,212,1558,1580]" captionStartId-5="14.[140,210,1558,1580]" captionStartId-6="15.[140,212,965,987]" captionStartId-7="16.[140,211,964,986]" captionTargetBox-0="[140,1108,506,1539]" captionTargetBox-1="[140,1108,407,1570]" captionTargetBox-2="[140,1108,826,1570]" captionTargetBox-3="[200,1045,807,1593]" captionTargetBox-4="[140,1108,385,1538]" captionTargetBox-5="[140,1108,384,1538]" captionTargetBox-6="[140,1108,165,945]" captionTargetBox-7="[219,1033,171,941]" captionTargetId-0="figure@8.[140,1108,505,1539]" captionTargetId-1="figure@9.[140,1108,407,1570]" captionTargetId-2="figure@10.[140,1108,825,1570]" captionTargetId-3="figure@11.[194,1054,799,1602]" captionTargetId-4="figure@13.[140,1108,384,1538]" captionTargetId-5="figure@14.[140,1108,384,1538]" captionTargetId-6="figure@15.[140,1108,165,946]" captionTargetId-7="figure@16.[195,1052,165,944]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionTargetPageId-7="16" captionText-0="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." captionText-1="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." captionText-2="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." captionText-3="Figure 4. Eutrichodesmus basalis sp. n., ♂ paratype; A, leg 9, caudal view; B &amp; C, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm." captionText-4="Figure 5. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, habitus, lateral view; B, midbody segments, lateral view; C, posterior part of body, lateral view; D, midbody segments, ventral view; E, posterior part of body, ventral view; F, telson, ventral view. − Scale bars: A, 1.0 mm, B, C, E, 0.2 mm, D, 0.5 mm &amp; F, 0.1 mm." captionText-5="Figure 6. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, anterior part of body, frontoventral view; B, cross-section of a midbody segment, caudal view; C, enlarged paraterga with ozopores, lateral view; D, paratergum with porostele, caudal view; E, porostele; F, texture of prozonite and limbus; G, cerotegument with a bisegmented seta. − Scale bars: A, B, 0.2 mm, C, D, 0.1 mm, E, G, 0.01 mm &amp; F, 0.05 mm." captionText-6="Figure 7. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, body segments 6 and 7, ventral view; B, gonopods, ventral view; C, same, ventral view; D, gonopod apices. − Scale bars: A, 0.5 mm, B, 0.2 mm, C, 0.05 mm &amp; D, 0.02 mm." captionText-7="Figure 8. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A &amp; B, right gonopod, mesal and lateral views, respectively (micropilosity not shown; dp = distofemoral process). − Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3787336" figureDoi-1="http://doi.org/10.5281/zenodo.3769071" figureDoi-2="http://doi.org/10.5281/zenodo.3769073" figureDoi-3="http://doi.org/10.5281/zenodo.3769075" figureDoi-4="http://doi.org/10.5281/zenodo.3769077" figureDoi-5="http://doi.org/10.5281/zenodo.3769079" figureDoi-6="http://doi.org/10.5281/zenodo.3769081" figureDoi-7="http://doi.org/10.5281/zenodo.3769084" httpUri-0="https://zenodo.org/record/3787336/files/figure.png" httpUri-1="https://zenodo.org/record/3769071/files/figure.png" httpUri-2="https://zenodo.org/record/3769073/files/figure.png" httpUri-3="https://zenodo.org/record/3769075/files/figure.png" httpUri-4="https://zenodo.org/record/3769077/files/figure.png" httpUri-5="https://zenodo.org/record/3769079/files/figure.png" httpUri-6="https://zenodo.org/record/3769081/files/figure.png" httpUri-7="https://zenodo.org/record/3769084/files/figure.png" pageId="27" pageNumber="28">Figure 18</figureCitation>
.
</emphasis>
<taxonomicName id="529F4D6DA77DA67FFEEAFB15FE4270B3" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[260,460,1162,1183]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="similis" status="sp. n.">
<emphasis id="A7EBEAFCA77DA67FFEEAFB15FE4270B3" box="[260,460,1162,1183]" italics="true" pageId="27" pageNumber="28">Eutrichodesmus similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77DA67FFE3DFB15FD8670B3" box="[467,520,1162,1183]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A77DA67FFDFAFB15FDAD708C" box="[532,547,1162,1184]" pageId="27" pageNumber="28">♂</specimenCount>
<typeStatus id="4A24884CA77DA67FFDC3FB15FDF270B3" box="[557,636,1162,1183]" pageId="27" pageNumber="28">paratype</typeStatus>
from Gui Dong 2 Cave; 
<emphasis id="A7EBEAFCA77DA67FFC80FB16FCF370B2" bold="true" box="[878,893,1161,1182]" pageId="27" pageNumber="28">A</emphasis>
, anterior part of body, lateral view; 
<emphasis id="A7EBEAFCA77DA67FFF11FB36FE807092" bold="true" box="[255,270,1193,1214]" pageId="27" pageNumber="28">B</emphasis>
, midbody segments, lateral view; 
<emphasis id="A7EBEAFCA77DA67FFDA8FB36FDD97093" bold="true" box="[582,599,1193,1215]" pageId="27" pageNumber="28">C</emphasis>
, posterior part of body, lateral view; 
<emphasis id="A7EBEAFCA77DA67FFC47FB36FC327092" bold="true" box="[937,956,1193,1214]" pageId="27" pageNumber="28">D</emphasis>
, anterior part of body, dorsal view; 
<emphasis id="A7EBEAFCA77DA67FFED6FB56FEC870F2" bold="true" box="[312,326,1225,1246]" pageId="27" pageNumber="28">E</emphasis>
, midbody segments, dorsal view; 
<emphasis id="A7EBEAFCA77DA67FFD6EFB56FD0370F2" bold="true" box="[640,653,1225,1246]" pageId="27" pageNumber="28">F</emphasis>
, posterior part of body, dorsal view. − Scale bars: A-E, 
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&amp; F, 
<quantity id="52679B0BA77DA67FFED5FB75FE0670D3" box="[315,392,1258,1279]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="952036EEA77DA67FFF62FACCFE6171FB" blockId="27.[140,1108,1363,1672]" pageId="27" pageNumber="28">
whereas the paraterga are often deeply incised caudolaterally or have clear radii, again much as in some 
<taxonomicName id="529F4D6DA77DA67FFEB9FAE8FD8871BD" authorityName="Silvestri" authorityYear="1896" box="[343,518,1399,1425]" class="Diplopoda" family="Pyrgodesmidae" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Pyrgodesmidae</taxonomicName>
. Therefore, it seems safe for the present to consider the 
<taxonomicName id="529F4D6DA77DA67FFF56FA05FEFD7198" authorityName="Karsch" authorityYear="1879" box="[184,371,1434,1460]" class="Diplopoda" family="Cryptodesmidae" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Cryptodesmidae</taxonomicName>
as a distinct family in the superfamily 
<taxonomicName id="529F4D6DA77DA67FFCDCFA05FC547198" baseAuthorityName="Hoffman" baseAuthorityYear="1980" box="[818,986,1434,1460]" higherTaxonomySource="Manual Input" pageId="27" pageNumber="28" rank="superFamily" superFamily="Polydesmoidea">Polydesmoidea</taxonomicName>
(
<bibRefCitation id="F10E4B1FA77DA67FFC07FA05FF4771FB" author="Hoffman" firstAuthor="Hoffman" pageId="27" pageNumber="28" refId="ref19420" refString="Hoffman RL (1980) Classification of the Diplopoda. Musee d'histoire naturelle: Geneve, 237 pp. (for 1979)." type="book" year="1980">
Hoffman 
<date id="E121102EA77DA67FFF62FA22FF4771FB" box="[140,201,1469,1495]" pageId="27" pageNumber="28" value="1980">1980</date>
</bibRefCitation>
, 
<bibRefCitation id="F10E4B1FA77DA67FFF3BFA22FEAF71FB" author="Hoffman" box="[213,289,1469,1495]" firstAuthor="Hoffman" pageId="27" pageNumber="28" pagination="689 - 724" refId="ref19473" refString="Hoffman RL (1982 b) Diplopoda. In: Parker SP (ed.) Synopsis and Classification of Living Organisms. McGraw-Hill Book Company, New York &amp; St. Louis, 2: 689 - 724." type="journal article" year="1982" yearSuffix="b">
<date id="E121102EA77DA67FFF3BFA22FE9C71FB" box="[213,274,1469,1495]" pageId="27" pageNumber="28" value="1982">1982</date>
b
</bibRefCitation>
; 
<bibRefCitation id="F10E4B1FA77DA67FFEC3FA22FE6C71FB" author="Simonsen" box="[301,482,1469,1495]" firstAuthor="Simonsen" pageId="27" pageNumber="28" refId="ref20144" refString="Simonsen A (1990) Phylogeny and biogeography of the millipede order Polydesmida, with special emphasis on the suborder Polydesmidea. Museum of Zoology, University of Bergen, 114 p." type="book" year="1990">
Simonsen 
<date id="E121102EA77DA67FFE4DFA22FE6C71FB" box="[419,482,1469,1495]" pageId="27" pageNumber="28" value="1990">1990</date>
</bibRefCitation>
).
</paragraph>
<paragraph id="952036EEA77DA67FFF57FA7FFBDA72A4" blockId="27.[140,1108,1363,1672]" pageId="27" pageNumber="28">
What is more remarkable, however, is that the above two new Chinese species have the body so strongly flattened, and the paraterga apparently too broad to allow a tight volvation. This condition represents the opposite to that observed for 
<taxonomicName id="529F4D6DA77DA67FFC4FF9B8FB89726D" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[929,1031,1575,1601]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="basalis" status="sp. n.">
<emphasis id="A7EBEAFCA77DA67FFC4FF9B8FB89726D" box="[929,1031,1575,1601]" italics="true" pageId="27" pageNumber="28">E. basalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77DA67FFBE1F9B8FBC0726D" box="[1039,1102,1575,1601]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
, in which the paraterga are obviously too short to permit complete volvation. In other words, within 
<taxonomicName id="529F4D6DA77DA67FFEC3F9F2FE5B72AB" authorityName="Silvestri" authorityYear="1910" box="[301,469,1645,1671]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="A7EBEAFCA77DA67FFEC3F9F2FE5B72AB" box="[301,469,1645,1671]" italics="true" pageId="27" pageNumber="28">Eutrichodesmus</emphasis>
</taxonomicName>
we find species representing virtually the entire spectrum
</paragraph>
<caption id="C1E06666A77AA678FF62FB31FBC27108" ID-DOI="http://doi.org/10.5281/zenodo.3787356" ID-Zenodo-Dep="3787356" httpUri="https://zenodo.org/record/3787356/files/figure.png" pageId="28" pageNumber="29" startId="28.[140,212,1198,1220]" targetBox="[140,1108,165,1179]" targetPageId="28">
<paragraph id="952036EEA77AA678FF62FB31FBC27108" blockId="28.[140,1108,1198,1316]" pageId="28" pageNumber="29">
<emphasis id="A7EBEAFCA77AA678FF62FB31FF7270E8" bold="true" box="[140,252,1198,1220]" pageId="28" pageNumber="29">
<figureCitation id="0DA42A6BA77AA678FF62FB31FF7870E8" box="[140,246,1198,1220]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStartId-0="8.[140,212,1558,1580]" captionStartId-1="9.[140,212,1590,1612]" captionStartId-2="10.[140,212,1590,1612]" captionStartId-3="11.[140,212,1622,1644]" captionStartId-4="13.[140,212,1558,1580]" captionStartId-5="14.[140,210,1558,1580]" captionStartId-6="15.[140,212,965,987]" captionStartId-7="16.[140,211,964,986]" captionStartId-8="17.[140,212,1091,1113]" captionTargetBox-0="[140,1108,506,1539]" captionTargetBox-1="[140,1108,407,1570]" captionTargetBox-2="[140,1108,826,1570]" captionTargetBox-3="[200,1045,807,1593]" captionTargetBox-4="[140,1108,385,1538]" captionTargetBox-5="[140,1108,384,1538]" captionTargetBox-6="[140,1108,165,945]" captionTargetBox-7="[219,1033,171,941]" captionTargetBox-8="[140,1108,165,1071]" captionTargetId-0="figure@8.[140,1108,505,1539]" captionTargetId-1="figure@9.[140,1108,407,1570]" captionTargetId-2="figure@10.[140,1108,825,1570]" captionTargetId-3="figure@11.[194,1054,799,1602]" captionTargetId-4="figure@13.[140,1108,384,1538]" captionTargetId-5="figure@14.[140,1108,384,1538]" captionTargetId-6="figure@15.[140,1108,165,946]" captionTargetId-7="figure@16.[195,1052,165,944]" captionTargetId-8="figure@17.[140,1108,165,1071]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionTargetPageId-7="16" captionTargetPageId-8="17" captionText-0="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." captionText-1="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." captionText-2="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." captionText-3="Figure 4. Eutrichodesmus basalis sp. n., ♂ paratype; A, leg 9, caudal view; B &amp; C, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm." captionText-4="Figure 5. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, habitus, lateral view; B, midbody segments, lateral view; C, posterior part of body, lateral view; D, midbody segments, ventral view; E, posterior part of body, ventral view; F, telson, ventral view. − Scale bars: A, 1.0 mm, B, C, E, 0.2 mm, D, 0.5 mm &amp; F, 0.1 mm." captionText-5="Figure 6. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, anterior part of body, frontoventral view; B, cross-section of a midbody segment, caudal view; C, enlarged paraterga with ozopores, lateral view; D, paratergum with porostele, caudal view; E, porostele; F, texture of prozonite and limbus; G, cerotegument with a bisegmented seta. − Scale bars: A, B, 0.2 mm, C, D, 0.1 mm, E, G, 0.01 mm &amp; F, 0.05 mm." captionText-6="Figure 7. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A, body segments 6 and 7, ventral view; B, gonopods, ventral view; C, same, ventral view; D, gonopod apices. − Scale bars: A, 0.5 mm, B, 0.2 mm, C, 0.05 mm &amp; D, 0.02 mm." captionText-7="Figure 8. Eutrichodesmus armatocaudatus sp. n., ♂ paratype from cave 4; A &amp; B, right gonopod, mesal and lateral views, respectively (micropilosity not shown; dp = distofemoral process). − Scale bar: 0.2 mm." captionText-8="Figure 9. Eutrichodesmus communicans sp. n., ♂ paratype from Malo Island, habitus." figureDoi-0="http://doi.org/10.5281/zenodo.3787336" figureDoi-1="http://doi.org/10.5281/zenodo.3769071" figureDoi-2="http://doi.org/10.5281/zenodo.3769073" figureDoi-3="http://doi.org/10.5281/zenodo.3769075" figureDoi-4="http://doi.org/10.5281/zenodo.3769077" figureDoi-5="http://doi.org/10.5281/zenodo.3769079" figureDoi-6="http://doi.org/10.5281/zenodo.3769081" figureDoi-7="http://doi.org/10.5281/zenodo.3769084" figureDoi-8="http://doi.org/10.5281/zenodo.3769086" httpUri-0="https://zenodo.org/record/3787336/files/figure.png" httpUri-1="https://zenodo.org/record/3769071/files/figure.png" httpUri-2="https://zenodo.org/record/3769073/files/figure.png" httpUri-3="https://zenodo.org/record/3769075/files/figure.png" httpUri-4="https://zenodo.org/record/3769077/files/figure.png" httpUri-5="https://zenodo.org/record/3769079/files/figure.png" httpUri-6="https://zenodo.org/record/3769081/files/figure.png" httpUri-7="https://zenodo.org/record/3769084/files/figure.png" httpUri-8="https://zenodo.org/record/3769086/files/figure.png" pageId="28" pageNumber="29">Figure 19</figureCitation>
.
</emphasis>
<taxonomicName id="529F4D6DA77AA678FEEAFB31FE4270EF" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[260,460,1198,1219]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="similis" status="sp. n.">
<emphasis id="A7EBEAFCA77AA678FEEAFB31FE4270EF" box="[260,460,1198,1219]" italics="true" pageId="28" pageNumber="29">Eutrichodesmus similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A77AA678FE3DFB30FD8670E8" box="[467,520,1199,1220]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A77AA678FDFAFB30FDAD70E9" box="[532,547,1199,1221]" pageId="28" pageNumber="29">♂</specimenCount>
<typeStatus id="4A24884CA77AA678FDC3FB30FDF270E8" box="[557,636,1199,1220]" pageId="28" pageNumber="29">paratype</typeStatus>
from Gui Dong 2 Cave; 
<emphasis id="A7EBEAFCA77AA678FC80FB31FCF370EF" bold="true" box="[878,893,1198,1219]" pageId="28" pageNumber="29">A</emphasis>
, anterior part of body, ventral view; 
<emphasis id="A7EBEAFCA77AA678FEE4FB51FE9770CF" bold="true" box="[266,281,1230,1251]" pageId="28" pageNumber="29">B</emphasis>
, segments 5-9, ventral view; 
<emphasis id="A7EBEAFCA77AA678FDC3FB51FDB070C8" bold="true" box="[557,574,1230,1252]" pageId="28" pageNumber="29">C</emphasis>
, posterior part of body, ventral view; 
<emphasis id="A7EBEAFCA77AA678FC4AFB51FC3970CF" bold="true" box="[932,951,1230,1251]" pageId="28" pageNumber="29">D</emphasis>
, cross-section of segment 4, caudal view; 
<emphasis id="A7EBEAFCA77AA678FE84FB71FEF6712F" bold="true" box="[362,376,1262,1283]" pageId="28" pageNumber="29">E</emphasis>
, texture of prozonite, limbus and bisegmented seta; 
<emphasis id="A7EBEAFCA77AA678FCB8FB71FCEC712F" bold="true" box="[854,866,1262,1283]" pageId="28" pageNumber="29">F</emphasis>
, metatergal cerotegument with a bisegmented seta. − Scale bars: A, D, 
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, B, 1.0 mm, C, 0.02, E, 
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&amp; F, 
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.
</paragraph>
</caption>
<paragraph id="952036EEA77AA678FF62FAE8FEDF7198" blockId="28.[140,1108,1399,1672]" pageId="28" pageNumber="29">of “doratodesmid” evolution of conglobation, including a few exceptional cases of imperfect volvation.</paragraph>
<paragraph id="952036EEA77AA679FF57FA22FBB07166" blockId="28.[140,1108,1399,1672]" lastBlockId="29.[140,1108,1293,1354]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Concerning the two new Chinese species that are not capable of complete conglobation, the situation somewhat parallels that observed in 
<taxonomicName id="529F4D6DA77AA678FCAFFA7FFEB17232" authority="Hoffman, 1990" authorityName="Hoffman" authorityYear="1990" class="Diplopoda" family="Sphaeriodesmidae" genus="Proeilodesmus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mecistonyx">
<emphasis id="A7EBEAFCA77AA678FCAFFA7FFBDD71D6" box="[833,1107,1504,1530]" italics="true" pageId="28" pageNumber="29">Proeilodesmus mecistonyx</emphasis>
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Hoffman, 
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</taxonomicName>
(
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Sphaeriodesmidae Humbert &amp; DeSaussure, 
<date id="E121102EA77AA678FCA2F99CFC077231" box="[844,905,1539,1565]" pageId="28" pageNumber="29" value="1869">1869</date>
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, Sphaeriodesmoidea), a cavernicole from 
<collectingCountry id="ED88767EA77AA678FE45F9B8FE71726D" box="[427,511,1575,1601]" name="Mexico" pageId="28" pageNumber="29">Mexico</collectingCountry>
that is apparently incapable of volvation (
<bibRefCitation id="F10E4B1FA77AA678FC07F9B8FF457248" author="Hoffman" firstAuthor="Hoffman" pageId="28" pageNumber="29" pagination="669 - 679" refId="ref19513" refString="Hoffman RL (1990) A phylogenetically interesting sphaeriodesmid millipede from Oaxaca, Mexico (Polydesmida: Sphaeriodesmidae). Revue suisse de Zoologie 97 (3): 669 - 679." type="journal article" year="1990">
Hoffman 
<date id="E121102EA77AA678FF62F9D5FF457248" box="[140,203,1610,1636]" pageId="28" pageNumber="29" value="1990">1990</date>
</bibRefCitation>
). In contrast to “doratodesmids”, however, most of the characters of this millipede seem plesiomorphic and suggest a link between a presumed flat-bodied chelodesmidean (= leptodesmidean) ancestor and the remaining, convex, truly volvatory members of this rather large, Neotropical family (
<bibRefCitation id="F10E4B1FA77BA679FD55FAAFFCE77166" author="Hoffman" box="[699,873,1328,1354]" firstAuthor="Hoffman" pageId="29" pageNumber="30" pagination="669 - 679" refId="ref19513" refString="Hoffman RL (1990) A phylogenetically interesting sphaeriodesmid millipede from Oaxaca, Mexico (Polydesmida: Sphaeriodesmidae). Revue suisse de Zoologie 97 (3): 669 - 679." type="journal article" year="1990">
Hoffman 
<date id="E121102EA77BA679FCC2FAAFFCE77166" box="[812,873,1328,1354]" pageId="29" pageNumber="30" value="1990">1990</date>
</bibRefCitation>
; 
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Golovatch 
<date id="E121102EA77BA679FC1CFAAFFBBF7166" box="[1010,1073,1328,1354]" pageId="29" pageNumber="30" value="2003">2003</date>
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>