321 lines
34 KiB
XML
321 lines
34 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.67.51332" ID-GBIF-Dataset="3af178c2-c3e5-49a3-8987-f37bcc82cdaf" ID-GBIF-Taxon="182223455" ID-Pensoft-Pub="1860-1324-1-69" ID-Pensoft-UUID="48A2D68FF09F5A41B01347C0DBEF72C8" ID-ZooBank="2B601B5DE1BD44B7BA89554E3AB5EAE1" LSID="urn:lsid:zoobank.org:act:7C19F1EE-EC88-4090-A25B-0089401B750A" ModsDocID="1860-1324-67-1-69" checkinTime="1621372352603" checkinUser="pensoft" docAuthor="Shimizu, So" docDate="2020" docId="1CE53E820CE655BABAECAA774EFB0B62" docLanguage="en" docName="DeutEntomolZeit 67(1): 69-126" docOrigin="Deutsche Entomologische Zeitschrift 67 (1)" docPubDate="2020-05-11" docSource="http://dx.doi.org/10.3897/dez.67.51332" docTitle="Enicospilus nepalensis Shimizu 2020, sp. nov." docType="treatment" docUuid="7C19F1EE-EC88-4090-A25B-0089401B750A" docUuidSource="ZooBank" docVersion="4" id="48A2D68FF09F5A41B01347C0DBEF72C8" lastPageNumber="69" masterDocId="48A2D68FF09F5A41B01347C0DBEF72C8" masterDocTitle="The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species" masterLastPageNumber="126" masterPageNumber="69" pageId="0" pageNumber="69" updateTime="1632029310057" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Shimizu, So</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5202-4552</mods:nameIdentifier>
|
||
<mods:affiliation>Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan & Research Fellow (DC 1 and Overseas Challenge Program for Young Researchers), Japan Society for the Promotion of Science, Tokyo, Japan & Department of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
|
||
<mods:nameIdentifier type="email">parasitoidwasp.sou@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2020-05-11</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>67</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>69</mods:start>
|
||
<mods:end>126</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/dez.67.51332</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.67.51332</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1860-1324-1-69</mods:identifier>
|
||
<mods:identifier type="ZooBank">2B601B5DE1BD44B7BA89554E3AB5EAE1</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">48A2D68FF09F5A41B01347C0DBEF72C8</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="182223455" LSID="urn:lsid:zoobank.org:act:7C19F1EE-EC88-4090-A25B-0089401B750A" httpUri="http://treatment.plazi.org/id/1CE53E820CE655BABAECAA774EFB0B62" lastPageNumber="69" pageId="0" pageNumber="69">
|
||
<subSubSection pageId="0" pageNumber="69" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<taxonomicName LSID="http://zoobank.org7C19F1EE-EC88-4090-A25B-0089401B750A" authority="Shimizu" authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus nepalensis" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">Enicospilus nepalensis Shimizu</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="69">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters for some species of Nepalese Enicospilus. A-D. Outer surface of mandible: A. E. nepalensis Shimizu sp. nov., B. E. tangi Shimizu sp. nov., C. E. tripartitus, D. E. yonezawanus; E-H. Scutellum, in dorsal view: E. E. formosensis, F. E. kakanicus Shimizu sp. nov., G. E. phulchokiensis Shimizu sp. nov., H. E. tangi Shimizu sp. nov.; I, J. Hind tarsal claw: I. E. biharensis, J. E. nikami Shimizu sp. nov." figureDoi="10.3897/dez.67.51332.figure2" httpUri="https://binary.pensoft.net/fig/408834" pageId="0" pageNumber="69">Figs 2A</figureCitation>
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">, 18</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="etymology">
|
||
<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">The specific name is derived from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="69">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<materialsCitation collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" country="Nepal" location="Nepal" specimenCount="2" specimenCount-female="2">
|
||
<specimenCount type="female">2♀♀</specimenCount>
|
||
:
|
||
<collectingCountry name="Nepal">Nepal</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Type series:
|
||
<typeStatus>holotype</typeStatus>
|
||
<specimenCount>♀</specimenCount>
|
||
,
|
||
<materialsCitation collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectorName="M. G. Allen" country="Nepal" elevation="950" location="Nepal" specimenCount="1">
|
||
Pokhara (
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</elevation>
|
||
</quantity>
|
||
),
|
||
<collectingCountry name="Nepal">Nepal</collectingCountry>
|
||
, VII-VIII.1983,
|
||
<collectorName>M.G. Allen</collectorName>
|
||
leg. (LT) (NHMUK) (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters for some species of Nepalese Enicospilus. A-D. Outer surface of mandible: A. E. nepalensis Shimizu sp. nov., B. E. tangi Shimizu sp. nov., C. E. tripartitus, D. E. yonezawanus; E-H. Scutellum, in dorsal view: E. E. formosensis, F. E. kakanicus Shimizu sp. nov., G. E. phulchokiensis Shimizu sp. nov., H. E. tangi Shimizu sp. nov.; I, J. Hind tarsal claw: I. E. biharensis, J. E. nikami Shimizu sp. nov." figureDoi="10.3897/dez.67.51332.figure2" httpUri="https://binary.pensoft.net/fig/408834" pageId="0" pageNumber="69">2A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<typeStatus>paratype</typeStatus>
|
||
<specimenCount>♀</specimenCount>
|
||
, same label and repository as
|
||
<typeStatus>holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="distribution">
|
||
<paragraph pageId="0" pageNumber="69">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">Nepal.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="description">
|
||
<paragraph pageId="0" pageNumber="69">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Female</emphasis>
|
||
(Holotype) (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18</figureCitation>
|
||
). Body length ca 16.5 mm.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Head</emphasis>
|
||
with GOI = 2.5 (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18C</figureCitation>
|
||
). Lower face 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as high, finely punctate with setae, strongly shiny (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B</figureCitation>
|
||
). Clypeus 1.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as high, finely punctate with setae, moderately convex in profile, its lower margin impressed (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B, C</figureCitation>
|
||
). Malar space 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basal mandibular width (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B, C</figureCitation>
|
||
). Mandible weakly twisted by ca 15°, moderately long, its proximal half evenly narrowed and distal half subparallel sided, its outer surface entirely almost flat with long and rather stout setae (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters for some species of Nepalese Enicospilus. A-D. Outer surface of mandible: A. E. nepalensis Shimizu sp. nov., B. E. tangi Shimizu sp. nov., C. E. tripartitus, D. E. yonezawanus; E-H. Scutellum, in dorsal view: E. E. formosensis, F. E. kakanicus Shimizu sp. nov., G. E. phulchokiensis Shimizu sp. nov., H. E. tangi Shimizu sp. nov.; I, J. Hind tarsal claw: I. E. biharensis, J. E. nikami Shimizu sp. nov." figureDoi="10.3897/dez.67.51332.figure2" httpUri="https://binary.pensoft.net/fig/408834" pageId="0" pageNumber="69">2A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B, C</figureCitation>
|
||
). Upper mandibular tooth 1.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as lower one, very slender and cylindrical (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters for some species of Nepalese Enicospilus. A-D. Outer surface of mandible: A. E. nepalensis Shimizu sp. nov., B. E. tangi Shimizu sp. nov., C. E. tripartitus, D. E. yonezawanus; E-H. Scutellum, in dorsal view: E. E. formosensis, F. E. kakanicus Shimizu sp. nov., G. E. phulchokiensis Shimizu sp. nov., H. E. tangi Shimizu sp. nov.; I, J. Hind tarsal claw: I. E. biharensis, J. E. nikami Shimizu sp. nov." figureDoi="10.3897/dez.67.51332.figure2" httpUri="https://binary.pensoft.net/fig/408834" pageId="0" pageNumber="69">2A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B</figureCitation>
|
||
). Frons, vertex and gena strongly shiny with fine setae (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B-D</figureCitation>
|
||
). Posterior ocellus rather small and separated from eye by 0.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
its own maximum diameter (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B-D</figureCitation>
|
||
). Ventral end of occipital carina joining oral carina. Antenna with 49 flagellomeres; first flagellomere 1.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as second; 20th flagellomere 2.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Mesosoma</emphasis>
|
||
entirely strongly shiny with setae (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Pronotum punctostriate dorsally and finely coriaceous ventrally (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Mesoscutum 1.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as its maximum width, almost smooth with very fine punctures with setae, and evenly rounded in profile (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Notauli absent (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Scutellum moderately convex, almost smooth with very fine and sparse punctures with setae, with lateral longitudinal carinae reaching posterior end (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Epicnemium from densely strigose dorsally to densely punctate ventrally with setae. Epicnemial carina present, evenly curved to anterior, its dorsal end not reaching anterior margin of mesopleuron (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Mesopleuron finely punctate dorsally and longitudinally punctostriate to strigose ventrally (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Submetapleural carina almost parallel sided centrally and weakly broadened anteriorly (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Metapleuron moderately punctate with setae (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
). Propodeum evenly rounded in profile; anterior transverse carina complete centrally, its lateral end almost joining pleural carina; anterior area longitudinally striate; spiracular area almost smooth with very fine and sparse punctures and setae; posterior area rather finely subconcentrically striate; propodeal spiracle elliptical, its outer margin not joining pleural carina by a ridge (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Wings</emphasis>
|
||
. Fore wing length ca 11.0 mm with AI = 0.4, CI = 0.3, DI = 0.4, ICI = 0.4, SDI = 1.2, SI = 0.2, SRI = 0.3; vein 1m-cu&M almost evenly curved; vein 2r&RS slightly sinuous and RS evenly curved; fenestra and sclerites of discosubmarginal cell as in Figure
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18F</figureCitation>
|
||
; proximal sclerite triangular, not confluent with distal sclerite, very strongly pigmented; central sclerite small and its major diameter subequal to thickness of vein 2r&RS, suboval, weakly sclerotised and pigmented, positioned in posterodistal part of fenestra; distal sclerite moderately pigmented; proximal corner of marginal cell evenly setose; posterodistal corner of second discal cell ca 95°; posterodistal corner of subbasal cell ca 95°; vein 1cu-a slightly antefurcal to M&RS by 0.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
1cu-a length (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18F</figureCitation>
|
||
). Hind wing with NI = 1.2, RI = 1.7; vein RS straight; vein RA with 6 uniform hamuli.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Legs</emphasis>
|
||
. Outer surface of fore tibia without dense and long spines. Hind leg with coxa in profile 1.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as deep; basitarsus 2.0
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as second tarsomere; fourth tarsomere 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as third tarsomere and 3.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide; tarsal claw simply pectinate.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Metasoma</emphasis>
|
||
with PI = 2.8, DMI = 1.3, THI = 2.5; dorsal margin of tergite 1 more or less sinuous; thyridium elongate (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Colour</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18</figureCitation>
|
||
). Entirely testaceous except for apex of mandible, posterior part of T5, and T6-8 black. Wings hyaline; proximal sclerite brown; central and distal sclerites amber; veins brown.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Variations</emphasis>
|
||
(n = 2): body length 15.5-16.5 mm; head with GOI = 2.4-2.5; clypeus 1.6-1.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as high; malar space 0.3-0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basal mandibular width; mandible twisted by 15-25°; upper mandibular tooth 1.6-2.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as lower one; antenna with first flagellomere 1.6-1.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as second; pronotum punctostriate dorsally and finely coriaceous ventrally or entirely almost smooth to weakly coriaceous with very sparse and fine punctures; metapleuron sparsely to moderately punctate; fore wing length 10.0-11.0 mm; hind coxa in profile 1.6-1.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as deep; fourth tarsomere 3.5-3.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide; metasoma with PI = 2.7-2.8; THI = 2.5-2.8; mandible proximally testaceous and apically black or entirely dark brown to black.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="69">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<taxonomicName authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus nepalensis" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="nepalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Enicospilus nepalensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is probably closely related to or belongs to the
|
||
<taxonomicName genus="E." lsidName="E. ramidulus" pageId="0" pageNumber="69" rank="species" species="ramidulus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. ramidulus</emphasis>
|
||
</taxonomicName>
|
||
complex. Among the complex,
|
||
<taxonomicName genus="E." lsidName="E. nepalensis" pageId="0" pageNumber="69" rank="species" species="nepalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. nepalensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most closely related to
|
||
<taxonomicName genus="E." lsidName="E. tricorniatus" pageId="0" pageNumber="69" rank="species" species="tricorniatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. tricorniatus</emphasis>
|
||
</taxonomicName>
|
||
Rao & Nikam, 1970 based on the rather small ocelli relative to other
|
||
<taxonomicName authorityName="Stephens" authorityYear="1835" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Enicospilus</emphasis>
|
||
</taxonomicName>
|
||
(posterior ocellus separated from eye by more than 0.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
its own maximum diameter) (e.g. Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B-D</figureCitation>
|
||
), highly shiny body (e.g. Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18A-E</figureCitation>
|
||
), shape of body (e.g. Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18A</figureCitation>
|
||
), shape and position of the fore wing veins and sclerites (e.g. Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18F</figureCitation>
|
||
), distribution, etc. However,
|
||
<taxonomicName genus="E." lsidName="E. nepalensis" pageId="0" pageNumber="69" rank="species" species="nepalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. nepalensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is readily distinguishable from
|
||
<taxonomicName genus="E." lsidName="E. tricorniatus" pageId="0" pageNumber="69" rank="species" species="tricorniatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. tricorniatus</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: lower face more or less elongate and 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as high (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B</figureCitation>
|
||
) (lower face subquadrate to transverse and 1.0-1.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as high in
|
||
<taxonomicName genus="E." lsidName="E. tricorniatus" pageId="0" pageNumber="69" rank="species" species="tricorniatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. tricorniatus</emphasis>
|
||
</taxonomicName>
|
||
), the central sclerite weakly sclerotised and pigmented (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18F</figureCitation>
|
||
) (moderately to strongly sclerotised and pigmented in
|
||
<taxonomicName genus="E." lsidName="E. tricorniatus" pageId="0" pageNumber="69" rank="species" species="tricorniatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. tricorniatus</emphasis>
|
||
</taxonomicName>
|
||
), moderate-sized, fore wing length 10.0-11.0 mm (small, fore wing length less than 8.5 mm in
|
||
<taxonomicName genus="E." lsidName="E. tricorniatus" pageId="0" pageNumber="69" rank="species" species="tricorniatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. tricorniatus</emphasis>
|
||
</taxonomicName>
|
||
), posterior ocellus separated from eye by 0.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
its own maximum diameter (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Enicospilus nepalensis Shimizu sp. nov., ♀, holotype. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing." figureDoi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69">18B-D</figureCitation>
|
||
) (posterior ocellus separated from eye by almost its own maximum diameter in
|
||
<taxonomicName genus="E." lsidName="E. tricorniatus" pageId="0" pageNumber="69" rank="species" species="tricorniatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">E. tricorniatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/dez.67.51332.figure18" httpUri="https://binary.pensoft.net/fig/408850" pageId="0" pageNumber="69" start="Figure 18" startId="F18">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 18.</emphasis>
|
||
<taxonomicName authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus nepalensis" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="nepalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Enicospilus nepalensis</emphasis>
|
||
</taxonomicName>
|
||
Shimizu sp. nov., ♀, holotype.
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A.</emphasis>
|
||
Habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B.</emphasis>
|
||
Head, frontal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C.</emphasis>
|
||
Head, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">D.</emphasis>
|
||
Head, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E.</emphasis>
|
||
Mesosoma, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="69">F.</emphasis>
|
||
Central part of fore wing.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |