242 lines
24 KiB
XML
242 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.339.4903" ID-GBIF-Dataset="f5e2c4d6-742c-4235-bf0e-b97e1ec678d2" ID-PMC="PMC3800827" ID-Pensoft-Pub="1313-2970-339-1" ID-PubMed="24146585" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-339-1" ModsDocOrigin="ZooKeys 339" ModsDocTitle="Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)" checkinTime="1451246844427" checkinUser="pensoft" docAuthor="Borkenhagen, Kai & Krupp, Friedhelm" docDate="2013" docId="1CDC01322A10AC251F77F62CE279BA62" docLanguage="en" docName="ZooKeys 339: 1-53" docOrigin="ZooKeys 339" docSource="http://dx.doi.org/10.3897/zookeys.339.4903" docTitle="Carasobarbus harterti Guenther 1901, comb. n." docType="treatment" docVersion="3" lastPageNumber="27" masterDocId="FF8B0A687873FFFAFFC0C22FFFF3FFD2" masterDocTitle="Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="25" updateTime="1668156576871" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Borkenhagen, Kai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Krupp, Friedhelm</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>339</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>53</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.339.4903</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.339.4903</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-339-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152048582" LSID="urn:lsid:plazi:treatment:1CDC01322A10AC251F77F62CE279BA62" httpUri="http://treatment.plazi.org/id/1CDC01322A10AC251F77F62CE279BA62" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
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<subSubSection pageId="24" pageNumber="25" type="nomenclature">
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<paragraph pageId="24" pageNumber="25">
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<taxonomicName LSID="http://species-id.net/wiki/Carasobarbus_harterti" authority="Guenther, 1901" authorityName="Guenther" authorityYear="1901" class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="harterti">
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<pageBreakToken pageId="24" pageNumber="25" start="start">Carasobarbus</pageBreakToken>
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harterti (
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<normalizedToken originalValue="Günther">Guenther</normalizedToken>
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, 1901)
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</taxonomicName>
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<taxonomicNameLabel pageId="24" pageNumber="25">comb. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="reference_group">
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<paragraph pageId="24" pageNumber="25">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus harterti" order="Cypriniformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="harterti">Barbus harterti</taxonomicName>
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<bibRefCitation author="Guenther, A" journalOrPublisher="A Journal of Zoology in Connection with the Tring Museum" pageId="44" pageNumber="45" pagination="367 - 368" title="Second notice of new species of fishes from Morocco. Novitates Zoologicae." volume="8" year="1901">
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<normalizedToken originalValue="Günther">Guenther</normalizedToken>
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1901
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</bibRefCitation>
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: 367.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="material">
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<paragraph pageId="24" pageNumber="25">Material.</paragraph>
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<paragraph pageId="24" pageNumber="25">
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Type material. Syntypes: BMNH 1901.7.26:4-5, 2, Morocco, Oued Oum er Rbia [
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.327778">33°19'40"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-8.333889">8°20'2"W</geoCoordinate>
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], E. Hartert.
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</paragraph>
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<paragraph pageId="24" pageNumber="25">
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Non-type material. Oued Oum er Rbia drainage. BMNH 1902.7.28:27-33, 7; BMNH 1903.10.29:11-15, 8, Morocco, Oued Oum er Rbia [
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.327778">33°19'40"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-8.333889">8°20'2"W</geoCoordinate>
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], F. Riggenbach. - BMNH 1903.7.1:5-7, 3, Morocco, Oued Oum er Rbia near El Jadida [
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.255">33°15'18"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-8.506111">8°30'22"W</geoCoordinate>
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], F. Riggenbach. - MNHN 1912-0089, 1; MNHN 1912-0090, 1; MNHN 1912-0091, 1; MNHN 1912-0092, 1; MNHN 1912-0093, 1, Morocco, Oued Oum er Rbia near Azemmour [
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.289444">33°17'22"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-8.3425">8°20'33"W</geoCoordinate>
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], C. du Gast, 1912. - SMF 33366, 1; SMF 33368, 1; SMF 33370, 1, Morocco, Oued Oum er Rbia (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="32.31472">32°18'53"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-6.909167">6°54'33"W</geoCoordinate>
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), A. Azeroual et al., 9 Apr 2011.
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</paragraph>
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<paragraph pageId="24" pageNumber="25">
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Oued Tennsift drainage. BMNH 1902.7.28:34, 1, Morocco, Oued Talmest [
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="31.870834">31°52'15"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-9.308611">9°18'31"W</geoCoordinate>
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], F. Riggenbach.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="diagnosis">
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<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
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<paragraph pageId="24" pageNumber="25">Two pairs of long barbels; 31 to 38 scales in the lateral line and 13 to 17 scales around the least circumference of the caudal peduncle; dorsal fin longer than anal fin and less than 15 % of the length of its last unbranched ray is flexible, dorsal profile of the head straight.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
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<paragraph pageId="24" pageNumber="25">Description.</paragraph>
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<paragraph pageId="24" pageNumber="25">The body is of moderate height and without a nuchal hump. The head is triangular with almost straight dorsal and ventral profile (Figs 19, 20). The head length is shorter than the body depth (Fig. 12). The mouth is subterminal with two pairs of long barbels (Table 2). The eyes are in the anterior half of the head and relatively big. The morphometric characters are summarised in Table 1.</paragraph>
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<caption pageId="24" pageNumber="25">
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<paragraph pageId="24" pageNumber="25">
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Figure 19.
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="harterti">Carasobarbus harterti</taxonomicName>
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, syntype (BMNH 1901.7.26:4-5) from Oued Oum er Rbia, © The Natural History Museum, London.
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</paragraph>
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</caption>
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<caption pageId="24" pageNumber="25">
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<paragraph pageId="24" pageNumber="25">
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Figure 20.
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="harterti">Carasobarbus harterti</taxonomicName>
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, live specimen from Oued Oum er Rbia.
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</paragraph>
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</caption>
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<paragraph pageId="24" pageNumber="25">The dorsal fin is long and strongly ossified and less than 15 % of the length of its last unbranched ray is flexible. Its last unbranched ray is as long as or longer than the head (Fig. 4). It usually has four unbranched and nine branched rays (Table 3). The anal fin usually has three unbranched and six or seven branched rays (Table 4). It does not reach the caudal fin origin.</paragraph>
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<paragraph pageId="24" pageNumber="25">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="harterti">Carasobarbus harterti</taxonomicName>
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has 31 to 38 scales in the lateral line (Table 5), usually 5.5 or 6.5 scales above the lateral line (Table 6), 4.5 to 6.5 scales below the lateral line (Table 7) and 13 to 17 scales around the least circumference of the caudal peduncle (Table 8). The scales are shown in Fig. 5.
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</paragraph>
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<paragraph pageId="24" pageNumber="25">The pharyngeal teeth count is -4.3.2 in four specimens examined. The pharyngeal teeth are hooked at their tips (Fig. 6).</paragraph>
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<paragraph pageId="24" pageNumber="25">Live specimens are silvery with an olive tinge and orange fins (Fig. 20). Ethanol-preserved specimens are yellow-brown, the back is darker than the belly and flanks.</paragraph>
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<paragraph pageId="24" pageNumber="25">The maximum length observed in the material examined is 250 mm SL.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="harterti">
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<pageBreakToken pageId="25" pageNumber="26" start="start">Carasobarbus</pageBreakToken>
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harterti
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</taxonomicName>
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differs from all congeners except
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus exulatus" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="exulatus">Carasobarbus exulatus</taxonomicName>
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and
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus fritschii" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="fritschii">Carasobarbus fritschii</taxonomicName>
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in having nine rather than 10 branched dorsal-fin rays. It differs from
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus exulatus" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="exulatus">Carasobarbus exulatus</taxonomicName>
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in having 31 to 38 scales in the lateral line vs 26 to 32 and modally 16 scales around the least circumference of the caudal peduncle vs. 12. It differs from
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus fritschii" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="fritschii">Carasobarbus fritschii</taxonomicName>
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in having a straight dorsal head profile and a last unbranched dorsal-fin ray that is strongly ossified and flexible for less than 15 % of its length vs. a convex dorsal head profile and a last unbranched dorsal-fin ray that is weakly ossified and flexible for more than 15 % of its length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="distribution">
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<paragraph pageId="25" pageNumber="26">Distribution.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="harterti">Carasobarbus harterti</taxonomicName>
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occurs in the rivers of the Oued Oum er Rbia and Tennsift drainage systems in Morocco (Fig. 18).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="habitats and biology">
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<paragraph pageId="25" pageNumber="26">Habitats and biology.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="harterti">Carasobarbus harterti</taxonomicName>
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is less common than
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus fritschii" order="Cypriniformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="fritschii">Carasobarbus fritschii</taxonomicName>
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and inhabits only the lower and middle course of big rivers.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="conservation status">
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<paragraph pageId="25" pageNumber="26">Conservation status.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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The IUCN rates this species as "Vulnerable A2ace" (
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<bibRefCitation author="Crivelli, AJ" journalOrPublisher="Environmental Biology of Fishes" pageId="43" pageNumber="44" title="IUCN (2012)" url="http://www.iucnredlist.org" year="2006 d">Crivelli 2006d</bibRefCitation>
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). The population has declined more than 30 % in the time from 1996 to 2006 due to urban, agricultural and industrial pollution (
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<bibRefCitation author="Crivelli, AJ" journalOrPublisher="Environmental Biology of Fishes" pageId="43" pageNumber="44" title="IUCN (2012)" url="http://www.iucnredlist.org" year="2006 d">Crivelli 2006d</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
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<paragraph pageId="25" pageNumber="26">Discussion.</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus harterti" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="harterti">
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<pageBreakToken pageId="26" pageNumber="27" start="start">Carasobarbus</pageBreakToken>
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harterti
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</taxonomicName>
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was described from Oued Oum er Rbia as
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus harterti" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="harterti">Barbus harterti</taxonomicName>
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(
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<bibRefCitation author="Guenther, A" journalOrPublisher="A Journal of Zoology in Connection with the Tring Museum" pageId="44" pageNumber="45" pagination="367 - 368" title="Second notice of new species of fishes from Morocco. Novitates Zoologicae." volume="8" year="1901">
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<normalizedToken originalValue="Günther">Guenther</normalizedToken>
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1901
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</bibRefCitation>
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). Some authors placed this species in the genus
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Barbus</taxonomicName>
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subgenus
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Labeobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labeobarbus" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Labeobarbus</taxonomicName>
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(
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<bibRefCitation author="Boulenger, GA" journalOrPublisher="Comptes Rendus Hebdomadaires des Seances de l'Academie des Sciences" pageId="42" pageNumber="43" pagination="1016 - 1018" title="La distribution en Afrique des Barbeaux du sous-genre Labeobarbus." volume="169" year="1919">Boulenger 1919</bibRefCitation>
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,
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<bibRefCitation author="Pellegrin, J" journalOrPublisher="Memoires de la Societe des Sciences Naturelles du Maroc" pageId="45" pageNumber="46" title="Les poissons des eaux douces de l'Afrique du Nord francaise Maroc: Algerie, Tunisie, Sahara." volume="1" year="1921">Pellegrin 1921</bibRefCitation>
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) while others continued using the genus
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Barbus</taxonomicName>
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||
(
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<bibRefCitation author="Pellegrin, JM" journalOrPublisher="Comptes rendus hebdomadaires des seances de l'Academie des sciences" pageId="46" pageNumber="47" pagination="809 - 811" title="Sur la faune ichthyologique des eaux douces du Maroc." volume="169" year="1919">Pellegrin 1919</bibRefCitation>
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,
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<bibRefCitation author="Pellegrin, J" journalOrPublisher="Bulletin de la Societe des sciences naturelles du Maroc" pageId="45" pageNumber="46" pagination="1 - 10" title="Les barbeaux de l'Afrique du nord Francaise: description d'une espece nouvelle." volume="19" year="1939">1939</bibRefCitation>
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).
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<bibRefCitation author="Karaman, MS" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="44" pageNumber="45" pagination="175 - 254" title="Suesswasserfische der Tuerkei. 8. Teil: Revision der Barben Europas, Vorderasiens und Nordafrikas." volume="67" year="1971">Karaman (1971)</bibRefCitation>
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synonymised it with
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus fritschii" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="fritschii">Carasobarbus fritschii</taxonomicName>
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, but regarded it as a distinct subspecies. He incorrectly synonymised
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus rothschildi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="rothschildi">Barbus rothschildi</taxonomicName>
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,
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus riggenbachi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="riggenbachi">Barbus riggenbachi</taxonomicName>
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and
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus paytonii" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="paytonii">Barbus paytonii</taxonomicName>
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with this subspecies and placed it in his newly erected genus
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Pseudotor" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotor" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Pseudotor</taxonomicName>
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. Subsequent authors did not accept
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<normalizedToken originalValue="Karaman’s">Karaman's</normalizedToken>
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proposal and continued using
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Barbus</taxonomicName>
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(
|
||
<bibRefCitation author="Fowler, HW" journalOrPublisher="Quarterly Journal of the Taiwan Museum" pageId="43" pageNumber="44" pagination="277 - 396" title="A Catalog of World Fishes (XXV)." volume="29" year="1976">Fowler 1976</bibRefCitation>
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,
|
||
<bibRefCitation author="El Gharbi, S" journalOrPublisher="Cahiers d'ethologie" pageId="43" pageNumber="44" pagination="223 - 226" title="Le genre Barbus (sous-genres Barbus et Labeobarbus) au Maroc. Genetique et parasitologie." volume="13" year="1993">El Gharbi et al. 1993</bibRefCitation>
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,
|
||
<bibRefCitation author="Azeroual, A" journalOrPublisher="Cybium" pageId="41" pageNumber="42" pagination="17 - 22" title="L'ichtyofaune des eaux continentals du Maroc." volume="23" year="2000">Azeroual et al. 2000</bibRefCitation>
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||
,
|
||
<bibRefCitation author="Leggatt, RA" journalOrPublisher="Reviews in Fish Biology and Fisheries" pageId="45" pageNumber="46" pagination="237 - 246" title="Occurrence of polyploidy in the fishes." url="10.1023/B:RFBF.0000033049.00668.fe" volume="13" year="2003">Leggatt and Iwama 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Colli, L" journalOrPublisher="Environmental Biology of Fishes" pageId="42" pageNumber="43" pagination="95 - 107" title="Molecular phylogeny of the blind cavefish Phreatichthys andruzzii and Garra barreimiae within the family Cyprinidae." url="10.1007/s10641-008-9393-z" volume="84" year="2009">Colli et al. 2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Borkenhagen, K" journalOrPublisher="Environmental Biology of Fishes" pageId="42" pageNumber="43" pagination="327 - 335" title="The molecular systematics of the Carasobarbus species from Iran and adjacent areas, with comments on Carasobarbus albus (Heckel, 1843)." url="10.1007/s10641-011-9787-1" volume="91" year="2011">Borkenhagen et al. 2011</bibRefCitation>
|
||
) or proposed using
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Labeobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labeobarbus" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Labeobarbus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Doadrio, I" journalOrPublisher="Annales du Musee royale de l'Afrique Centrale, Zoologie" pageId="43" pageNumber="44" pagination="21 - 34" title="Freshwater fish fauna of North Africa and its biogeography." volume="275" year="1994">Doadrio 1994</bibRefCitation>
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||
,
|
||
<bibRefCitation author="Tsigenopoulos, CS" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="46" pageNumber="47" pagination="851 - 856" title="Phylogenetic relationships of hexaploid large-sized barbs (genus Labeobarbus, Cyprinidae) based on mtDNA data." url="10.1016/j.ympev.2010.02.006" volume="56" year="2010">Tsigenopoulos et al. 2010</bibRefCitation>
|
||
). We transfer this species to the genus
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Carasobarbus</taxonomicName>
|
||
, based on the possession of a smooth last unbranched dorsal-fin ray, nine branched dorsal-fin rays, six branched rays in the anal fin and shield-shaped scales with numerous parallel radii. Analysis of molecular genetic characters (
|
||
<bibRefCitation author="Durand, JD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="43" pageNumber="44" pagination="91 - 100" title="Phylogeny and biogeography of the family Cyprinidae in the Middle East inferred from cytochrome b DNA - Evolutionary significance of this region." url="10.1006/mpev.2001.1040" volume="22" year="2002">Durand et al. 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Tsigenopoulos, CS" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="46" pageNumber="47" pagination="851 - 856" title="Phylogenetic relationships of hexaploid large-sized barbs (genus Labeobarbus, Cyprinidae) based on mtDNA data." url="10.1016/j.ympev.2010.02.006" volume="56" year="2010">Tsigenopoulos et al. 2010</bibRefCitation>
|
||
, KB unpublished data) support this decision.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |