230 lines
34 KiB
XML
230 lines
34 KiB
XML
<document id="64F3E756C164048B6D1D5802919D86C1" ID-CLB-Dataset="30537" ID-DOI="10.11646/zootaxa.4394.1.5" ID-GBIF-Dataset="0b5f9b4e-fceb-4175-a008-07b6aaa8a228" ID-ISSN="1175-5326" ID-Zenodo-Dep="1196942" ID-ZooBank="B37194D0-F2E7-46DE-B057-D6CDDA997B8B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520938471869" checkinUser="plazi" docAuthor="Salnitska, Maria & Solodovnikov, Aleхey" docDate="2018" docId="18024119C43A4D3052C1FD14920C5145" docLanguage="en" docName="zootaxa.4394.1.5.pdf.imf" docOrigin="Zootaxa 4394 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius przewalskii Riittir 1887" docType="treatment" docVersion="7" lastPageNumber="99" masterDocId="E43B3961C4384D345256FFEE91695379" masterDocTitle="Hypogean presumably sister species Quedius repentinus sp. n. from Altai and Q. roma from Sikhote-Alin (Coleoptera: Staphylinidae): a disjunct distribution or poorly sampled Siberia?" masterLastPageNumber="104" masterPageNumber="95" pageNumber="97" updateTime="1698529850080" updateUser="ExternalLinkService">
|
||
<mods:mods id="FBCD718DE0E00271EC160954F41CCA36" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3034655366E32890382F9FDC7CAB45DD">
|
||
<mods:title id="19869194010559085AEF5B2A1C631C92">Hypogean presumably sister species Quedius repentinus sp. n. from Altai and Q. roma from Sikhote-Alin (Coleoptera: Staphylinidae): a disjunct distribution or poorly sampled Siberia?</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A085156BBFF09B0156F723643C4723E2" type="personal">
|
||
<mods:role id="02A33960E944E16D918EE31EBA435E9D">
|
||
<mods:roleTerm id="3F894DDD46F5E2AC7A875DE65F09E999">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2CD1CD6E2B1C05A2C2EDCA5C3949F421">Salnitska, Maria</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="BF0473BD8479C4DC3A56CCBB84363DF4" type="personal">
|
||
<mods:role id="12617468ED1A8CC8D9E24D28E0613CD7">
|
||
<mods:roleTerm id="E1007F0491E906229B220B41AF547C21">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5F47C80B2F11F2117885BEB8537400B8">Solodovnikov, Aleхey</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="EADFE07F227A39A95C0B0AE8EF187C68">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9384C06CC2DC96B5D04289C1FEDDE39C" type="host">
|
||
<mods:titleInfo id="48680AADBF14480EFEE5B871D4A188C9">
|
||
<mods:title id="600D3A9A46C197E5E79F9EA289B1391F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="43C11E73225F96B4C0B69F44780FF65B">
|
||
<mods:date id="4324BBB6AE5A51CA22BC002AABE0D1BC">2018</mods:date>
|
||
<mods:detail id="F7647CC0E1D5F8A5F7715BF407671C78" type="pubDate">
|
||
<mods:number id="291D515A271CD8852382601205209A41">2018-03-13</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DEFF3E0ED0F9D1F389D9EEE01CD0A2A5" type="volume">
|
||
<mods:number id="16A5B0746570F02D5688FC11D1B62670">4394</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AC52D0C9CB91169E76A83FFF59205E40" type="issue">
|
||
<mods:number id="216FF37DA75C4CE1C1A72C939A0E7FFB">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="44FC3F9AD3FE0DFD9FDA64C9FFBC38F1" unit="page">
|
||
<mods:start id="3C5E9917F9C523C8E94F561A3A25D2AF">95</mods:start>
|
||
<mods:end id="0E37E097B2A903AA4F3A794690D8D8E6">104</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="9C22A40E8D425629E17D2E41DDCDCEF1">journal article</mods:classification>
|
||
<mods:identifier id="EB956E56837D4EF929151D479EF803C8" type="CLB-Dataset">30537</mods:identifier>
|
||
<mods:identifier id="B943FDBA2781BF7BDF77855ECEC3DB7B" type="DOI">10.11646/zootaxa.4394.1.5</mods:identifier>
|
||
<mods:identifier id="95A5CBE4DE7A59E5822EE6BA5CD2F57E" type="GBIF-Dataset">0b5f9b4e-fceb-4175-a008-07b6aaa8a228</mods:identifier>
|
||
<mods:identifier id="34A87BCEFE4A6F7BE13305D131936F5C" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="2D90F49D885C90924DDE5A25A08623AA" type="Zenodo-Dep">1196942</mods:identifier>
|
||
<mods:identifier id="238E2A7E8908D94D3F6E99EE0E582D3A" type="ZooBank">B37194D0-F2E7-46DE-B057-D6CDDA997B8B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="18024119C43A4D3052C1FD14920C5145" ID-DOI="http://doi.org/10.5281/zenodo.5990947" ID-GBIF-Taxon="142225610" ID-Zenodo-Dep="5990947" LSID="urn:lsid:plazi:treatment:18024119C43A4D3052C1FD14920C5145" httpUri="http://treatment.plazi.org/id/18024119C43A4D3052C1FD14920C5145" lastPageId="4" lastPageNumber="99" pageId="2" pageNumber="97">
|
||
<subSubSection id="D8B1A384C43A4D3652C1FD149359506D" box="[151,560,762,788]" pageId="2" pageNumber="97" type="nomenclature">
|
||
<paragraph id="9014F00FC43A4D3652C1FD149359506D" blockId="2.[151,560,762,788]" box="[151,560,762,788]" pageId="2" pageNumber="97">
|
||
<heading id="CB5C4763C43A4D3652C1FD149359506D" bold="true" box="[151,560,762,788]" fontSize="11" level="1" pageId="2" pageNumber="97" reason="1">
|
||
<taxonomicName id="57AB8B8CC43A4D3652C1FD149359506D" ID-CoL="4R2TW" authority="Riittir, 1887" authorityName="Riittir" authorityYear="1887" box="[151,560,762,788]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D3652C1FD149359506D" bold="true" box="[151,560,762,788]" pageId="2" pageNumber="97">
|
||
<emphasis id="A2DF2C1DC43A4D3652C1FD1490E3506D" bold="true" box="[151,394,762,788]" italics="true" pageId="2" pageNumber="97">Quedius przewalskii</emphasis>
|
||
Rеittеr, 1887
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8B1A384C43A4D3652C1FCAF9223576D" pageId="2" pageNumber="97" type="reference_group">
|
||
<paragraph id="9014F00FC43A4D3652C1FCAF9094502E" blockId="2.[151,509,833,855]" box="[151,509,833,855]" pageId="2" pageNumber="97">
|
||
<taxonomicName id="57AB8B8CC43A4D3652C1FCAF90A4502E" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" box="[151,461,833,855]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="lamus">
|
||
<emphasis id="A2DF2C1DC43A4D3652C1FCAF9046502E" box="[151,303,833,855]" italics="true" pageId="2" pageNumber="97">Quedius lamus</emphasis>
|
||
<bibRefCitation id="F43A8DFEC43A4D365362FCAF90A4502E" box="[308,461,833,855]" pageId="2" pageNumber="97" refId="ref4930" refString="Smetana, A. (1995) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 1. Some species of the genus Quedius Stephens, 1829, subgenus Microsaurus Dejean, 1833. Elytra (Tokyo), 23 (2), 235 - 244." type="journal article">Smetana, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
, 239
|
||
</paragraph>
|
||
<paragraph id="9014F00FC43A4D3652C1FC6A9223576D" blockId="2.[151,1436,900,1044]" pageId="2" pageNumber="97">
|
||
<bibRefCitation id="F43A8DFEC43A4D3652C1FC6A904A50E3" box="[151,291,900,922]" pageId="2" pageNumber="97" refId="ref4832" refString="Reitter, E. (1887) Insecta in itinere Cl. N. Przewalskii in Asia Centrali novisime lecta. VI. Clavicronia, Lamellicornia et Serricornia. Horae Societatis Entomologicae Rossicae, 21, 201 - 234." type="journal article">Reitter, 1887</bibRefCitation>
|
||
, 211 (original description);
|
||
<bibRefCitation id="F43A8DFEC43A4D365003FC6A92F650E3" box="[597,927,900,922]" pageId="2" pageNumber="97" refId="ref4718" refString="Bernhauer, M., and Schubert, K. (1916) Staphylinidae V. In S. Schenkling, Coleopterorum Catalogus, 5 (67), 409 - 498. [Berlin: Junk.]" type="journal article">Bernhauer and Schubert, 1916</bibRefCitation>
|
||
, 432 (catalog);
|
||
<bibRefCitation id="F43A8DFEC43A4D365606FC6A958F50E3" box="[1104,1254,900,922]" pageId="2" pageNumber="97" refId="ref4789" refString="Gridelli, E. (1924) Studi sul genere Quedius Steph. (Col. Staph.). Secondo contributo. Specie della regione Paleartica. Memorie della Societa Entomologica Italiana, 3 (1), 5 - 180." type="journal article">Gridelli, 1924</bibRefCitation>
|
||
, 71 (characters); Scheerpeltz, 1933, 1458 (catalog);
|
||
<bibRefCitation id="F43A8DFEC43A4D36506CFC4C93A850C1" box="[570,705,930,952]" pageId="2" pageNumber="97" refId="ref4757" refString="Bohac, J. (1988) New and little known Staphylininae (Coleoptera, Staphylinidae). Entomologicheskoe Obozrenie, 67, 549 - 557. [in Russian]." type="journal article">Boháč, 1988</bibRefCitation>
|
||
, 554 (cited as
|
||
<taxonomicName id="57AB8B8CC43A4D365133FC4D929650C1" box="[869,1023,930,952]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskyi">
|
||
<emphasis id="A2DF2C1DC43A4D365133FC4D929650C1" box="[869,1023,930,952]" italics="true" pageId="2" pageNumber="97">Q. przewalskyi</emphasis>
|
||
</taxonomicName>
|
||
; redescription based on
|
||
<typeStatus id="4F104EADC43A4D36575FFC4C940C50C1" box="[1289,1381,930,952]" pageId="2" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
; first illustration of aedeagus; records);
|
||
<bibRefCitation id="F43A8DFEC43A4D365078FC2F93A250AE" box="[558,715,961,983]" pageId="2" pageNumber="97" refId="ref4987" refString="Smetana, A. (1999) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 15. 15. Genus Strouhalium Scheerpeltz, 1962. Section 3. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 9. Elytra (Tokyo), 27 (2), 519 - 534." type="journal article">Smetana, 1999</bibRefCitation>
|
||
, 521, 524 (revision of the
|
||
<typeStatus id="4F104EADC43A4D3651BCFC2F952F50AE" box="[1002,1094,961,983]" pageId="2" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
; redescription; characters; notes; records; synonymy with
|
||
<taxonomicName id="57AB8B8CC43A4D365396FC0F9372508F" authorityName="Smetana" authorityYear="1995" box="[448,539,992,1014]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="lamus">
|
||
<emphasis id="A2DF2C1DC43A4D365396FC0F9372508F" box="[448,539,992,1014]" italics="true" pageId="2" pageNumber="97">Q. lamus</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="F43A8DFEC43A4D365067FC0E9200508F" box="[561,873,992,1014]" pageId="2" pageNumber="97" refId="ref5113" refString="Solodovnikov, A. & Hansen, A. K. (2016) Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini). Zootaxa, 4170 (3), 475 - 490. http: // dx. doi. org / 10.11646 / zootaxa. 4170.3.3" type="journal article">Solodovnikov & Hansen, 2016</bibRefCitation>
|
||
(notes; discussion on the composition of
|
||
<taxonomicName id="57AB8B8CC43A4D36575CFC0F94F5508F" ID-CoL="4R2TW" authorityName="Riittir" authorityYear="1887" box="[1290,1436,992,1014]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D36575CFC0F94F5508F" box="[1290,1436,992,1014]" italics="true" pageId="2" pageNumber="97">Q. przewalskii</emphasis>
|
||
</taxonomicName>
|
||
species group);
|
||
<bibRefCitation id="F43A8DFEC43A4D365331FC109369576D" box="[359,512,1022,1044]" pageId="2" pageNumber="97" refId="ref5054" refString="Smetana, A. (2014) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, series A, 21 (1 - 4), 1 - 39." type="journal article">Smetana, 2014</bibRefCitation>
|
||
, 13 (notes; distribution records).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8B1A384C43A4D3652C1FBAF929256BB" pageId="2" pageNumber="97" type="description">
|
||
<paragraph id="9014F00FC43A4D3652C1FBAF929256BB" blockId="2.[151,1437,1089,1870]" pageId="2" pageNumber="97">
|
||
The taxonomic history of
|
||
<taxonomicName id="57AB8B8CC43A4D365387FBAC93115720" authorityName="Riittir" authorityYear="1887" box="[465,632,1089,1113]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D365387FBAC93115720" box="[465,632,1089,1113]" italics="true" pageId="2" pageNumber="97">Q. przewalskii</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="57AB8B8CC43A4D3650BCFBAC92F85720" authorityName="Riittir" authorityYear="1887" box="[746,913,1089,1113]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D3650BCFBAC92F85720" box="[746,913,1089,1113]" italics="true" pageId="2" pageNumber="97">Q. przewalskii</emphasis>
|
||
</taxonomicName>
|
||
species group was recently summarized by
|
||
<bibRefCitation id="F43A8DFEC43A4D3652C1FB8B936D5707" box="[151,516,1125,1150]" pageId="2" pageNumber="97" refId="ref5113" refString="Solodovnikov, A. & Hansen, A. K. (2016) Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini). Zootaxa, 4170 (3), 475 - 490. http: // dx. doi. org / 10.11646 / zootaxa. 4170.3.3" type="journal article">Solodovnikov & Hansen (2016)</bibRefCitation>
|
||
. As follows from this paper, the diagnosis and composition of the
|
||
<taxonomicName id="57AB8B8CC43A4D3656AFFB8894F55704" authorityName="Riittir" authorityYear="1887" box="[1273,1436,1125,1149]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D3656AFFB8894F55704" box="[1273,1436,1125,1149]" italics="true" pageId="2" pageNumber="97">Q. przewalskii</emphasis>
|
||
</taxonomicName>
|
||
group remains ambiguous. Moreover, in view of the similar
|
||
<taxonomicName id="57AB8B8CC43A4D365106FB6492C657D8" authorityName="Solodovnikov et Hansen" authorityYear="2016" box="[848,943,1162,1185]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="roma">
|
||
<emphasis id="A2DF2C1DC43A4D365106FB6492C657D8" box="[848,943,1162,1185]" italics="true" pageId="2" pageNumber="97">Q. roma</emphasis>
|
||
</taxonomicName>
|
||
described in that paper, and the similar
|
||
<taxonomicName id="57AB8B8CC43A4D3657D7FB64919657BC" authorityName="Eppelsheim A.Smetana" authorityYear="1888" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="A2DF2C1DC43A4D3657D7FB64919657BC" italics="true" pageId="2" pageNumber="97">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
group newly defined in
|
||
<bibRefCitation id="F43A8DFEC43A4D365047FB4392E557BF" box="[529,908,1197,1222]" pageId="2" pageNumber="97" refId="ref4872" refString="Salnitska, M. & Solodovnikov, A. (2018) Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini). European Journal of Taxonomy, 401, 1 - 17. https: // doi. org / 10.5852 / ejt. 2018.401" type="journal article">Salnitska & Solodovnikov (2018)</bibRefCitation>
|
||
, it became apparent that the morphology of the parameral side of the median lobe (often best observed in lateral view, or when the paramere is dissected) is important for the identification of these species and species groups. In that respect the aedeagi of
|
||
<taxonomicName id="57AB8B8CC43A4D3656AEFB1894F55674" authorityName="Riittir" authorityYear="1887" box="[1272,1436,1269,1293]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D3656AEFB1894F55674" box="[1272,1436,1269,1293]" italics="true" pageId="2" pageNumber="97">Q. przewalskii</emphasis>
|
||
</taxonomicName>
|
||
and other species assigned to this group were not sufficiently described or illustrated, even though the former species was subject to several publications.
|
||
<bibRefCitation id="F43A8DFEC43A4D3650F7FAD39256562F" box="[673,831,1341,1366]" pageId="2" pageNumber="97" refId="ref4757" refString="Bohac, J. (1988) New and little known Staphylininae (Coleoptera, Staphylinidae). Entomologicheskoe Obozrenie, 67, 549 - 557. [in Russian]." type="journal article">Boháč (1988)</bibRefCitation>
|
||
provided somewhat distorted illustrations not fully matching the real structure, while
|
||
<bibRefCitation id="F43A8DFEC43A4D36507AFA8C938B5603" box="[556,738,1378,1402]" pageId="2" pageNumber="97" refId="ref4987" refString="Smetana, A. (1999) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 15. 15. Genus Strouhalium Scheerpeltz, 1962. Section 3. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 9. Elytra (Tokyo), 27 (2), 519 - 534." type="journal article">Smetana (1999)</bibRefCitation>
|
||
did not provide the lateral view of the aedeagus at all. To facilitate informative comparisons for the new species described in this paper and in the future, here we redescribe the
|
||
<typeStatus id="4F104EADC43A4D365296FA47904B56BB" box="[192,290,1449,1474]" pageId="2" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="57AB8B8CC43A4D365311FA44908356B8" authorityName="Riittir" authorityYear="1887" box="[327,490,1449,1473]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskii">
|
||
<emphasis id="A2DF2C1DC43A4D365311FA44908356B8" box="[327,490,1449,1473]" italics="true" pageId="2" pageNumber="97">Q. przewalskii</emphasis>
|
||
</taxonomicName>
|
||
and provide such missing illustrations (
|
||
<figureCitation id="0890EC8AC43A4D3651FEFA47928456BB" box="[936,1005,1449,1474]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="2" pageNumber="97">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8B1A384C43A4D365291FA2393415557" pageId="2" pageNumber="97" type="materials_examined">
|
||
<paragraph id="9014F00FC43A4D365291FA2393415557" blockId="2.[151,1437,1089,1870]" pageId="2" pageNumber="97">
|
||
<emphasis id="A2DF2C1DC43A4D365291FA239328569F" bold="true" box="[199,577,1485,1510]" pageId="2" pageNumber="97">
|
||
<typeStatus id="4F104EADC43A4D365291FA23906B569F" box="[199,258,1485,1510]" pageId="2" pageNumber="97">Type</typeStatus>
|
||
material examined:
|
||
<collectingCountry id="E8BCB09FC43A4D3653AEFA239328569F" box="[504,577,1485,1510]" name="China" pageId="2" pageNumber="97">China</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="A2DF2C1DC43A4D365019FA2393D5569F" bold="true" box="[591,700,1485,1510]" pageId="2" pageNumber="97">
|
||
<typeStatus id="4F104EADC43A4D365019FA2393D5569F" box="[591,700,1485,1510]" pageId="2" pageNumber="97" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="86AD3B86C43A4D36509BFA20938C569F" box="[717,741,1486,1510]" pageId="2" pageNumber="97" type="male">♂</specimenCount>
|
||
): ”Prj 84 [green label]/ VII Huangho super. [superior]
|
||
<date id="E415D6CFC43A4D365711FA2094E5569F" box="[1351,1420,1486,1510]" pageId="2" pageNumber="97" value="2013-02-01">13 1/2</date>
|
||
– 14000 p. [handwritten label]/
|
||
<taxonomicName id="57AB8B8CC43A4D3653BFFA1F92205570" box="[489,841,1521,1546]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="species" species="przewalskyi" subGenus="Ediquus">
|
||
<emphasis id="A2DF2C1DC43A4D3653BFFA1F932E5570" box="[489,583,1521,1545]" italics="true" pageId="2" pageNumber="97">Quedius</emphasis>
|
||
(
|
||
<emphasis id="A2DF2C1DC43A4D36500EFA1F93DD5570" box="[600,692,1521,1545]" italics="true" pageId="2" pageNumber="97">Ediquus</emphasis>
|
||
)
|
||
<emphasis id="A2DF2C1DC43A4D365093FA1F92205570" box="[709,841,1521,1545]" italics="true" pageId="2" pageNumber="97">przewalskyi</emphasis>
|
||
</taxonomicName>
|
||
[sic!] m. n. sp. unicum [handwritten label]/
|
||
<typeStatus id="4F104EADC43A4D365715FA1C94F55573" box="[1347,1436,1522,1546]" pageId="2" pageNumber="97">
|
||
<collectionCode id="F6BA68CAC43A4D365715FA1C94F55573" box="[1347,1436,1522,1546]" pageId="2" pageNumber="97">TYPUS</collectionCode>
|
||
</typeStatus>
|
||
[red printed label]” (
|
||
<figureCitation id="0890EC8AC43A4D36532BF9FB90B95557" box="[381,464,1557,1582]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="2" pageNumber="97">Fig. 2F</figureCitation>
|
||
) (
|
||
<collectionCode id="F6BA68CAC43A4D3653B1F9F893725557" box="[487,539,1558,1582]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="97">ZIN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8B1A384C43A4D305291F9D7920C5145" lastPageId="4" lastPageNumber="99" pageId="2" pageNumber="97" type="description">
|
||
<paragraph id="9014F00FC43A4D365291F9D79518550F" blockId="2.[151,1437,1089,1870]" pageId="2" pageNumber="97">
|
||
<emphasis id="A2DF2C1DC43A4D365291F9D7901D552B" bold="true" box="[199,372,1593,1618]" pageId="2" pageNumber="97">Redescription.</emphasis>
|
||
Measurements and ratios: HL: 1.93; HW: 1.89; PL: 2.25; PW: 2.59; EL: 1.98; EW: 2.66; FB: 6.16; TL: 15.7; HL/HW: 1.02; PL/PW: 0.87; EL/EW: 0.92; PL/EL: 1.24; PW/EW: 0.97.
|
||
</paragraph>
|
||
<paragraph id="9014F00FC43A4D365291F96F92775437" blockId="2.[151,1437,1089,1870]" pageId="2" pageNumber="97">
|
||
Body light brown (somewhat reddish, possibly because of the age of the specimen according to
|
||
<bibRefCitation id="F43A8DFEC43A4D36576AF96C918A55C7" pageId="2" pageNumber="97" refId="ref4930" refString="Smetana, A. (1995) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 1. Some species of the genus Quedius Stephens, 1829, subgenus Microsaurus Dejean, 1833. Elytra (Tokyo), 23 (2), 235 - 244." type="journal article">Smetana (1995)</bibRefCitation>
|
||
, rather unicolorous, appendages only slightly lighter, anterior margins of abdominal tergites dark brown; elytra and abdomen with dense yellowish pubescence; legs rather long, body flattened dorso-ventrally. Head barely longer than wide with distinctly rounded posterior angles; with dense and shallow microsculpture of transverse waves uniformly covering the whole head. Eyes small and flat, slightly protruding from lateral contour of head; temples about 2.7 as long as longitudinal diameter of eye.
|
||
</paragraph>
|
||
<caption id="C4D4A087C43B4D3752C1FB8A902457C1" httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="3" pageNumber="98" startId="3.[151,250,1124,1146]" targetBox="[216,1353,187,1076]" targetPageId="3">
|
||
<paragraph id="9014F00FC43B4D3752C1FB8A902457C1" blockId="3.[151,1435,1124,1208]" pageId="3" pageNumber="98">
|
||
<emphasis id="A2DF2C1DC43B4D3752C1FB8A907F5700" bold="true" box="[151,278,1124,1146]" pageId="3" pageNumber="98">FIGURE 2.</emphasis>
|
||
<emphasis id="A2DF2C1DC43B4D375377FB8A90935703" box="[289,506,1124,1146]" italics="true" pageId="3" pageNumber="98">
|
||
<taxonomicName id="57AB8B8CC43B4D375377FB8A909F5703" authorityName="Riittir" authorityYear="1887" box="[289,502,1124,1146]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="przewalskii">Quedius przewalskii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels. Scale bars: A–C, E–G = 1 mm, D = 0.4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9014F00FC43B4D375291FAED91BF56B6" blockId="3.[151,1437,1283,2028]" pageId="3" pageNumber="98">Head disk on each side with the following setiferous punctures: anterior frontal puncture situated very close to inner margin of eye, posterior frontal puncture at about the same distance from posterior margin of eye and nuchal ridge, and a pair of small barely visible vertical punctures between nuchal ridge and posterior frontal puncture. Each temple with two temporal punctures of which the posterior temporal puncture is closer to posterior margin of head than to posterior margin of eye; additionally, temples with fine setiferous punctures bearing short yellowish setae.</paragraph>
|
||
<paragraph id="9014F00FC43B4D375291FA3290155545" blockId="3.[151,1437,1283,2028]" pageId="3" pageNumber="98">Antennae moderately long, with antennal segments: third distinctly longer than second, fourth and fifth as long as wide, sixth to tenth wider than long gradually widening towards apex of antenna, last segment longer than preceding segments.</paragraph>
|
||
<paragraph id="9014F00FC43B4D375291F9A690C85589" blockId="3.[151,1437,1283,2028]" pageId="3" pageNumber="98">Pronotum barely wider than long, widest at about middle, more strongly narrowing anteriad than posteriad; posterior lateral portions distinctly explanate. Dorsal rows of setae each with three equal sized small punctures, the basalmost puncture located at a level slightly before large lateral puncture; sublateral group with two punctures; with three shallow hardly visible punctures along basal margin at each side of pronotum; microsculpture slightly shallower than on head.</paragraph>
|
||
<paragraph id="9014F00FC43B4D375291F9159204546D" blockId="3.[151,1437,1283,2028]" box="[199,877,1787,1812]" pageId="3" pageNumber="98">Scutellum impunctate, with faint transversal microsculpture.</paragraph>
|
||
<paragraph id="9014F00FC43B4D375291F8F1952E54F9" blockId="3.[151,1437,1283,2028]" pageId="3" pageNumber="98">Elytra widening posteriad, very short and wider than long; slightly wider and notably shorter than pronotum; punctation fine and dense with interspaces between setiferous punctures as large as diameter of punctures; interspaces shiny, with minute irregularities; pubescence yellowish. Wings vestigial.</paragraph>
|
||
<paragraph id="9014F00FC43B4D375291F86591885495" blockId="3.[151,1437,1283,2028]" pageId="3" pageNumber="98">Abdomen: punctation fine and dense, becoming sparser toward posterior part of each tergite and in general toward apex of abdomen; interspaces with minute irregularities; posterior margin of tergite VII without palisade fringe.</paragraph>
|
||
<paragraph id="9014F00FC43C4D305291FF79920C5145" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="99">
|
||
First four segments of front tarsus dilated; second segment as wide as apex of tibia. Sternite VIII with three long setae on each side (on the left side one is broken off), with shallow subangulate triangular medio-apical emargination, area before emargination slightly depressed (
|
||
<figureCitation id="0890EC8AC43C4D30517BFF3192E15381" box="[813,904,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2G</figureCitation>
|
||
). Sternite IХ asymmetrical, elongate, with wide and glabrous basal portion, and with sparsely setose and weakly emarginated apical portion (
|
||
<figureCitation id="0890EC8AC43C4D305695FEEA94735265" box="[1219,1306,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2F</figureCitation>
|
||
). Tergite Х triangular, gradually narrowed apicad, with numerous variably long setae apically (
|
||
<figureCitation id="0890EC8AC43C4D305614FEC995F25239" box="[1090,1179,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2E</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="0890EC8AC43C4D30577BFEC991C6521D" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Figs. 2B– D</figureCitation>
|
||
): median lobe (in parameral or anteparameral view) narrowed into distinctly asymmetrical, pointed apex (
|
||
<figureCitation id="0890EC8AC43C4D305739FEA291D252F1" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2C</figureCitation>
|
||
); (in lateral view) with slightly arcuate and elongated narrow apical portion with subacutely triangular apex and minute tooth situated close to apex (
|
||
<figureCitation id="0890EC8AC43C4D305016FE7A93F552D5" box="[576,668,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2B</figureCitation>
|
||
). Paramere (in parameral view) much narrower than median lobe, asymmetrical, gradually narrowed apicad, with distinct apical emargination (
|
||
<figureCitation id="0890EC8AC43C4D3051ABFE5995EC52A9" box="[1021,1157,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Figs. 2C, D</figureCitation>
|
||
); with one pair of apical setae on each side of emargination and two pairs of lateral setae (
|
||
<figureCitation id="0890EC8AC43C4D3051FFFE329560528D" box="[937,1033,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2D</figureCitation>
|
||
); (underside) with 13 sensory peg arranged in two dense groups on each side of apical emargination; (in lateral view) nearly reaching apex of median lobe (
|
||
<figureCitation id="0890EC8AC43C4D305280FDCA90475145" box="[214,302,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[216,1353,187,1076]" captionTargetId="figure@3.[216,1353,187,1076]" captionTargetPageId="3" captionText="FIGURE 2. Quedius przewalskii, holotype: A, habitus; B, aedeagus (laterally); C, same (in parameral view); D, apex of paramere, underside (side facing median lobe); E, tergite X; F, sternite IX; G, sternite VIII; H, labels.Scale bars:A–C, E–G = 1 mm, D = 0.4 mm." httpUri="https://zenodo.org/record/1196946/files/figure.png" pageId="4" pageNumber="99">Fig. 2B</figureCitation>
|
||
). Internal sac without larger sclerotized structures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |