199 lines
18 KiB
XML
199 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.962.52759" ID-GBIF-Dataset="0aa621ef-de45-41be-8c10-73ba877577b9" ID-PMC="PMC7471473" ID-Pensoft-Pub="1313-2970-962-1" ID-Pensoft-UUID="AD14C60754A7579492E50291C659E3C6" ID-PubMed="32943974" ID-ZooBank="CA7DEA7CBFEB4BC8A598A802941FD59F" ModsDocID="1313-2970-962-1" checkinTime="1598026755870" checkinUser="pensoft" docAuthor="Laudee, Pongsak, Seetapan, Kriengkrai, Vongsombath, Chanda & Malicky, Hans" docDate="2020" docId="1292136E4EBC5481968F185FD740F3F7" docLanguage="en" docName="ZooKeys 962: 1-11" docOrigin="ZooKeys 962" docSource="http://dx.doi.org/10.3897/zookeys.962.52759" docTitle="Ecnomus petchanaae Laudee & Malicky 2020, sp. nov." docType="treatment" docUuid="61596C7B-1396-41B1-9DC1-60E344674F81" docUuidSource="ZooBank" docVersion="4" id="AD14C60754A7579492E50291C659E3C6" lastPageNumber="1" masterDocId="AD14C60754A7579492E50291C659E3C6" masterDocTitle="New species of caddisflies (Trichoptera, Ecnomidae, Polycentropodidae, Psychomyiidae) from Mekong tributaries, Laos" masterLastPageNumber="11" masterPageNumber="1" pageNumber="1" updateTime="1668169335156" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of caddisflies (Trichoptera, Ecnomidae, Polycentropodidae, Psychomyiidae) from Mekong tributaries, Laos</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Laudee, Pongsak</mods:namePart>
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<mods:affiliation>Department of Fishery and Costal Resources, Faculty of Science and Industrial Technology, Prince of Songkla University, Surat Thani Campus, Muang District, Surat Thani Province, Thailand</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3819-7980</mods:nameIdentifier>
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<mods:nameIdentifier type="email">pongsak.l@psu.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Seetapan, Kriengkrai</mods:namePart>
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<mods:affiliation>School of Agriculture and Natural Resources, University of Phayao, Tumbol Maeka, Muang District, Phayao Province, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vongsombath, Chanda</mods:namePart>
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<mods:affiliation>Faculty of Environmental Sciences, National University of Laos, Dong Dok Campus, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Malicky, Hans</mods:namePart>
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<mods:affiliation>Sonnengasse 13, A- 3293 Lunz am See, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>962</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>11</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.962.52759</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.962.52759</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-962-1</mods:identifier>
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<mods:identifier type="ZooBank">CA7DEA7CBFEB4BC8A598A802941FD59F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">AD14C60754A7579492E50291C659E3C6</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="167287972" LSID="urn:lsid:zoobank.org:act:61596C7B-1396-41B1-9DC1-60E344674F81" httpUri="http://treatment.plazi.org/id/1292136E4EBC5481968F185FD740F3F7" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="1292136E-4EBC-5481-968F-185FD740F3F7" authority="Laudee & Malicky" authorityName="Laudee & Malicky" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Ecnomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecnomus petchanaae" order="Trichoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petchanaae" status="sp. nov.">Ecnomus petchanaae Laudee & Malicky</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">Figure 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The male genitalia of
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<taxonomicName lsidName="E. petchanaae" pageId="0" pageNumber="1" rank="species" species="petchanaae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. petchanaae</emphasis>
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</taxonomicName>
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sp. nov. are similar to
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<taxonomicName authorityName="Cartwright" authorityYear="1994" class="Insecta" family="Ecnomidae" genus="Ecnomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecnomus gapit" order="Trichoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gapit">
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<emphasis italics="true" pageId="0" pageNumber="1">Ecnomus gapit</emphasis>
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</taxonomicName>
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Cartwright, 1994,
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<taxonomicName lsidName="E. yuleae" pageId="0" pageNumber="1" rank="species" species="yuleae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. yuleae</emphasis>
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</taxonomicName>
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Cartwright, 1994,
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<taxonomicName lsidName="E. dares" pageId="0" pageNumber="1" rank="species" species="dares">
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<emphasis italics="true" pageId="0" pageNumber="1">E. dares</emphasis>
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</taxonomicName>
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Malicky, 2000, and
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<taxonomicName lsidName="E. perseis" pageId="0" pageNumber="1" rank="species" species="perseis">
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<emphasis italics="true" pageId="0" pageNumber="1">E. perseis</emphasis>
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</taxonomicName>
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Malicky, 2008 described from Borneo. The superior appendages of all these species, including the new species, are particularly large and the subapical part of the superior appendages is covered by numerous spiny setae. However,
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<taxonomicName lsidName="E. petchanaae" pageId="0" pageNumber="1" rank="species" species="petchanaae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. petchanaae</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished by the shape of its superior appendages, which, in lateral view, are slender and club-shaped, but basally broad in
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<taxonomicName lsidName="E. gapit" pageId="0" pageNumber="1" rank="species" species="gapit">
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<emphasis italics="true" pageId="0" pageNumber="1">E. gapit</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. yuleae" pageId="0" pageNumber="1" rank="species" species="yuleae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. yuleae</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. dares" pageId="0" pageNumber="1" rank="species" species="dares">
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<emphasis italics="true" pageId="0" pageNumber="1">E. dares</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="E. perseis" pageId="0" pageNumber="1" rank="species" species="perseis">
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<emphasis italics="true" pageId="0" pageNumber="1">E. perseis</emphasis>
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</taxonomicName>
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. In addition, in ventral view of the outer surface of the inferior appendages of the new species is crescent-shaped, whereas they are curved and claw-shaped in
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<taxonomicName lsidName="E. gapit" pageId="0" pageNumber="1" rank="species" species="gapit">
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<emphasis italics="true" pageId="0" pageNumber="1">E. gapit</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. yuleae" pageId="0" pageNumber="1" rank="species" species="yuleae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. yuleae</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. dares" pageId="0" pageNumber="1" rank="species" species="dares">
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<emphasis italics="true" pageId="0" pageNumber="1">E. dares</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="E. perseis" pageId="0" pageNumber="1" rank="species" species="perseis">
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<emphasis italics="true" pageId="0" pageNumber="1">E. perseis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
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<taxonomicName authorityName="Laudee & Malicky" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Ecnomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecnomus petchanaae" order="Trichoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petchanaae">
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<emphasis italics="true" pageId="0" pageNumber="1">Ecnomus petchanaae</emphasis>
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</taxonomicName>
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, sp. nov. Male genitalia.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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Segment IX and superior appendages, dorsal
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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segments IX and superior appendages, left lateral
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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segment IX and inferior appendages, ventral
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult, male, length of each male forewing 5.6-6.0 mm; color in alcohol of head, thorax, forewings, abdomen, and legs brown. Male genitalia as in Figure
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1A-D</figureCitation>
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. Tergum IX somewhat square, anterior margin truncated, posterior margin bilobed in dorsal view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1A</figureCitation>
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); trapezoid and rounded anterodorsally in lateral view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1B</figureCitation>
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). Sternum IX ovoid in lateral view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1B</figureCitation>
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); rectangular with
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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concave incision posteriorly, bilobed and rounded anteriorly in ventral view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1C</figureCitation>
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). Superior appendages long, slender, with expanded base, curved inward posteriorly with numerous long spiny setae subapically in dorsal view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1A</figureCitation>
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); in lateral view, superior appendages, relatively large, long, slightly curved upward, bulb-like apically, with numerous spiny setae (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1B</figureCitation>
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). Inferior appendages tubular, bent inward, beak-like apically in lateral view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1B</figureCitation>
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); in ventral view, crescent-shaped, with a submediate knot, overlapping each other subapically (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1C</figureCitation>
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). Phallus long, tubular, curved upward, pointed apex with dorsal process in lateral view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1B</figureCitation>
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); in ventral view, bulb-like, with pointed apex (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ecnomus petchanaae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segments IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus tip, ventral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure1" httpUri="https://binary.pensoft.net/fig/443670" pageId="0" pageNumber="1">1D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Male. Laos: Pakse Province
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</emphasis>
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: Paksong, Vang Ngao River,
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<geoCoordinate degrees="15" direction="north" minutes="11" orientation="latitude" precision="15" seconds="37" value="15.193611">15°11'37"N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="06" orientation="longitude" precision="15" seconds="40" value="106.11111">106°06'40"E</geoCoordinate>
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, elev. 920 m, 7.iv.2019, Pongsak Laudee. (PSUNHM).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: same data as the holotype, 3 males: 1 male (PSUNHM), 1 male (CHM), 1 male (NMPC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species epithet honors Mrs Kanchanaluk Petchana, Director of Administration and Strategic Development Division, Prince of Songkla University, Surat Thani Campus.</paragraph>
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</subSubSection>
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</treatment>
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</document> |