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<document id="8ACEEC3CA42F52EED76236C8D9B2AB48" ID-DOI="10.24199/j.mmv.2005.62.5" ID-ISSN="1447-2554" ID-Zenodo-Dep="12208025" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718988401551" checkinUser="felipe" docAuthor="O'Loughlin, P. Mark &amp; Ahearn, Cynthia" docDate="2005" docId="116087F5FFA7FFF0FCA97D21FF0D1D03" docLanguage="en" docName="MemMusVictoria.62.2.147-179.pdf" docOrigin="Memoirs of Museum Victoria 62 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-62-issue-2-2005/pages-147-179/" docStyle="DocumentStyle:99484BB52C5EA01CDE4EA5EECB5782C2.2:MemMusVictoria.2003-.journal_article.type2" docStyleId="99484BB52C5EA01CDE4EA5EECB5782C2" docStyleName="MemMusVictoria.2003-.journal_article.type2" docStyleVersion="2" docTitle="Molpadiodemas pediculus O'Loughlin &amp; Ahearn, 2005, sp. nov." docType="treatment" docVersion="2" lastPageNumber="163" masterDocId="ED59FF8DFFA8FFE0FFB7783EFFFE1F73" masterDocTitle="A review of pygal-furrowed Synallactidae (Echinodermata: Holothuroidea), with new species from the Antarctic, Atlantic and Pacific oceans" masterLastPageNumber="179" masterPageNumber="147" pageNumber="162" updateTime="1718989876917" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="10CCB9BE6EC030801C1EE3372EDEE583" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0BA52BAA601C4F9E99194DC874EA151B">
<mods:title id="63182DB1E6CC359A36F47795F4F631CA">A review of pygal-furrowed Synallactidae (Echinodermata: Holothuroidea), with new species from the Antarctic, Atlantic and Pacific oceans</mods:title>
</mods:titleInfo>
<mods:name id="71949D9C72DDCFB1162B80518FA80CAF" type="personal">
<mods:role id="3BAABB211DEEB8EA028782FC1E2CF541">
<mods:roleTerm id="47E0EF484354BF32A986C7E09ACC120F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5425E59D15EA3412321560BF34BC252E">O'Loughlin, P. Mark</mods:namePart>
</mods:name>
<mods:name id="DA5030C3AF4106958C22375E8510B9AF" type="personal">
<mods:role id="51629B290A35FD3A4EE9B1C934E86474">
<mods:roleTerm id="5D7AD3F5D92DA80B44FB5313873FD83D">Author</mods:roleTerm>
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<mods:namePart id="ACA95F682E89CC34D55D2A5F630209D1">Ahearn, Cynthia</mods:namePart>
</mods:name>
<mods:typeOfResource id="131F91437BF21BAE5153757CAC4B8689">text</mods:typeOfResource>
<mods:relatedItem id="9564DF0183A504EACE999759F5DD6CAF" type="host">
<mods:titleInfo id="738306E774E6477A9E356621D6532A79">
<mods:title id="F46560359329C42AA01ADFCA76964584">Memoirs of Museum Victoria</mods:title>
</mods:titleInfo>
<mods:part id="A4CE280775B1B01EAC4087574181F21E">
<mods:date id="F9A2C5A9239FEBBD09FE0825F6D32ECD">2005</mods:date>
<mods:detail id="C972D0730EC2BFA476B33FFEC5D46DD1" type="pubDate">
<mods:number id="DD7C1864359DAAD78D147BCB7E529399">2005-12-31</mods:number>
</mods:detail>
<mods:detail id="EE8E122183693EE876BA862658CA366A" type="volume">
<mods:number id="BE87CE02DB183FD9B1AA221550EF98A9">62</mods:number>
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<mods:detail id="A35C115818945EE35C07AFBA8FEDCC8B" type="issue">
<mods:number id="C4DD4019B9CCF06AE82703CDEDFECD22">2</mods:number>
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<mods:extent id="82A38491CF33EECF53ABEE0F25491DFC" unit="page">
<mods:start id="F3E709D649D874E0FC668622EA15AFD7">147</mods:start>
<mods:end id="37242D0CBB3B9C1AF62C1970BEC55E1E">179</mods:end>
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<mods:url id="C418DD146853593035EEB19879561E70">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-62-issue-2-2005/pages-147-179/</mods:url>
</mods:location>
<mods:classification id="5AF1BDE4DEBFEC0D7647AF08174BC162">journal article</mods:classification>
<mods:identifier id="1AAAE101FD234814E6D13829F927D42B" type="DOI">10.24199/j.mmv.2005.62.5</mods:identifier>
<mods:identifier id="C200EBBC80E0263685C150A0C93AAFDE" type="ISSN">1447-2554</mods:identifier>
<mods:identifier id="4573DFE60E41C5D8B81F4A0CFF9837EB" type="Zenodo-Dep">12208025</mods:identifier>
</mods:mods>
<treatment id="116087F5FFA7FFF0FCA97D21FF0D1D03" LSID="urn:lsid:plazi:treatment:116087F5FFA7FFF0FCA97D21FF0D1D03" httpUri="http://treatment.plazi.org/id/116087F5FFA7FFF0FCA97D21FF0D1D03" lastPageId="16" lastPageNumber="163" pageId="15" pageNumber="162">
<subSubSection id="D1D36568FFA7FFEFFCA97D21FB7E1A46" box="[798,1152,1311,1333]" pageId="15" pageNumber="162" type="nomenclature">
<paragraph id="997636E3FFA7FFEFFCA97D21FB7E1A46" blockId="15.[798,1152,1311,1333]" box="[798,1152,1311,1333]" pageId="15" pageNumber="162">
<heading id="C23E818FFFA7FFEFFCA97D21FB7E1A46" box="[798,1152,1311,1333]" fontSize="9" level="2" pageId="15" pageNumber="162" reason="5">
<taxonomicName id="5EC94D60FFA7FFEFFCA97D21FBD21A46" ID-CoL="43YMT" authorityName="O'Loughlin &amp; Ahearn" authorityYear="2005" box="[798,1068,1311,1333]" class="Holothuroidea" family="Molpadiodemidae" genus="Molpadiodemas" kingdom="Animalia" order="Persiculida" pageId="15" pageNumber="162" phylum="Echinodermata" rank="species" species="pediculus" status="sp. nov.">
<emphasis id="ABBDEAF1FFA7FFEFFCA97D21FBD21A46" bold="true" box="[798,1068,1311,1333]" italics="true" pageId="15" pageNumber="162">Molpadiodemas pediculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B08E578AFFA7FFEFFB857D21FB7E1A46" box="[1074,1152,1311,1333]" pageId="15" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D1D36568FFA7FFF0FCA97D74FF0D1D03" lastPageId="16" lastPageNumber="163" pageId="15" pageNumber="162" type="description">
<paragraph id="997636E3FFA7FFEFFCA97D74FC1F1A13" blockId="15.[798,993,1354,1376]" box="[798,993,1354,1376]" pageId="15" pageNumber="162">
<figureCitation id="01F22A66FFA7FFEFFCA97D74FC5A1A13" box="[798,932,1354,1376]" captionStart="Figure 5" captionStartId="9.[130,187,1647,1667]" captionTargetBox="[129,1432,297,1621]" captionTargetId="figure-26@9.[569,995,833,1087]" captionTargetPageId="9" captionText="Figure 5. a–d, Meseres morbillus sp. nov., a, lateral view (USNM 1008093, paratype, 91 mm long); b, tentacles, ventrolateral tube feet “spotting” (NMV F101865); c, tentacles, digits (USNM E49243); d, calcareous ring (radial plate right half) (USNM 1008093, paratype). e–g, M. neovillosus sp. nov.; e, dorsal view (USNM 1008458, paratype, 71 mm long); f, pygal end, tube feet (USNM 1008458); g, ventrolateral tube feet (BMNH 86.10.2.151, holotype). h–k, M. pediculus sp. nov.; h, dorsal view, elongate form (USNM 1008318, holotype, 56 mm long); i, ventrolateral tube feet (NMV F104796, paratype); j, tentacles (holotype); k, broadly attached, relatively narrow, slightly thickened, longitudinal muscles (arrows) (paratype). l–m, M. porphyrus sp. nov. (USNM E38795, holotype, 114 mm long); l, dorsal view, thick rounded brim; m, gonad. n–o, M. translucens sp. nov. (USNM E48652, holotype, 78 mm long); n, dorsal view, longitudinal muscles visible through translucent body wall (arrows); o, tentacles." figureDoi="http://doi.org/10.5281/zenodo.12208036" httpUri="https://zenodo.org/record/12208036/files/figure.png" pageId="15" pageNumber="162">Figures 5h–k</figureCitation>
,
<figureCitation id="01F22A66FFA7FFEFFC187D74FC1F1A13" box="[943,993,1354,1376]" captionStart="Figure 8" captionStartId="19.[130,188,1603,1623]" captionTargetBox="[127,1432,295,1577]" captionTargetPageId="19" captionText="Figure 8. Tentacle ossicles (SEM). a–d, Meseres morbillus sp. nov. (USNM E49251). e–f, M. neovillosus sp. nov. (USNM 1008458, paratype). g–h, M. pediculus sp. nov. (USNM 1008318, holotype). i–l, M. porphyrus sp. nov. (USNM E38795, holotype). m–p, M. translucens sp. nov. (USNM E48652, holotype). q–t, M. villosus, q–r (USNM E48640), s–t (USNM E9929). u–x, M. violaceus, u–v (USNM 1022608), w–x (USNM E49366)." figureDoi="http://doi.org/10.5281/zenodo.12208042" httpUri="https://zenodo.org/record/12208042/files/figure.png" pageId="15" pageNumber="162">
<quantity id="5E319B06FFA7FFEFFC187D74FC381A13" box="[943,966,1354,1376]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="15" pageNumber="162" unit="g" value="8.0">8g</quantity>
–h
</figureCitation>
</paragraph>
<paragraph id="997636E3FFA7FFEFFCA97D4AFB9A1ACF" blockId="15.[798,1429,1395,1547]" pageId="15" pageNumber="162">
<emphasis id="ABBDEAF1FFA7FFEFFCA97D4AFC321AF4" box="[798,972,1396,1415]" italics="true" pageId="15" pageNumber="162">Material examined</emphasis>
.
<materialsCitation id="29A13CBEFFA7FFEFFC6A7D4DFB9A1ACF" collectingDate="1963-04-29" collectionCode="USNM" collectorName="RV Eltanin &amp; Cr" elevation="5444" latitude="-55.116665" location="South Atlantic Ocean" longLatPrecision="1066" longitude="-25.983334" pageId="15" pageNumber="162" specimenCode="USNM 1008318" specimenCount="1" typeStatus="holotype">
<typeStatus id="46728841FFA7FFEFFC6A7D4DFBCD1AF4" box="[989,1075,1395,1415]" pageId="15" pageNumber="162" type="holotype">Holotype</typeStatus>
.
<location id="9C166038FFA7FFEFFBF77D4DFAF21AF4" LSID="urn:lsid:plazi:treatment:116087F5FFA7FFF0FCA97D21FF0D1D03:9C166038FFA7FFEFFBF77D4DFAF21AF4" box="[1088,1292,1395,1415]" latitude="-55.116665" longLatPrecision="1066" longitude="-25.983334" name="South Atlantic Ocean" pageId="15" pageNumber="162">South Atlantic Ocean</location>
,
<locationDeviation id="88C42FC1FFA7FFEFFAAC7D4AFC6E1AD2" location="South Sandwich Is" pageId="15" pageNumber="162">NE of South Sandwich Is</locationDeviation>
,
<geoCoordinate id="FCFD5024FFA7FFEFFC2B7DB0FC1A1AD1" box="[924,996,1422,1442]" degrees="55" direction="south" minutes="07" orientation="latitude" pageId="15" pageNumber="162" precision="925" value="-55.116665">55°07'S</geoCoordinate>
,
<geoCoordinate id="FCFD5024FFA7FFEFFC477DB0FBC01AD1" box="[1008,1086,1422,1442]" degrees="25" direction="west" minutes="59" orientation="longitude" pageId="15" pageNumber="162" precision="925" value="-25.983334">25°59'W</geoCoordinate>
,
<quantity id="5E319B06FFA7FFEFFBFD7DB0FB371AD2" box="[1098,1225,1422,1442]" metricMagnitude="3" metricUnit="m" metricValue="5.444" metricValueMax="5.452999999999999" metricValueMin="5.435" pageId="15" pageNumber="162" unit="m" value="5444.0" valueMax="5453.0" valueMin="5435.0">
<elevation id="12E4D1D0FFA7FFEFFBFD7DB0FB371AD2" box="[1098,1225,1422,1442]" metricMagnitude="3" metricUnit="m" metricValue="5.444" metricValueMax="5.452999999999999" metricValueMin="5.435" pageId="15" pageNumber="162" unit="m" value="5444.0" valueMax="5453.0" valueMin="5435.0">5435–5453 m</elevation>
</quantity>
,
<collectorName id="343C5335FFA7FFEFFB627DB0FAC21AD2" box="[1237,1340,1422,1441]" pageId="15" pageNumber="162">
RV
<emphasis id="ABBDEAF1FFA7FFEFFB4D7DB0FAC21AD2" box="[1274,1340,1422,1441]" italics="true" pageId="15" pageNumber="162">Eltanin</emphasis>
</collectorName>
,
<collectorName id="343C5335FFA7FFEFFAFE7DB0FAA11AD1" box="[1353,1375,1422,1442]" pageId="15" pageNumber="162">Cr</collectorName>
8 stn 591,
<date id="ED771023FFA7FFEFFCFC7D96FC401ACF" box="[843,958,1448,1468]" pageId="15" pageNumber="162" value="1963-04-29">
<collectingDate id="FD33E9CBFFA7FFEFFCFC7D96FC401ACF" box="[843,958,1448,1468]" pageId="15" pageNumber="162" value="1963-04-29">29 Apr 1963</collectingDate>
</date>
,
<specimenCode id="C96F9E98FFA7FFEFFC7F7D96FB9E1ACF" box="[968,1120,1448,1468]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="162" type="Museum">USNM 1008318</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="997636E3FFA7FFEFFC8F7DFDFAE31AA5" blockId="15.[798,1429,1395,1547]" box="[824,1309,1474,1494]" pageId="15" pageNumber="162">
<materialsCitation id="29A13CBEFFA7FFEFFC8F7DFDFB091AA5" box="[824,1271,1474,1494]" collectionCode="NMV" pageId="15" pageNumber="162" specimenCode="F104796" specimenCount="1" typeStatus="paratype">
<typeStatus id="46728841FFA7FFEFFC8F7DFDFC6C1AA5" box="[824,914,1475,1494]" pageId="15" pageNumber="162" type="paratype">Paratypes</typeStatus>
. Type locality and date,
<collectionCode id="FFD8AE26FFA7FFEFFBD97DFDFB5C1AA5" box="[1134,1186,1475,1494]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="15" pageNumber="162" type="Museum">NMV</collectionCode>
<specimenCode id="C96F9E98FFA7FFEFFB107DFCFB091AA5" box="[1191,1271,1474,1494]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="15" pageNumber="162" type="Museum">F104796</specimenCode>
</materialsCitation>
(1).
</paragraph>
<paragraph id="997636E3FFA7FFEFFC8F7DE3FC771978" blockId="15.[798,1429,1395,1547]" pageId="15" pageNumber="162">
Other material.
<materialsCitation id="29A13CBEFFA7FFEFFC7F7DE3FC9A1978" collectionCode="USNM" elevation="2800" location="Scotia Sea" pageId="15" pageNumber="162" specimenCode="E48635" specimenCount="1" stateProvince="Orkney Islands">
<location id="9C166038FFA7FFEFFC7F7DE3FBD71A82" LSID="urn:lsid:plazi:treatment:116087F5FFA7FFF0FCA97D21FF0D1D03:9C166038FFA7FFEFFC7F7DE3FBD71A82" box="[968,1065,1501,1521]" name="Scotia Sea" pageId="15" pageNumber="162" stateProvince="Orkney Islands">Scotia Sea</location>
,
<locationDeviation id="88C42FC1FFA7FFEFFB837DE3FB661A82" box="[1076,1176,1501,1521]" location="South" pageId="15" pageNumber="162">N of South</locationDeviation>
<collectingRegion id="5B0DF801FFA7FFEFFB287DE3FB1C1A82" box="[1183,1250,1501,1521]" country="United Kingdom" name="Orkney Islands" pageId="15" pageNumber="162">Orkney</collectingRegion>
I.,
<quantity id="5E319B06FFA7FFEFFAB47DE3FAB71A83" box="[1283,1353,1501,1521]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="162" unit="m" value="2800.0">
<elevation id="12E4D1D0FFA7FFEFFAB47DE3FAB71A83" box="[1283,1353,1501,1521]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="162" unit="m" value="2800.0">2800 m</elevation>
</quantity>
,
<collectionCode id="FFD8AE26FFA7FFEFFAE37DE3FA6B1A82" box="[1364,1429,1501,1521]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="162" type="Museum">USNM</collectionCode>
<specimenCode id="C96F9E98FFA7FFEFFCA97DC9FC9A1978" box="[798,868,1527,1547]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="162" type="Museum">E48635</specimenCode>
</materialsCitation>
(1).
</paragraph>
<paragraph id="997636E3FFA7FFEFFCA97E1FFAD818DD" blockId="15.[798,1429,1568,1966]" pageId="15" pageNumber="162">
<emphasis id="ABBDEAF1FFA7FFEFFCA97E1FFC6A1944" box="[798,916,1569,1591]" italics="true" pageId="15" pageNumber="162">Description</emphasis>
. Up to
<quantity id="5E319B06FFA7FFEFFC4F7E1EFBB81945" box="[1016,1094,1568,1590]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="162" unit="mm" value="60.0">60 mm</quantity>
long; body small, rounded in section, elongate, tapered anteriorly and posteriorly, lacking brim; body wall firm, semi-gelatinous, surface smooth, not covered with attachments; tube feet all over body, overlapping ventrolaterally, distinctly cylindrical, frequently erect, close cover, anteriorly and pygally, typically 1.0 mm long,
<quantity id="5E319B06FFA7FFEFFA8C7E8FFA6B19B4" box="[1339,1429,1713,1735]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="162" unit="mm" value="0.25">0.25 mm</quantity>
diameter, up to
<quantity id="5E319B06FFA7FFEFFC087EF0FBF31997" box="[959,1037,1742,1764]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="162" unit="mm" value="1.5">1.5 mm</quantity>
long; tube feet typically shorter, more thread-like, dorsally and ventrally than ventrolaterally; longitudinal muscles broadly attached, relatively narrow, slightly thickened; some unbranched gonad tubules; ossicles in tentacles only, predominantly irregular, thick, rough to smooth, sometimes bluntly spinous rods, swollen mid-rod, tapering at ends, rare branches joined to create thick intertwining mesh, sometimes irregularly tuberous, up to 280 Μm long.
</paragraph>
<paragraph id="997636E3FFB8FFF0FF3D7915FD8D1E32" blockId="16.[138,627,299,321]" box="[138,627,299,321]" pageId="16" pageNumber="163">
<emphasis id="ABBDEAF1FFB8FFF0FF3D7915FF2F1E32" box="[138,209,299,321]" italics="true" pageId="16" pageNumber="163">Colour</emphasis>
. Body, tube feet off-white to pale brown.
</paragraph>
<paragraph id="997636E3FFB8FFF0FF3D7968FD6F1EFA" blockId="16.[138,769,342,393]" pageId="16" pageNumber="163">
<emphasis id="ABBDEAF1FFB8FFF0FF3D7968FF0B1E1F" box="[138,245,342,364]" italics="true" pageId="16" pageNumber="163">Etymology</emphasis>
. From the Latin
<taxonomicName id="5EC94D60FFB8FFF0FE107968FDF91E1F" authorityName="O'Loughlin &amp; Ahearn" authorityYear="2005" box="[423,519,342,364]" class="Holothuroidea" family="Molpadiodemidae" genus="Molpadiodemas" kingdom="Animalia" order="Persiculida" pageId="16" pageNumber="163" phylum="Echinodermata" rank="species" species="pediculus">
<emphasis id="ABBDEAF1FFB8FFF0FE107968FDF91E1F" box="[423,519,342,364]" italics="true" pageId="16" pageNumber="163">pediculus</emphasis>
</taxonomicName>
(little foot), referring to the prominent ventrolateral band of small tube feet.
</paragraph>
<paragraph id="997636E3FFB8FFF0FF3D79A1FE301EA2" blockId="16.[138,769,414,465]" pageId="16" pageNumber="163">
<emphasis id="ABBDEAF1FFB8FFF0FF3D79A1FEFA1EC6" box="[138,260,415,437]" italics="true" pageId="16" pageNumber="163">Distribution</emphasis>
. South Atlantic Ocean, off South Sandwich and South
<collectingRegion id="5B0DF801FFB8FFF0FF7C7985FEE81EA2" box="[203,278,443,465]" country="United Kingdom" name="Orkney Islands" pageId="16" pageNumber="163">Orkney</collectingRegion>
Is,
<quantity id="5E319B06FFB8FFF0FE8B7985FE341EA2" box="[316,458,443,465]" metricMagnitude="3" metricUnit="m" metricValue="4.1265" metricValueMax="5.452999999999999" metricValueMin="2.8" pageId="16" pageNumber="163" unit="m" value="4126.5" valueMax="5453.0" valueMin="2800.0">2800–5453 m</quantity>
.
</paragraph>
<paragraph id="997636E3FFB8FFF0FF3D79D9FF0D1D03" blockId="16.[138,769,487,624]" pageId="16" pageNumber="163">
<emphasis id="ABBDEAF1FFB8FFF0FF3D79D9FF1F1E8E" box="[138,225,487,509]" italics="true" pageId="16" pageNumber="163">Remarks</emphasis>
. Amongst
<taxonomicName id="5EC94D60FFB8FFF0FEED79D9FDAF1E8E" box="[346,593,487,509]" pageId="16" pageNumber="163">
<emphasis id="ABBDEAF1FFB8FFF0FEED79D9FE051E8E" box="[346,507,487,509]" italics="true" pageId="16" pageNumber="163">Molpadiodemas</emphasis>
species
</taxonomicName>
, the distinguishing characters are: body tapering orally and anally; close cylindrical tube feet overlapping ventrolaterally; longitudinal muscles broadly attached, relatively narrow, slightly thickened.
</paragraph>
</subSubSection>
</treatment>
</document>
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