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<document id="A81CD6F833527F86FE13C2C12585853E" ID-CLB-Dataset="54099" ID-DOI="10.5852/ejt.2022.807.1717" ID-GBIF-Dataset="e5917c1e-b93d-4ffe-882e-8fcc8cdcbfc4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6419306" ID-ZooBank="67DAF0D7-9D9F-4F0F-8004-9CCE0071BF33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649272462660" checkinUser="felipe" docAuthor="Cruz-Gómez, Christopher" docDate="2022" docId="03CB87CFC652FFBEB82CFB64B38EFCF5" docLanguage="en" docName="ejt-807_1-59.pdf" docOrigin="European Journal of Taxonomy 807" docSource="http://dx.doi.org/10.5852/ejt.2022.807.1717" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hartmanipsammolyce pettiboneae Cruz-Gómez 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FFF2FFB7C65FFFAFBA6CFFD0B13EFFE3" masterDocTitle="Pelogeniinae Chamberlin, 1919 (Annelida, Sigalionidae) from the Grand Caribbean Region" masterLastPageNumber="59" masterPageNumber="1" pageNumber="14" updateTime="1699319485203" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="98D710C2894110DA497A73465701DCEB">Pelogeniinae Chamberlin, 1919 (Annelida, Sigalionidae) from the Grand Caribbean Region</mods:title>
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<mods:name id="E1C8EF8F2C93A688131695B4A963F81B" type="personal">
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<mods:namePart id="D267928103DA6ED4B4CFEC0DB3A4B7F0">Cruz-Gómez, Christopher</mods:namePart>
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<mods:affiliation id="7272990829248892131B9B786728E334">Departamento de Sistemática y Ecología Acuática, El Colegio de la Frontera Sur, Unidad Chetumal, Chetumal, Quintana Roo, 77014, Mexico.</mods:affiliation>
<mods:nameIdentifier id="D6DFFF7B6D7E3942DAA825588DB936F9" type="email">christopher.cruz-gomez@hotmail.com</mods:nameIdentifier>
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<mods:title id="9A4A960E973E4C7E681E9AAB8E1DF842">European Journal of Taxonomy</mods:title>
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<mods:date id="2566F7D4DE79FF07AA83DE960BDA889C">2022</mods:date>
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<mods:identifier id="596396173F7BBA08E5A227C6181A2128" type="DOI">10.5852/ejt.2022.807.1717</mods:identifier>
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<treatment id="03CB87CFC652FFBEB82CFB64B38EFCF5" ID-DOI="http://doi.org/10.5281/zenodo.6421751" ID-GBIF-Taxon="194474794" ID-Zenodo-Dep="6421751" LSID="urn:lsid:plazi:treatment:03CB87CFC652FFBEB82CFB64B38EFCF5" httpUri="http://treatment.plazi.org/id/03CB87CFC652FFBEB82CFB64B38EFCF5" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
<subSubSection id="C3786552C652FFA2B82CFB64B501FB2C" box="[576,1087,1204,1231]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BDD36D9C652FFA2B82CFB64B501FB2C" blockId="13.[395,1268,1204,1301]" box="[576,1087,1204,1231]" pageId="13" pageNumber="14">
<heading id="D09581B5C652FFA2B82CFB64B501FB2C" box="[576,1087,1204,1231]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<taxonomicName id="4C624D5AC652FFA2B82CFB64B2E0FB2C" ID-CoL="8ZF2V" authority="Cruz-Gómez, 2022" authorityName="Cruz-Gómez" authorityYear="2022" box="[576,990,1204,1231]" class="Polychaeta" family="Sigalionidae" genus="Hartmanipsammolyce" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
<emphasis id="B916EACBC652FFA2B82CFB64B2E0FB2C" bold="true" box="[576,990,1204,1231]" italics="true" pageId="13" pageNumber="14">Hartmanipsammolyce pettiboneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557B0C652FFA2B989FB65B501FB2C" box="[997,1087,1205,1231]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="C3786552C652FFA2BBE7FB07B2A7FAF6" pageId="13" pageNumber="14" type="description">
<paragraph id="8BDD36D9C652FFA2BBE7FB07B5CAFB11" blockId="13.[395,1268,1204,1301]" box="[395,1268,1239,1266]" pageId="13" pageNumber="14">
<uri id="FFF33ADBC652FFA2BBE7FB07B5CAFB11" box="[395,1268,1239,1266]" pageId="13" pageNumber="14">
urn:lsid:zoobank.org:act:
<uuid id="FFC40C0CC652FFA2B8C5FB07B5CAFB11" box="[681,1268,1239,1266]" pageId="13" pageNumber="14">B129D523-4BFB-45BB-AADD-70EF1B644E73</uuid>
</uri>
</paragraph>
<paragraph id="8BDD36D9C652FFA2B889FB2AB2A7FAF6" blockId="13.[395,1268,1204,1301]" box="[741,921,1274,1301]" pageId="13" pageNumber="14">
<figureCitation id="13592A5CC652FFA2B889FB2AB25FFAF6" box="[741,865,1274,1301]" captionStart="Fig" captionStartId="6.[227,270,1269,1295]" captionTargetBox="[351,1311,302,1227]" captionTargetId="figure-75@6.[351,1311,302,1227]" captionTargetPageId="6" captionText="Fig. 2. Elytral microstructures. A–B, E–F, M–N. Dayipsammolyce paulayi sp. nov. C–D, I–J. Pelogenia kinbergi (Hansen, 1882). G–H. Neopsammolyce floccifera (Augener, 1906). K–L. Neopsammolyce aff. floccifera. O–P. Hartmanipsammolyce pettiboneae sp. nov. Q–R. Pelogenia capitata sp. nov. A–H. Non-adherent microestructures. A–B. Microtubercles. C–D. Pedunculate papillae with puffed tips. E–F. Short dendritic papillae. G–H. Long dendritic papillae. I–R. Adherent papillae. I–J. Flat papillae. K–L. Pedunculate papillae with flat tip. M–N. Pedunculate papillae with truncated tips. O–P. Pedunculate globular papillae. Q–R. Pedunculate capitate papillae. Scale bars: A–B, N, Q–R = 10 μm; C–F, I–M, O = 20 μm; G–H = 100 μm; P = 5 μm." figureDoi="http://doi.org/10.5281/zenodo.6419310" httpUri="https://zenodo.org/record/6419310/files/figure.png" pageId="13" pageNumber="14">Figs 2O–P</figureCitation>
,
<figureCitation id="13592A5CC652FFA2B902FB2AB2A7FAF6" box="[878,921,1274,1301]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[227,270,1753,1779]" captionStartId-1="16.[227,270,1859,1885]" captionTargetBox-0="[349,1314,302,1713]" captionTargetBox-1="[314,1350,302,1825]" captionTargetId-0="figure-174@15.[349,1314,302,1714]" captionTargetId-1="figure-120@16.[314,1350,302,1825]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." captionText-1="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi-0="http://doi.org/10.5281/zenodo.6419316" figureDoi-1="http://doi.org/10.5281/zenodo.6419318" httpUri-0="https://zenodo.org/record/6419316/files/figure.png" httpUri-1="https://zenodo.org/record/6419318/files/figure.png" pageId="13" pageNumber="14">5–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3786552C652FFA2BA8FFAEBB579FA29" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BDD36D9C652FFA2BA8FFAEBB062FAB6" blockId="13.[227,348,1339,1365]" box="[227,348,1339,1365]" pageId="13" pageNumber="14">
<heading id="D09581B5C652FFA2BA8FFAEBB062FAB6" bold="true" box="[227,348,1339,1365]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B916EACBC652FFA2BA8FFAEBB062FAB6" bold="true" box="[227,348,1339,1365]" pageId="13" pageNumber="14">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD36D9C652FFA2BA8FFABAB579FA29" blockId="13.[226,1436,1386,1482]" pageId="13" pageNumber="14">
A species of
<taxonomicName id="4C624D5AC652FFA2BB15FABAB342FA67" authorityName="Pettibone" authorityYear="1997" box="[377,636,1386,1412]" class="Polychaeta" family="Sigalionidae" genus="Hartmanipsammolyce" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
<emphasis id="B916EACBC652FFA2BB15FABAB342FA67" box="[377,636,1386,1412]" italics="true" pageId="13" pageNumber="14">Hartmanipsammolyce</emphasis>
</taxonomicName>
with an oblong neuropodial lobe on segment III; neurochaetae from median segments with blades with long distal and subdistal teeth; posterior elytra with a medial process and 6–7 medial projections; marginal elytral papillae long and articulated.
</paragraph>
</subSubSection>
<subSubSection id="C3786552C652FFA2BA8FFA22B085F940" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BDD36D9C652FFA2BA8FFA22B054F9EF" blockId="13.[227,362,1522,1548]" box="[227,362,1522,1548]" pageId="13" pageNumber="14">
<heading id="D09581B5C652FFA2BA8FFA22B054F9EF" bold="true" box="[227,362,1522,1548]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B916EACBC652FFA2BA8FFA22B054F9EF" bold="true" box="[227,362,1522,1548]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD36D9C652FFA2BA8FF9F0B085F940" blockId="13.[226,1436,1568,1699]" pageId="13" pageNumber="14">
The species is named after the late Dr Marian H. Pettibone in recognition of her sustained efforts in studying polychaetes during many years, especially regarding the members of
<taxonomicName id="4C624D5AC652FFA2BE1BF993B430F9BE" authorityName="Kinberg" authorityYear="1856" box="[1143,1294,1603,1629]" class="Polychaeta" family="Sigalionidae" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="14" phylum="Annelida" rank="family">Sigalionidae</taxonomicName>
. Her studies are the backbone for this and further contributions. The epithet is a noun in the genitive case (
<bibRefCitation id="EFF34B28C652FFA2BF39F9B6B01EF940" author="ICZN" pageId="13" pageNumber="14" refId="ref29913" refString="ICZN. 1999. International Code of Zoological Nomenclature, 4 th Ed. International Trust for Zoological Nomenclature. The Natural History Museum, London." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 31.1.2).
</paragraph>
</subSubSection>
<subSubSection id="C3786552C652FFA2BA8FF91AB3C4F8BA" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BDD36D9C652FFA2BA8FF91AB0EEF907" blockId="13.[227,464,1738,1764]" box="[227,464,1738,1764]" pageId="13" pageNumber="14">
<heading id="D09581B5C652FFA2BA8FF91AB0EEF907" bold="true" box="[227,464,1738,1764]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B916EACBC652FFA2BA8FF91AB0EEF907" bold="true" box="[227,464,1738,1764]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD36D9C652FFA2BB7CF929B3C4F8BA" blockId="13.[227,1437,1785,1881]" pageId="13" pageNumber="14">
<materialsCitation id="3B0A3C84C652FFA2BB7CF929B3C4F8BA" ID-GBIF-Occurrence="3748376301" collectingDate="1967-05-22" collectionCode="R, V" collectorName="R / V Pillsbury" country="Mexico" county="Cancun" elevation="351" latitude="21.083334" location="Quintana Roo" longLatPrecision="1265" longitude="-86.38333" municipality="Pillsbury" pageId="13" pageNumber="14" specimenCode="UMML 22.1087" specimenCount="1" stateProvince="Quintana Roo" typeStatus="holotype">
<emphasis id="B916EACBC652FFA2BB7CF929B0BCF8F0" bold="true" box="[272,386,1785,1811]" pageId="13" pageNumber="14">
<typeStatus id="54D9887BC652FFA2BB7CF929B0BCF8F0" box="[272,386,1785,1811]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3757649C652FFA2BA8FF8CCB069F8D5" box="[227,343,1820,1846]" name="Mexico" pageId="13" pageNumber="14">MEXICO</collectingCountry>
•
<specimenCount id="9D64FD50C652FFA2BB1DF8CBB309F8D5" box="[369,567,1819,1846]" pageId="13" pageNumber="14">incomplete spec</specimenCount>
.;
<collectingRegion id="49A6F83BC652FFA2B82BF8CBB3D1F8D5" box="[583,751,1819,1846]" country="Mexico" name="Quintana Roo" pageId="13" pageNumber="14">Quintana Roo</collectingRegion>
,
<locationDeviation id="9A6F2FFBC652FFA2B890F8CBB2BCF8D5" box="[764,898,1819,1846]" pageId="13" pageNumber="14">off Cancun</locationDeviation>
;
<geoCoordinate id="EE56501EC652FFA2B9E3F8CAB2C2F8D5" box="[911,1020,1818,1846]" degrees="21" direction="north" minutes="05" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="21.083334">21°05′ N</geoCoordinate>
,
<geoCoordinate id="EE56501EC652FFA2BE65F8CAB542F8D5" box="[1033,1148,1818,1846]" degrees="86" direction="west" minutes="23" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="-86.38333">86°23′ W</geoCoordinate>
;
<collectorName id="2697530FC652FFA2BEE5F8CCB412F8D6" box="[1161,1324,1819,1846]" pageId="13" pageNumber="14">R/V Pillsbury</collectorName>
, Stn 580; depth
<quantity id="4C9A9B3CC652FFA2BB47F8EEB047F8BA" box="[299,377,1854,1881]" metricMagnitude="2" metricUnit="m" metricValue="3.51" pageId="13" pageNumber="14" unit="m" value="351.0">
<elevation id="004FD1EAC652FFA2BB47F8EEB047F8BA" box="[299,377,1854,1881]" metricMagnitude="2" metricUnit="m" metricValue="3.51" pageId="13" pageNumber="14" unit="m" value="351.0">351 m</elevation>
</quantity>
;
<date id="FFDC1019C652FFA2BBE9F8EFB31BF8BB" box="[389,549,1854,1881]" pageId="13" pageNumber="14" value="1967-05-22">
<collectingDate id="EF98E9F1C652FFA2BBE9F8EFB31BF8BB" box="[389,549,1854,1881]" pageId="13" pageNumber="14" value="1967-05-22">22 May 1967</collectingDate>
</date>
;
<specimenCode id="DBC49EA2C652FFA2B85DF8EFB3CBF8BA" box="[561,757,1855,1881]" collectionCode="UMML" country="USA" httpUri="http://grbio.org/cool/qx5p-wgw2" name="University of Miami Marine Laboratory" pageId="13" pageNumber="14">UMML 22.1087</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3786552C652FFA1BA8FF8AFB0C9F7EC" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8BDD36D9C652FFA2BA8FF8AFB0CBF878" blockId="13.[227,501,1919,1947]" box="[227,501,1919,1947]" pageId="13" pageNumber="14">
<emphasis id="B916EACBC652FFA2BA8FF8AFB04AF87A" bold="true" box="[227,372,1919,1945]" pageId="13" pageNumber="14">Description</emphasis>
(
<typeStatus id="54D9887BC652FFA2BBEFF850B0D3F878" box="[387,493,1920,1947]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BDD36D9C652FFA2BA8FF87FB23DF7EC" blockId="13.[226,1438,1966,2063]" pageId="13" pageNumber="14">
BODY. Orange, long, broad (
<figureCitation id="13592A5CC652FFA2B859F87EB3ABF82A" box="[565,661,1966,1993]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
); 136 segments,
<quantity id="4C9A9B3CC652FFA2B90DF87EB2A3F82A" box="[865,925,1966,1993]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="cm" value="9.0">9 cm</quantity>
long,
<quantity id="4C9A9B3CC652FFA2B987F87EB503F82A" box="[1003,1085,1966,1993]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="14" unit="cm" value="1.1">1.1 cm</quantity>
to segment 30,
<quantity id="4C9A9B3CC652FFA2BF6CF87EB46DF82A" box="[1280,1363,1966,1993]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="14" unit="cm" value="1.1">1.1 cm</quantity>
wide. Middorsal line covered with white foreign particles attached to adhesive papillae (
<figureCitation id="13592A5CC652FFA2BEC9F801B43DF80F" box="[1189,1283,2001,2028]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
). Venter only covered with short globular papillae (
<figureCitation id="13592A5CC652FFA2B8F6F824B3CBF7EC" box="[666,757,2036,2063]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="8BDD36D9C651FFA1BA8FFEFFB4A6FDC5" blockId="14.[226,1438,303,550]" pageId="14" pageNumber="15">
PROSTOMIUM. Oval, wider than long. Two pairs of eyes, anterior eyes larger. Lateral antennae small, slender; ceratophores as long as styles, dorsally fused with tentacular segment. Median antennal ceratophore bulbous, as long as prostomium, with a lateral ridge; basally with ctenidia; style long, as long as ceratophore. Middorsal lobe of segment II subtriangular (
<figureCitation id="13592A5CC651FFA1B9C9FE4DB53CFE5B" box="[933,1026,413,440]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5B</figureCitation>
). First segment directed anteriorly, fused with tentacular segment; biramous, chaetae simple verticillate. Dorsal tentacular cirrus slightly longer than neuropodia including chaetae, ventral tentacular cirrus slightly shorter than dorsal one; palps laterally displaced, short, reaching segment five, with inner sheaths. Pharynx partially everted (
<figureCitation id="13592A5CC651FFA1BF43FDDBB4B4FDC5" box="[1327,1418,523,550]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8BDD36D9C651FFA1BA8FFD9FB341FC6C" blockId="14.[225,1438,590,911]" pageId="14" pageNumber="15">
ELYTRA. First right elytron oval, with a mid-distal notch, covered with few fragments of shells attached to several kinds of papillae (
<figureCitation id="13592A5CC651FFA1B844FDA3B3BCFD6D" box="[552,642,627,654]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">Fig. 6E</figureCitation>
); elytral surface with pedunculate papillae with flat tips, pedunculate papillae with truncated tips, and pedunculate globular papillae, more abundant in the mid-posterior section; elytral margins with short dendritic papillae and segmented, long dendritic papillae (
<figureCitation id="13592A5CC651FFA1BF77FD6DB4A9FD34" box="[1307,1431,701,727]" captionStart="Fig" captionStartId="6.[227,270,1269,1295]" captionTargetBox="[351,1311,302,1227]" captionTargetId="figure-75@6.[351,1311,302,1227]" captionTargetPageId="6" captionText="Fig. 2. Elytral microstructures. A–B, E–F, M–N. Dayipsammolyce paulayi sp. nov. C–D, I–J. Pelogenia kinbergi (Hansen, 1882). G–H. Neopsammolyce floccifera (Augener, 1906). K–L. Neopsammolyce aff. floccifera. O–P. Hartmanipsammolyce pettiboneae sp. nov. Q–R. Pelogenia capitata sp. nov. A–H. Non-adherent microestructures. A–B. Microtubercles. C–D. Pedunculate papillae with puffed tips. E–F. Short dendritic papillae. G–H. Long dendritic papillae. I–R. Adherent papillae. I–J. Flat papillae. K–L. Pedunculate papillae with flat tip. M–N. Pedunculate papillae with truncated tips. O–P. Pedunculate globular papillae. Q–R. Pedunculate capitate papillae. Scale bars: A–B, N, Q–R = 10 μm; C–F, I–M, O = 20 μm; G–H = 100 μm; P = 5 μm." figureDoi="http://doi.org/10.5281/zenodo.6419310" httpUri="https://zenodo.org/record/6419310/files/figure.png" pageId="14" pageNumber="15">Figs 2O–P</figureCitation>
,
<figureCitation id="13592A5CC651FFA1BA8EFD31B01FFD1F" box="[226,289,737,764]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">6I, L</figureCitation>
). Second right elytron oblong, covered with few fragments of shells and same kinds of papillae (
<figureCitation id="13592A5CC651FFA1BA81FCD6B07AFCC2" box="[237,324,774,801]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">Fig. 6F</figureCitation>
) but sparser, lateral margin with one mid-distal projection (
<figureCitation id="13592A5CC651FFA1B997FCD6B566FCC2" box="[1019,1112,774,801]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">Fig. 6H</figureCitation>
). Third right elytron similar to second one. Posterior elytra subrectangular with papillae only along posterolateral elytral surface (
<figureCitation id="13592A5CC651FFA1BA87FC80B050FC89" box="[235,366,848,874]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">Fig. 6G, M</figureCitation>
), with a large medial process, lateral margin with 6–7 mid-distal projections with segmented, long dendritic papillae (
<figureCitation id="13592A5CC651FFA1BB95FCA4B34CFC6C" box="[505,626,884,911]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">Fig. 6J–K</figureCitation>
).
</paragraph>
<paragraph id="8BDD36D9C651FFA1BA8FFC68B0D6FB37" blockId="14.[226,1437,951,1236]" pageId="14" pageNumber="15">
RIGHT PARAPODIUM FROM SEGMENT II (
<figureCitation id="13592A5CC651FFA1B8CDFC67B3C1FC31" box="[673,767,951,978]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5F</figureCitation>
). Notopodia truncated, papillate, short, half as long as neuropodia. With up to 50 simple verticillate notochaetae, shortest ones as long as notopodia, longest ones 4× as long (
<figureCitation id="13592A5CC651FFA1BBD0FBD1B320FBFF" box="[444,542,1025,1052]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5G</figureCitation>
). Neuropodia heart-shaped, smooth (non-papillate), larger than notopodia. Neurochaetae only bifid falcigers: unit A, eight falcigers directed upwards with handles slender, with 30–36 transverse rows of spines, blades long, 16 × as long as wide (
<figureCitation id="13592A5CC651FFA1BE6BFB9AB558FB86" box="[1031,1126,1098,1125]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5H</figureCitation>
); unit B, 15 falcigers with handles thick, with 27–34 transverse rows of spines, blades long, 35× as long as wide (
<figureCitation id="13592A5CC651FFA1BE8EFBBFB40AFB69" box="[1250,1332,1135,1162]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5I</figureCitation>
); unit D, seven falcigers with handles slender with 15–18 transverse rows of spines, blades medium-sized, 10 × as long as wide (
<figureCitation id="13592A5CC651FFA1BBEAFB69B0E4FB37" box="[390,474,1209,1236]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5J</figureCitation>
).
</paragraph>
<paragraph id="8BDD36D9C651FFA1BA8FFB2CB595F9F4" blockId="14.[225,1438,1276,1560]" pageId="14" pageNumber="15">
RIGHT PARAPODIUM FROM SEGMENT III (
<figureCitation id="13592A5CC651FFA1B8FBFB2CB3CBFAF4" box="[663,757,1276,1303]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5K</figureCitation>
). Notopodia heart-shaped, papillate, short, half as long as neuropodia. With up to 80 simple verticillate notochaetae, shortest ones as long as notopodia, longest ones 4× as long (
<figureCitation id="13592A5CC651FFA1BBDAFA95B32CFA83" box="[438,530,1349,1376]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5L</figureCitation>
). Neuropodia truncated, smooth (non-papillate), larger than notopodia, with one large oblong distal lobe (twice as long as neuropodia). Neurochaetae only falcigers: unit A, four falcigers with handles thick with 14–15 transverse rows of spines, blades long, 14–16× as long as wide (
<figureCitation id="13592A5CC651FFA1BA87FA64B076FA2D" box="[235,328,1459,1486]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5O</figureCitation>
); unit B, 12 falcigers with handles slender, smooth, blades long, 12 × as long as wide (
<figureCitation id="13592A5CC651FFA1BF40FA64B4B0FA2D" box="[1324,1422,1459,1486]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5M</figureCitation>
); unit C and D undifferentiated, 13 falcigers with handles slender, smooth, blades medium-sized, 8–10 × as long as wide (
<figureCitation id="13592A5CC651FFA1BBE9FA2DB0DAF9FB" box="[389,484,1533,1560]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5N</figureCitation>
). Unit A with bifid blades, units B and C with falcate blades.
</paragraph>
<paragraph id="8BDD36D9C651FFA1BA8FF991B027F828" blockId="14.[227,1437,1600,1995]" pageId="14" pageNumber="15">
RIGHT PARAPODIUM FROM SEGMENT 81 (MIDDLE SEGMENT) (
<figureCitation id="13592A5CC651FFA1B917F990B2E8F9B8" box="[891,982,1600,1627]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Figs 5P</figureCitation>
,
<figureCitation id="13592A5CC651FFA1B989F990B537F9B9" box="[997,1033,1600,1626]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">6A</figureCitation>
). Notopodia, truncated, papillate, short, as long as half of neuropodia. With up to 120 simple verticillate notochaetae, shortest ones half as long as notopodia, longest ones 4× as long (
<figureCitation id="13592A5CC651FFA1B97CF95AB24CF946" box="[784,882,1674,1701]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Figs 5Q</figureCitation>
,
<figureCitation id="13592A5CC651FFA1B913F95AB29AF946" box="[895,932,1674,1701]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">6B</figureCitation>
). Neuropodia leaf-shaped, papillate, larger than notopodia. Neurochaetae only bifid falcigers: unit A, nine falcigers with handles thick, with 5–10 transverse rows of spines, blades medium-sized, 8–10× as long as wide (
<figureCitation id="13592A5CC651FFA1BE0CF903B5FAF90D" box="[1120,1220,1747,1774]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Figs 5R</figureCitation>
,
<figureCitation id="13592A5CC651FFA1BEBFF903B5C7F90D" box="[1235,1273,1747,1774]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">6C</figureCitation>
); unit B, two falcigers with handle thick with 5–6 rows of spines, blades lost (in posterior segments, blades mediumsized, 5–6× as long as wide) (
<figureCitation id="13592A5CC651FFA1B82FF8CDB3A1F8DB" box="[579,671,1821,1848]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Figs 5S</figureCitation>
,
<figureCitation id="13592A5CC651FFA1B8C0F8CDB3ECF8DB" box="[684,722,1821,1848]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">6C</figureCitation>
); unit C, 13 falcigers with handles thick, with 3–4 transverse rows of spines, blades medium-sized, 5–6 × as long as wide (
<figureCitation id="13592A5CC651FFA1B9A4F892B51BF8BF" box="[968,1061,1857,1884]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5T</figureCitation>
); subunit 2, four falcigers with handles thick, smooth, blades long, 20× as long as wide (
<figureCitation id="13592A5CC651FFA1B9E5F8B6B2D6F862" box="[905,1000,1894,1921]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Fig. 5U</figureCitation>
); unit D, eight falcigers with handles slender, with two barely noticeable rows of denticles, blades long, 15–20 × as long as wide (
<figureCitation id="13592A5CC651FFA1BF54F85BB4A6F845" box="[1336,1432,1931,1958]" captionStart="Fig" captionStartId="15.[227,270,1753,1779]" captionTargetBox="[349,1314,302,1713]" captionTargetId="figure-174@15.[349,1314,302,1714]" captionTargetPageId="15" captionText="Fig. 5. Hartmanipsamolyce pettiboneae sp. nov., holotype (UMML 22.1087). A. Incomplete worm, dorsal view. B. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx. C. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx. D. Dorsal papillae (close-up in box). E. Ventral papillae (close-up in box). F. Right parapodium from segment II. G. Notochaeta from same. H. Unit A. I. Unit B. J. Units C and D. K. Right parapodium from segment III, arrow indicates neuropodial lobe. L. Notochaeta from same. M. Unit B. N. Units C and D. O. Unit A. P. Right parapodium from segment 81. Q. Notochaeta from same. R. Unit A. S. Unit B. T. Unit C. U. Subunit 2. V. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O, R–V = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="14" pageNumber="15">Figs 5V</figureCitation>
,
<figureCitation id="13592A5CC651FFA1BA88F860B032F828" box="[228,268,1968,1995]" captionStart="Fig" captionStartId="16.[227,270,1859,1885]" captionTargetBox="[314,1350,302,1825]" captionTargetId="figure-120@16.[314,1350,302,1825]" captionTargetPageId="16" captionText="Fig. 6. Hartmanipsammolyce pettiboneae sp. nov., holotype (UMML 22.10887). A–D, K–M. SEM micrographs. A. Right parapodium from segment 30.B. Notochaeta from same.C. UnitsA and B.D. Unit D. E. First right elytron. F. Second right elytron. G. Posterior right elytron. H. Proximal margin of E. I. Posterior margin from E. J. Posterior processes from G. K. Segmented long papillae. L. Pedunculate papilla with flat tip. M. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G = 1 mm; L = 10 μm; M = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="14" pageNumber="15">6D</figureCitation>
).
</paragraph>
<paragraph id="8BDD36D9C651FFA1BA8FF825B0C9F7EC" blockId="14.[227,503,2037,2063]" box="[227,503,2037,2063]" pageId="14" pageNumber="15">POSTERIOR REGION. Lost.</paragraph>
</subSubSection>
<caption id="DF1D6651C650FFA0BA8FF909B38AF7ED" ID-DOI="http://doi.org/10.5281/zenodo.6419316" ID-Zenodo-Dep="6419316" httpUri="https://zenodo.org/record/6419316/files/figure.png" pageId="15" pageNumber="16" startId="15.[227,270,1753,1779]" targetBox="[349,1314,302,1713]" targetPageId="15">
<paragraph id="8BDD36D9C650FFA0BA8FF909B38AF7ED" blockId="15.[225,1437,1753,2062]" pageId="15" pageNumber="16">
<emphasis id="B916EACBC650FFA0BA8FF909B00AF917" bold="true" box="[227,308,1753,1780]" pageId="15" pageNumber="16">Fig. 5.</emphasis>
<taxonomicName id="4C624D5AC650FFA0BB53F90AB3FFF917" authority="Cruz-Gómez, 2022" authorityName="Cruz-Gómez" authorityYear="2022" box="[319,705,1754,1780]" class="Polychaeta" family="Sigalionidae" genus="Hartmanipsamolyce" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="16" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
<emphasis id="B916EACBC650FFA0BB53F90AB3FFF917" box="[319,705,1754,1780]" italics="true" pageId="15" pageNumber="16">Hartmanipsamolyce pettiboneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557B0C650FFA0B8A0F90AB213F917" box="[716,813,1754,1780]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (UMML 22.1087).
<emphasis id="B916EACBC650FFA0BEF6F909B591F910" bold="true" box="[1178,1199,1753,1779]" pageId="15" pageNumber="16">A</emphasis>
. Incomplete worm, dorsal view.
<emphasis id="B916EACBC650FFA0BBEDF92DB0ABF8F4" bold="true" box="[385,405,1789,1815]" pageId="15" pageNumber="16">B</emphasis>
. Anterior end, dorsal view; arrowhead indicates ctenidia; asterisk indicates partially everted pharynx.
<emphasis id="B916EACBC650FFA0BBD9F8F0B0F4F8D9" bold="true" box="[437,458,1824,1850]" pageId="15" pageNumber="16">C</emphasis>
. Anterior end, ventral view (close-up in box); asterisk indicates partially everted pharynx.
<emphasis id="B916EACBC650FFA0BB3CF893B05BF8BE" bold="true" box="[336,357,1859,1885]" pageId="15" pageNumber="16">D</emphasis>
. Dorsal papillae (close-up in box).
<emphasis id="B916EACBC650FFA0B895F893B233F8BE" bold="true" box="[761,781,1859,1885]" pageId="15" pageNumber="16">E</emphasis>
. Ventral papillae (close-up in box).
<emphasis id="B916EACBC650FFA0BEC5F893B585F8BE" bold="true" box="[1193,1211,1859,1885]" pageId="15" pageNumber="16">F</emphasis>
. Right parapodium from segment II.
<emphasis id="B916EACBC650FFA0BBC5F8B6B0FEF863" bold="true" box="[425,448,1894,1920]" pageId="15" pageNumber="16">G</emphasis>
. Notochaeta from same.
<emphasis id="B916EACBC650FFA0B88EF8B7B3C7F862" bold="true" box="[738,761,1895,1921]" pageId="15" pageNumber="16">H</emphasis>
. Unit A.
<emphasis id="B916EACBC650FFA0B90CF8B7B255F862" bold="true" box="[864,875,1895,1921]" pageId="15" pageNumber="16">I</emphasis>
. Unit B.
<emphasis id="B916EACBC650FFA0B9B9F8B7B2DAF862" bold="true" box="[981,996,1895,1921]" pageId="15" pageNumber="16">J</emphasis>
. Units C and D.
<emphasis id="B916EACBC650FFA0BECFF8B7B584F862" bold="true" box="[1187,1210,1895,1921]" pageId="15" pageNumber="16">K</emphasis>
. Right parapodium from segment III, arrow indicates neuropodial lobe.
<emphasis id="B916EACBC650FFA0B93EF85AB25AF847" bold="true" box="[850,868,1930,1956]" pageId="15" pageNumber="16">L</emphasis>
. Notochaeta from same.
<emphasis id="B916EACBC650FFA0BEFDF85AB595F847" bold="true" box="[1169,1195,1930,1956]" pageId="15" pageNumber="16">M</emphasis>
. Unit B.
<emphasis id="B916EACBC650FFA0BF71F85AB40FF847" bold="true" box="[1309,1329,1930,1956]" pageId="15" pageNumber="16">N</emphasis>
. Units C and D.
<emphasis id="B916EACBC650FFA0BB54F87DB071F824" bold="true" box="[312,335,1965,1991]" pageId="15" pageNumber="16">O</emphasis>
. Unit A.
<emphasis id="B916EACBC650FFA0BBD5F87DB0F5F824" bold="true" box="[441,459,1965,1991]" pageId="15" pageNumber="16">P</emphasis>
. Right parapodium from segment 81.
<emphasis id="B916EACBC650FFA0B9E7F87DB29CF824" bold="true" box="[907,930,1965,1991]" pageId="15" pageNumber="16">Q</emphasis>
. Notochaeta from same.
<emphasis id="B916EACBC650FFA0BEA4F87DB5E3F824" bold="true" box="[1224,1245,1965,1991]" pageId="15" pageNumber="16">R</emphasis>
. Unit A.
<emphasis id="B916EACBC650FFA0BF2BF87DB469F824" bold="true" box="[1351,1367,1965,1991]" pageId="15" pageNumber="16">S</emphasis>
. Unit B.
<emphasis id="B916EACBC650FFA0BB69F800B027F809" bold="true" box="[261,281,2000,2026]" pageId="15" pageNumber="16">T</emphasis>
. Unit C.
<emphasis id="B916EACBC650FFA0BBEBF800B0A2F809" bold="true" box="[391,412,2000,2026]" pageId="15" pageNumber="16">U</emphasis>
. Subunit 2.
<emphasis id="B916EACBC650FFA0B841F800B37CF809" bold="true" box="[557,578,2000,2026]" pageId="15" pageNumber="16">V</emphasis>
. Unit D. Scale bars: A = 1 cm; B–C = 1 mm; D–F, K, P = 500 μm; G, L, Q = 40 μm; H–J, M–O,
<collectionCode id="ED73AE1CC650FFA0BB90F824B331F7ED" box="[508,527,2036,2062]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="16">R</collectionCode>
–
<collectionCode id="ED73AE1CC650FFA0B873F824B30DF7ED" box="[543,563,2036,2062]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="16" type="Museum">V</collectionCode>
= 100 μm.
</paragraph>
</caption>
<caption id="DF1D6651C64FFFBFBA8FF893B527F7ED" ID-DOI="http://doi.org/10.5281/zenodo.6419318" ID-Zenodo-Dep="6419318" httpUri="https://zenodo.org/record/6419318/files/figure.png" pageId="16" pageNumber="17" startId="16.[227,270,1859,1885]" subCaptionStartIDs="16.[663,834,2000,2026]" subCaptionStarts="Abbr" targetBox="[314,1350,302,1825]" targetPageId="16">
<paragraph id="8BDD36D9C64FFFBFBA8FF893B527F7ED" blockId="16.[224,1437,1859,2062]" pageId="16" pageNumber="17">
<emphasis id="B916EACBC64FFFBFBA8FF893B00BF8BE" bold="true" box="[227,309,1859,1885]" pageId="16" pageNumber="17">Fig. 6.</emphasis>
<taxonomicName id="4C624D5AC64FFFBFBB2DF893B3E7F8BE" authority="Cruz-Gómez, 2022" authorityName="Cruz-Gómez" authorityYear="2022" box="[321,729,1859,1885]" class="Polychaeta" family="Sigalionidae" genus="Hartmanipsammolyce" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
<emphasis id="B916EACBC64FFFBFBB2DF893B3E7F8BE" box="[321,729,1859,1885]" italics="true" pageId="16" pageNumber="17">Hartmanipsammolyce pettiboneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557B0C64FFFBFB888F894B278F8BD" box="[740,838,1860,1886]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (UMML 22.10887). A–D, K–M. SEM micrographs.
<emphasis id="B916EACBC64FFFBFBBEEF8B6B0A9F863" bold="true" box="[386,407,1894,1920]" pageId="16" pageNumber="17">A</emphasis>
. Right parapodium from segment 30.
<emphasis id="B916EACBC64FFFBFB92BF8B7B265F862" bold="true" box="[839,859,1895,1921]" pageId="16" pageNumber="17">B</emphasis>
. Notochaeta from same.
<emphasis id="B916EACBC64FFFBFBE14F8B6B5B3F863" bold="true" box="[1144,1165,1894,1920]" pageId="16" pageNumber="17">C</emphasis>
. Units A and B.
<emphasis id="B916EACBC64FFFBFBF28F8B7B467F862" bold="true" box="[1348,1369,1895,1921]" pageId="16" pageNumber="17">D</emphasis>
. Unit D.
<emphasis id="B916EACBC64FFFBFBB64F85AB022F847" bold="true" box="[264,284,1930,1956]" pageId="16" pageNumber="17">E</emphasis>
. First right elytron.
<emphasis id="B916EACBC64FFFBFB862F85AB321F847" bold="true" box="[526,543,1930,1956]" pageId="16" pageNumber="17">F</emphasis>
. Second right elytron.
<emphasis id="B916EACBC64FFFBFB95DF859B279F840" bold="true" box="[817,839,1929,1955]" pageId="16" pageNumber="17">G</emphasis>
. Posterior right elytron.
<emphasis id="B916EACBC64FFFBFBE01F85AB5BDF847" bold="true" box="[1133,1155,1930,1956]" pageId="16" pageNumber="17">H</emphasis>
. Proximal margin of E.
<emphasis id="B916EACBC64FFFBFBA88F87DB1D0F824" bold="true" box="[228,238,1965,1991]" pageId="16" pageNumber="17">I</emphasis>
. Posterior margin from E.
<emphasis id="B916EACBC64FFFBFB845F87DB306F824" bold="true" box="[553,568,1965,1991]" pageId="16" pageNumber="17">J</emphasis>
. Posterior processes from G.
<emphasis id="B916EACBC64FFFBFB9F8F87DB295F824" bold="true" box="[916,939,1965,1991]" pageId="16" pageNumber="17">K</emphasis>
. Segmented long papillae.
<emphasis id="B916EACBC64FFFBFBE87F87DB5C1F824" bold="true" box="[1259,1279,1965,1991]" pageId="16" pageNumber="17">L</emphasis>
. Pedunculate papilla with flat tip.
<emphasis id="B916EACBC64FFFBFBBA2F800B0D4F809" bold="true" box="[462,490,2000,2026]" pageId="16" pageNumber="17">M</emphasis>
. Surface of E. Abbreviations: a = anterior; p = posterior. Scale bars: A = 200 μm; B = 5 μm; C–D, H–K = 50 μm; E–G =
<quantity id="4C9A9B3CC64FFFBFB8DCF823B3C9F7ED" box="[688,759,2035,2062]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="1.0">1 mm</quantity>
; L = 10 μm; M = 20 μm.
</paragraph>
</caption>
<subSubSection id="C3786552C64EFFBEBA8FFEFEB3C1FD41" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BDD36D9C64EFFBEBA8FFEFEB06BFEAB" blockId="17.[227,341,302,328]" box="[227,341,302,328]" pageId="17" pageNumber="18">
<heading id="D09581B5C64EFFBEBA8FFEFEB06BFEAB" bold="true" box="[227,341,302,328]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B916EACBC64EFFBEBA8FFEFEB06BFEAB" bold="true" box="[227,341,302,328]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD36D9C64EFFBEBA8FFE8FB3C1FD41" blockId="17.[224,1439,351,675]" pageId="17" pageNumber="18">
<taxonomicName id="4C624D5AC64EFFBEBA8FFE8FB34DFE9A" authority="Cruz-Gómez, 2022" authorityName="Cruz-Gómez" authorityYear="2022" box="[227,627,351,377]" class="Polychaeta" family="Sigalionidae" genus="Hartmanipsammolyce" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
<emphasis id="B916EACBC64EFFBEBA8FFE8FB34DFE9A" box="[227,627,351,377]" italics="true" pageId="17" pageNumber="18">Hartmanipsammolyce pettiboneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557B0C64EFFBEB81BFE8FB3F1FE9A" box="[631,719,351,377]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C624D5AC64EFFBEB931FEB0B59FFE9A" authority="(Hartman, 1942)" baseAuthorityName="Hartman" baseAuthorityYear="1942" box="[861,1185,351,377]" class="Polychaeta" family="Sigalionidae" genus="Psammolyce" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="pendula">
<emphasis id="B916EACBC64EFFBEB931FEB0B2E3FE9A" box="[861,989,351,377]" italics="true" pageId="17" pageNumber="18">H. pendula</emphasis>
(Hartman, 1942)
</taxonomicName>
because in the former the large lobe on the margin of segment III is oblong; the middle segments have neurochaetal blades with long distal and subdistal teeth, even with tips touching each other; the first elytron is short, oval; and the posterior elytra have one large posterior process and 6–7 mid-distal projections with segmented, long dendritic papillae. By contrast, in
<taxonomicName id="4C624D5AC64EFFBEB8DEFE24B20BFDEE" baseAuthorityName="Hartman" baseAuthorityYear="1942" box="[690,821,499,525]" class="Polychaeta" family="Sigalionidae" genus="Psammolyce" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="pendula">
<emphasis id="B916EACBC64EFFBEB8DEFE24B20BFDEE" box="[690,821,499,525]" italics="true" pageId="17" pageNumber="18">H. pendula</emphasis>
</taxonomicName>
segment III has a balloon-like (
<bibRefCitation id="EFF34B28C64EFFBEBEC1FE24B4A6FDED" author="Hartman O." box="[1197,1432,499,526]" pageId="17" pageNumber="18" refId="ref29666" refString="Hartman O. 1965. Deep-water Benthic Polychaetous Annelids off New England to Bermuda and Other North Atlantic Areas. Allan Hancock Foundation Publications, Occasional Paper 28." type="journal volume" year="1965">Hartman 1965: 265</bibRefCitation>
, pl. 3 fig. a) or drop-like (
<bibRefCitation id="EFF34B28C64EFFBEB864FDC8B3D2FDD0" author="Pettibone M. H." box="[520,748,536,563]" pageId="17" pageNumber="18" pagination="1 - 89" refId="ref30430" refString="Pettibone M. H. 1997. Revision of the sigalionid species (Polychaeta) referred to Psammolyce Kinberg, 1856, Pelogenia Schmarda, 1861, and belonging to the subfamily Pelogeniinae Chamberlin, 1919. Smithsonian Contributions to Zoology 581: 1 - 89. https: // doi. org / 10.5479 / si. 00810282.581" type="journal article" year="1997">Pettibone 1997: 24</bibRefCitation>
, fig. 15e) lobe; middle segments have neurochaetal blades with short distal and subdistal teeth (
<bibRefCitation id="EFF34B28C64EFFBEB8FDFDEEB245FDBB" author="Hartman O." box="[657,891,573,600]" pageId="17" pageNumber="18" refId="ref29695" refString="Hartman O. 1968. Atlas of the Errantiate Polychaetous Annelids from California. Allan Hancock Foundation, University of Southern California, California." type="book" year="1968">Hartman 1968: 265</bibRefCitation>
, pl. 3 fig. b;
<bibRefCitation id="EFF34B28C64EFFBEBE63FDEEB5D1FDBB" author="Pettibone M. H." box="[1039,1263,573,600]" pageId="17" pageNumber="18" pagination="1 - 89" refId="ref30430" refString="Pettibone M. H. 1997. Revision of the sigalionid species (Polychaeta) referred to Psammolyce Kinberg, 1856, Pelogenia Schmarda, 1861, and belonging to the subfamily Pelogeniinae Chamberlin, 1919. Smithsonian Contributions to Zoology 581: 1 - 89. https: // doi. org / 10.5479 / si. 00810282.581" type="journal article" year="1997">Pettibone 1997: 25</bibRefCitation>
, fig. 16b); first elytra are enlarged and oblong; and posterior elytra have one large posterior process and no medial processes, with entire, long dendritic papillae.
</paragraph>
</subSubSection>
<subSubSection id="C3786552C64EFFBEBA8FFD1BB38EFCF5" box="[227,688,715,790]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BDD36D9C64EFFBEBA8FFD1BB042FD06" blockId="17.[227,380,715,741]" box="[227,380,715,741]" pageId="17" pageNumber="18">
<heading id="D09581B5C64EFFBEBA8FFD1BB042FD06" bold="true" box="[227,380,715,741]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B916EACBC64EFFBEBA8FFD1BB042FD06" bold="true" box="[227,380,715,741]" pageId="17" pageNumber="18">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD36D9C64EFFBEBA8FFD2CB38EFCF5" blockId="17.[227,688,764,790]" box="[227,688,764,790]" pageId="17" pageNumber="18">
Caribbean Sea.
<collectingRegion id="49A6F83BC64EFFBEBBF0FD2CB37DFCF5" box="[412,579,764,790]" country="Mexico" name="Quintana Roo" pageId="17" pageNumber="18">Quintana Roo</collectingRegion>
,
<collectingCountry id="F3757649C64EFFBEB822FD2CB395FCF5" box="[590,683,764,790]" name="Mexico" pageId="17" pageNumber="18">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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