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<document id="B9951BC06D4552D5C2AD7C5F4EC0AC16" ID-DOI="10.11646/zootaxa.5443.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11064278" ID-ZooBank="A91A8D67-A81B-427D-8096-59E41326A8E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714028852870" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2024" docId="03CB87B7E954FFFBF2CFFED7FAFAF94F" docLanguage="en" docName="zootaxa.5443.4.2.pdf" docOrigin="Zootaxa 5443 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Octavius lothenicus Janak 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="505" masterDocId="FFF2FFCFE95EFFF1F258FF9CFFC6FFD6" masterDocTitle="On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)" masterLastPageNumber="522" masterPageNumber="495" pageNumber="505" updateTime="1714029516812" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C16D26764ADA33395C9B06EF540EB040" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DDF4A38CA81852E2589E6FF9FA24CC88">
<mods:title id="BCEDB96B08E1C197139C58F45FFB0FFA">On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)</mods:title>
</mods:titleInfo>
<mods:name id="3E7FB6103FB75B3AFBE41E002457CE4E" type="personal">
<mods:role id="56C71652E63BB875DEB232EA2B916F6B">
<mods:roleTerm id="D12E870EE8E6061209CA06D8FFA1EDD0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7DBF007F4237A57C7CC7FF209CBA5DE">Janák, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="4BEC15606540FEE34BEC31A6BD61AB19">text</mods:typeOfResource>
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<mods:titleInfo id="8B82BA40FD18E84986E60ECDE0108F26">
<mods:title id="FA2C6BC1ECB6BF98C8791E051790A358">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CFED6C346757B19250A9333D578F282E">
<mods:date id="46895E7A55C925D029E738C7B159E945">2024</mods:date>
<mods:detail id="EF0CC281A861B1035F6EC99880B0E1D9" type="pubDate">
<mods:number id="2E0889975C3FB68877354390C79D2DB5">2024-04-24</mods:number>
</mods:detail>
<mods:detail id="DEBE0636B0FA17D1C250D63061039E6E" type="volume">
<mods:number id="2DE7737E4F5CE96F80D61394E3AA5F5B">5443</mods:number>
</mods:detail>
<mods:detail id="3760DDD1C63394C9B48AAACBB7F335F5" type="issue">
<mods:number id="74122B5AAE55E4BAE3C4FBE452753785">4</mods:number>
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<mods:extent id="8FAA606CFB593AB7FC5C274DBE17404C" unit="page">
<mods:start id="D01B00F67AA9913AFCA0135CA07C138E">495</mods:start>
<mods:end id="D2F79D8B46EBFF75068EA424ABD3CF2F">522</mods:end>
</mods:extent>
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<mods:location id="070D2FAF01EF29B007E00ADF015D48C0">
<mods:url id="F2ECE672FB8D68B72F9BA3986037FD37">http://dx.doi.org/10.11646/zootaxa.5443.4.2</mods:url>
</mods:location>
<mods:classification id="0B27697C08347A4C003350EA1C3E77AB">journal article</mods:classification>
<mods:identifier id="9587661CAE083B83EB7FA6E5C12276CC" type="DOI">10.11646/zootaxa.5443.4.2</mods:identifier>
<mods:identifier id="D87E6F057BD745660BFC7F7FF5F0A3F3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A08FD4B8D1464366BD29C429ADAF6207" type="Zenodo-Dep">11064278</mods:identifier>
<mods:identifier id="F77D21D31BED82638AF8515AF1DB158A" type="ZooBank">A91A8D67-A81B-427D-8096-59E41326A8E3</mods:identifier>
</mods:mods>
<treatment id="03CB87B7E954FFFBF2CFFED7FAFAF94F" ID-DOI="http://doi.org/10.5281/zenodo.11064363" ID-Zenodo-Dep="11064363" LSID="urn:lsid:plazi:treatment:03CB87B7E954FFFBF2CFFED7FAFAF94F" httpUri="http://treatment.plazi.org/id/03CB87B7E954FFFBF2CFFED7FAFAF94F" lastPageNumber="505" pageId="10" pageNumber="505">
<subSubSection id="C378652AE954FFFBF2CFFED7FD80FEB3" box="[151,582,330,357]" pageId="10" pageNumber="505" type="nomenclature">
<paragraph id="8BDD36A1E954FFFBF2CFFED7FD80FEB3" blockId="10.[151,582,330,393]" box="[151,582,330,357]" pageId="10" pageNumber="505">
<heading id="D09581CDE954FFFBF2CFFED7FD80FEB3" bold="true" box="[151,582,330,357]" fontSize="11" level="1" pageId="10" pageNumber="505" reason="1">
<emphasis id="B916EAB3E954FFFBF2CFFED7FD80FEB3" bold="true" box="[151,582,330,357]" pageId="10" pageNumber="505">
<taxonomicName id="4C624D22E954FFFBF2CFFED7FE1DFEB3" authority="Janak" authorityName="Janak" authorityYear="2024" box="[151,475,330,357]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="lothenicus" status="sp. nov.">
<emphasis id="B916EAB3E954FFFBF2CFFED7FE40FEB3" bold="true" box="[151,390,330,357]" italics="true" pageId="10" pageNumber="505">Octavius lothenicus</emphasis>
Janák
</taxonomicName>
,
<taxonomicNameLabel id="A22557C8E954FFFBF3B1FED7FD80FEB3" box="[489,582,331,357]" pageId="10" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378652AE954FFFBF2CFFEF3FE4DFE5F" box="[151,395,367,393]" pageId="10" pageNumber="505" type="description">
<paragraph id="8BDD36A1E954FFFBF2CFFEF3FE4DFE5F" blockId="10.[151,582,330,393]" box="[151,395,367,393]" pageId="10" pageNumber="505">
<figureCitation id="13592A24E954FFFBF2CFFEF3FED8FE5F" box="[151,286,367,393]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Figs. 44–48</figureCitation>
,
<figureCitation id="13592A24E954FFFBF371FEF3FE93FE5F" box="[297,341,367,393]" captionStart="FIGURES 113–121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113–121. Octavius undulatus Janák, sp. nov.; 113–116, 118–119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116–117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114–115; 116–117; 118–119; o: ommatidium. 120: Distribution of Octavius – subgroup A. 121: Distribution of Octavius – subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="10" pageNumber="505">120</figureCitation>
,
<figureCitation id="13592A24E954FFFBF339FEF3FE4DFE5F" box="[353,395,367,393]" captionStart="FIGURES 128–133" captionStartId="25.[151,264,1651,1675]" captionTargetBox="[154,1434,182,1627]" captionTargetId="figure-15@25.[154,1434,181,1627]" captionTargetPageId="25" captionText="FIGURES 128–133. 128: Cathedral Peak, Masongwane river banks. 129: Injisuthi. 130: Monkʼs Cowl, Hlatikulu forest. 131: Lotheni. 132: Kamberg, locality 1. 133: Kamberg locality 2." figureDoi="http://doi.org/10.5281/zenodo.11064306" httpUri="https://zenodo.org/record/11064306/files/figure.png" pageId="10" pageNumber="505">131</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C378652AE954FFFBF2CFFE2BFA4FFDB7" pageId="10" pageNumber="505" type="materials_examined">
<paragraph id="8BDD36A1E954FFFBF2CFFE2BFA95FE07" blockId="10.[151,1437,438,1689]" box="[151,1363,438,465]" pageId="10" pageNumber="505">
<materialsCitation id="3B0A3CFCE954FFFBF2CFFE2BFA89FE07" box="[151,1359,438,465]" country="South Africa" location="Lotheni National Reserve" municipality="Maloti-Drakensberg Park" pageId="10" pageNumber="505" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<emphasis id="B916EAB3E954FFFBF2CFFE2BFEF0FE06" bold="true" box="[151,310,438,465]" pageId="10" pageNumber="505">
<typeStatus id="54D98803E954FFFBF2CFFE2BFF14FE07" box="[151,210,439,465]" pageId="10" pageNumber="505">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3757631E954FFFBF364FE2BFE0BFE07" box="[316,461,439,465]" name="South Africa" pageId="10" pageNumber="505">South Africa</collectingCountry>
,
<collectingRegion id="49A6F843E954FFFBF381FE2BFD33FE07" box="[473,757,439,465]" country="South Africa" name="KwaZulu-Natal" pageId="10" pageNumber="505">KwaZulu-Natal Province</collectingRegion>
,
<collectingMunicipality id="6BB9ACDBE954FFFBF159FE2BFBE6FE07" box="[769,1056,439,465]" pageId="10" pageNumber="505">Maloti-Drakensberg Park</collectingMunicipality>
,
<location id="8EBD607AE954FFFBF674FE2BFA89FE07" LSID="urn:lsid:plazi:treatment:03CB87B7E954FFFBF2CFFED7FAFAF94F:8EBD607AE954FFFBF674FE2BFA89FE07" box="[1068,1359,439,465]" country="South Africa" municipality="Maloti-Drakensberg Park" name="Lotheni National Reserve" pageId="10" pageNumber="505" stateProvince="KwaZulu-Natal">Lotheni National Reserve</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFE47FA4FFDB7" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FFE47FEB2FE22" bold="true" box="[199,372,474,501]" pageId="10" pageNumber="505">Type material</emphasis>
(
<specimenCount id="9D64FD28E954FFFBF3D2FE47FDFEFE23" box="[394,568,475,501]" count="149" pageId="10" pageNumber="505" type="generic">149 specimens</specimenCount>
).
<emphasis id="B916EAB3E954FFFBF00CFE47FD07FE23" bold="true" box="[596,705,475,501]" pageId="10" pageNumber="505">
<typeStatus id="54D98803E954FFFBF00CFE47FD07FE23" box="[596,705,475,501]" pageId="10" pageNumber="505" type="holotype">Holotype</typeStatus>
</emphasis>
<materialsCitation id="3B0A3CFCE954FFFBF097FE47FD79FDEB" collectingDate="2019-01-14" collectorName="J. Janak" country="South Africa" determinerName="J. Janak" elevation="1570" location="Maloti-Drakensberg Park" pageId="10" pageNumber="505" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="9D64FD28E954FFFBF097FE47FD21FE23" box="[719,743,475,501]" count="1" pageId="10" pageNumber="505" type="male">♂</specimenCount>
: “
<collectingCountry id="F3757631E954FFFBF15BFE47FC5AFE23" box="[771,924,475,501]" name="South Africa" pageId="10" pageNumber="505">South Africa</collectingCountry>
,
<collectingRegion id="49A6F843E954FFFBF1F6FE47FBA7FE23" box="[942,1121,475,501]" country="South Africa" name="KwaZulu-Natal" pageId="10" pageNumber="505">KwaZulu-Natal</collectingRegion>
,
<location id="8EBD607AE954FFFBF62AFE47FA5EFE23" LSID="urn:lsid:plazi:treatment:03CB87B7E954FFFBF2CFFED7FAFAF94F:8EBD607AE954FFFBF62AFE47FA5EFE23" box="[1138,1432,475,501]" country="South Africa" name="Maloti-Drakensberg Park" pageId="10" pageNumber="505" stateProvince="KwaZulu-Natal">Maloti-Drakensberg Park</location>
, Lotheni NR, ind. forest patch 29°26.3ʼS 29°31.7ʼE,
<quantity id="4C9A9B44E954FFFBF0A0FE62FC8EFDCF" box="[760,840,510,537]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="10" pageNumber="505" unit="m" value="1570.0">
<elevation id="004FD192E954FFFBF0A0FE62FC8EFDCF" box="[760,840,510,537]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="10" pageNumber="505" unit="m" value="1570.0">1570m</elevation>
</quantity>
,
<date id="FFDC1061E954FFFBF10FFE63FC05FDCF" box="[855,963,511,537]" pageId="10" pageNumber="505" value="2019-01-14">
<collectingDate id="EF98E989E954FFFBF10FFE63FC05FDCF" box="[855,963,511,537]" pageId="10" pageNumber="505" value="2019-01-14">14.i.2019</collectingDate>
</date>
,
<collectorName id="26975377E954FFFBF18AFE63FBE9FDCF" box="[978,1071,511,537]" pageId="10" pageNumber="505">J. Janák</collectorName>
lgt.”, “
<typeStatus id="54D98803E954FFFBF6DDFE63FAE8FDCF" box="[1157,1326,511,537]" pageId="10" pageNumber="505" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C624D22E954FFFBF761FE63FECCFDEB" authority="Janák, 2024" authorityName="Janák" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="lothenicus" status="sp. nov.">
<emphasis id="B916EAB3E954FFFBF761FE63FECCFDEB" italics="true" pageId="10" pageNumber="505">Octavius lothenicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557C8E954FFFBF349FDBFFEA1FDEB" box="[273,359,547,573]" pageId="10" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="7E01DA65E954FFFBF335FDBFFE03FDEB" box="[365,453,547,573]" pageId="10" pageNumber="505">J. Janák</determinerName>
det. 2021”.
<typeStatus id="54D98803E954FFFBF016FDBFFD79FDEB" box="[590,703,547,573]" pageId="10" pageNumber="505" type="paratype">Paratypes</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B0A3CFCE954FFFBF091FDBFFA43FDB7" collectionCode="TMSA, JJRC" country="South Africa" determinerName="J. Janak" location="All" municipality="Maloti-Drakensberg Park" pageId="10" pageNumber="505" specimenCount="150" specimenCount-female="87" specimenCount-male="63" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="9D64FD28E954FFFBF091FDBFFCC4FDEB" box="[713,770,547,573]" count="62" pageId="10" pageNumber="505" type="male">62 ♂</specimenCount>
,
<specimenCount id="9D64FD28E954FFFBF156FDBFFC81FDEB" box="[782,839,547,573]" count="86" pageId="10" pageNumber="505" type="female">86 ♀</specimenCount>
: same data as holotype (
<collectionCode id="ED73AE64E954FFFBF60FFDBFFB6FFDEB" box="[1111,1193,547,573]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="505">TMSA</collectionCode>
,
<collectionCode id="ED73AE64E954FFFBF6ECFDBFFB34FDEB" box="[1204,1266,547,573]" pageId="10" pageNumber="505">JJRC</collectionCode>
,
<specimenCount id="9D64FD28E954FFFBF6A5FDBFFAEEFDEB" box="[1277,1320,547,573]" count="1" pageId="10" pageNumber="505" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D64FD28E954FFFBF76CFDBFFA9BFDEB" box="[1332,1373,547,573]" count="1" pageId="10" pageNumber="505" type="female">1 ♀</specimenCount>
in 96 % alcohol).
<location id="8EBD607AE954FFFBF346FDDBFE84FDB7" LSID="urn:lsid:plazi:treatment:03CB87B7E954FFFBF2CFFED7FAFAF94F:8EBD607AE954FFFBF346FDDBFE84FDB7" box="[286,322,583,609]" name="All" pageId="10" pageNumber="505">All</location>
<typeStatus id="54D98803E954FFFBF310FDDBFE75FDB7" box="[328,435,583,609]" pageId="10" pageNumber="505" type="paratype">paratypes</typeStatus>
with additional labels: “
<typeStatus id="54D98803E954FFFBF09CFDDBFCA3FDB7" box="[708,869,583,609]" pageId="10" pageNumber="505" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="4C624D22E954FFFBF134FDDBFB8FFDB7" authority="Janák, 2024" authorityName="Janák" authorityYear="2024" box="[876,1097,583,609]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="lothenicus" status="sp. nov.">
<emphasis id="B916EAB3E954FFFBF134FDDBFB8FFDB7" box="[876,1097,583,609]" italics="true" pageId="10" pageNumber="505">Octavius lothenicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557C8E954FFFBF608FDDBFB61FDB7" box="[1104,1191,583,609]" pageId="10" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="7E01DA65E954FFFBF6F5FDDBFAC0FDB7" box="[1197,1286,583,609]" pageId="10" pageNumber="505">J. Janák</determinerName>
det. 2021”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C378652AE954FFFBF29FFDF6FC00FAE7" pageId="10" pageNumber="505" type="description">
<paragraph id="8BDD36A1E954FFFBF29FFDF6FE82FD1B" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FFDF6FE9FFD52" bold="true" box="[199,345,618,644]" pageId="10" pageNumber="505">Description.</emphasis>
Body length
<quantity id="4C9A9B44E954FFFBF3ABFDF7FDBDFD53" box="[499,635,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="10" pageNumber="505" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
(M
<quantity id="4C9A9B44E954FFFBF0F5FDF6FCD2FD53" box="[685,788,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="10" pageNumber="505" unit="mm" value="1.75">1.75 mm</quantity>
, HT
<quantity id="4C9A9B44E954FFFBF115FDF7FC6EFD53" box="[845,936,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="505" unit="mm" value="1.8">1.8 mm</quantity>
), forebody length
<quantity id="4C9A9B44E954FFFBF62FFDF7FB39FD53" box="[1143,1279,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="10" pageNumber="505" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
(M
<quantity id="4C9A9B44E954FFFBF769FDF7FA5EFD53" box="[1329,1432,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="505" unit="mm" value="0.86">0.86 mm</quantity>
, HT
<quantity id="4C9A9B44E954FFFBF29AFD13FEDDFD7F" box="[194,283,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="505" unit="mm" value="0.9">0.9 mm</quantity>
). Macrophthalmous, apterous, rusty, head dull, pronotum slightly shiny, elytra and abdomen moderately shiny (
<figureCitation id="13592A24E954FFFBF2BBFD2FFEFEFD1B" box="[227,312,691,717]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 44</figureCitation>
).
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFD4BFBD3FC8B" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
Head (
<figureCitation id="13592A24E954FFFBF349FD4BFEA5FD27" box="[273,355,727,753]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 44</figureCitation>
) slightly narrower than pronotum (R 0.89–0.95, M 0.91, HT 0.90), eyes relatively large, temples less than half longer than eyes (R 1.21–1.67, M 1.44, HT 1.36), sides of head almost straight, not distinctly widened towards posterior angles, posterior angles moderately angular, median impression on disc absent, lateral parts of head moderately granulose, median part very densely and very finely reticulate.
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFCFBFCC5FC3B" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
Pronotum (
<figureCitation id="13592A24E954FFFBF31DFCFBFE51FC57" box="[325,407,871,897]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 44</figureCitation>
) slightly broader than long (R 1.07–1.14, M 1.11, HT 1.12), strongly narrowed posteriorly; anterior angles slightly angular, dorsal impressions moderately deep, transverse impression deep, lateral impressions deep, delimited by sharp longitudinal ridge laterally; lateral parts beside lateral impressions densely granulose, remainder of surface very densely and finely reticulate.
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFC6BFE5DFB8F" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
Elytra (
<figureCitation id="13592A24E954FFFBF343FC6BFEAAFBC7" box="[283,364,1015,1041]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 44</figureCitation>
) subquadrate, much broader than long (R 1.46–1.58, M 1.52, HT 1.48), with sharp longitudinal ridge laterally; between latter and suture sparsely, irregularly granulate, moderately reticulate and moderately sparsely shortly setose.
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFBFFFB4AFB77" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">Abdomen subparallel, slightly widened to segment IV or V, with one paratergite and microsculpture consisting of triangular or rhomboid fields, leaving here and there small smooth places, finely setose.</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FFB37FC00FAE7" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
Male. Sternite VIII moderately triangularly emarginated in posterior one-sixth (
<figureCitation id="13592A24E954FFFBF612FB37FB58FB13" box="[1098,1182,1195,1221]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 47</figureCitation>
), sternite IX as in
<figureCitation id="13592A24E954FFFBF728FB37FF70FB3F" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 48</figureCitation>
. Aedeagus only slightly asymmetrical, moderately long and narrow (
<quantity id="4C9A9B44E954FFFBF1E1FB53FB99FB3F" box="[953,1119,1231,1257]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="10" pageNumber="505" unit="mm" value="0.435" valueMax="0.44" valueMin="0.43">0.43–0.44 mm</quantity>
, M
<quantity id="4C9A9B44E954FFFBF6D3FB53FB37FB3F" box="[1163,1265,1231,1257]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="505" unit="mm" value="0.43">0.43 mm</quantity>
, HT
<quantity id="4C9A9B44E954FFFBF770FB53FA56FB3F" box="[1320,1424,1231,1257]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="10" pageNumber="505" unit="mm" value="0.44">0.44 mm</quantity>
), apical part divided in two short protrusions, internal structure undulate, strongly sclerotised; parameres markedly shorter than median lobe, with about 5–7 apicolateral setae (
<figureCitation id="13592A24E954FFFBF163FA8BFC7CFAE7" box="[827,954,1302,1329]" captionStart="FIGURES 39–48" captionStartId="9.[151,264,1791,1815]" captionTargetBox="[185,1403,194,1748]" captionTargetId="figure-15@9.[172,1416,181,1766]" captionTargetPageId="9" captionText="FIGURES 39–48. 39–43: Octavius kambergensis Janák, sp. nov., 39–40, 42–43, holotype; 41, paratype. 44–48: Octavius lothenicus Janák, sp. nov., 44–45, 47–48, holotype; 46, paratype. 39, 44, habitus; 40–41, 45–46, aedeagus ventral; 42, 47, male sternite VIII; 43, 48, male sternite IX. Scale 0.5 mm: 39, 44; 0.1 mm: remaining figures." figureDoi="http://doi.org/10.5281/zenodo.11064286" httpUri="https://zenodo.org/record/11064286/files/figure.png" pageId="10" pageNumber="505">Fig. 45–46</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C378652AE954FFFBF29FFAA6FAB8FA33" pageId="10" pageNumber="505" type="diagnosis">
<paragraph id="8BDD36A1E954FFFBF29FFAA6FAB8FA33" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FFAA6FE0AFA82" bold="true" box="[199,460,1338,1364]" pageId="10" pageNumber="505">Differential diagnosis.</emphasis>
<taxonomicName id="4C624D22E954FFFBF389FAA7FD32FA83" authority="Janak" authorityName="Janak" authorityYear="2024" box="[465,756,1339,1365]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="lothenicus" status="sp. nov.">
<emphasis id="B916EAB3E954FFFBF389FAA7FD6AFA83" box="[465,684,1339,1365]" italics="true" pageId="10" pageNumber="505">Octavius lothenicus</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E954FFFBF0A5FAA7FC93FA83" bold="true" box="[765,853,1339,1365]" pageId="10" pageNumber="505">
<taxonomicNameLabel id="A22557C8E954FFFBF0A5FAA7FC93FA83" box="[765,853,1339,1365]" pageId="10" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs among species with large eyes, with temples at most twice as long as eyes, with the head not or at most slightly widened posteriorly, with large body and the disc of the head without a median impression. In the most recent key of South African
<taxonomicName id="4C624D22E954FFFBF61AFA1FFA87FA4B" authority="(Janak 2014)" baseAuthorityName="Janak" baseAuthorityYear="2014" box="[1090,1345,1411,1437]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B916EAB3E954FFFBF61AFA1FFB60FA4B" box="[1090,1190,1411,1437]" italics="true" pageId="10" pageNumber="505">Octavius</emphasis>
(
<bibRefCitation id="EFF34B50E954FFFBF6EDFA1FFAFFFA4B" author="Janak, J." box="[1205,1337,1411,1437]" pageId="10" pageNumber="505" pagination="195 - 231" refId="ref16165" refString="Janak, J. (2014) New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae). Acta Entomologica Musei Nationalis Pragae, 54 (1), 195 - 231." type="journal article" year="2014">Janák 2014</bibRefCitation>
)
</taxonomicName>
is to be arranged at the couplet 154 (151), but it differs from the species arranged there (e. g.
<taxonomicName id="4C624D22E954FFFBF609FA3BFA9AFA17" authority="Janak, 2014" authorityName="Janak" authorityYear="2014" box="[1105,1372,1447,1473]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B916EAB3E954FFFBF609FA3BFB09FA17" box="[1105,1231,1447,1473]" italics="true" pageId="10" pageNumber="505">O. mikhaili</emphasis>
<bibRefCitation id="EFF34B50E954FFFBF68EFA3BFA9AFA17" author="Janak, J." box="[1238,1372,1447,1473]" pageId="10" pageNumber="505" pagination="195 - 231" refId="ref16165" refString="Janak, J. (2014) New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae). Acta Entomologica Musei Nationalis Pragae, 54 (1), 195 - 231." type="journal article" year="2014">Janák, 2014</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="4C624D22E954FFFBF7D9FA3BFE76FA33" authority="Janak, 2014" authorityName="Janak" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="multisetosus">
<emphasis id="B916EAB3E954FFFBF7D9FA3BFEE7FA33" italics="true" pageId="10" pageNumber="505">O. multisetosus</emphasis>
<bibRefCitation id="EFF34B50E954FFFBF371FA57FE76FA33" author="Janak, J." box="[297,432,1483,1509]" pageId="10" pageNumber="505" pagination="195 - 231" refId="ref16165" refString="Janak, J. (2014) New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae). Acta Entomologica Musei Nationalis Pragae, 54 (1), 195 - 231." type="journal article" year="2014">Janák, 2014</bibRefCitation>
</taxonomicName>
) by almost symmetrical aedeagus with the apical part divided in two short protrusions.
</paragraph>
</subSubSection>
<subSubSection id="C378652AE954FFFBF29FFA72FB0AF9DF" box="[199,1228,1518,1545]" pageId="10" pageNumber="505" type="etymology">
<paragraph id="8BDD36A1E954FFFBF29FFA72FB0AF9DF" blockId="10.[151,1437,438,1689]" box="[199,1228,1518,1545]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FFA72FE62F9DE" bold="true" box="[199,420,1518,1544]" pageId="10" pageNumber="505">Derivatio nominis.</emphasis>
This species is named after the
<typeStatus id="54D98803E954FFFBF152FA73FCFCF9DF" box="[778,826,1519,1545]" pageId="10" pageNumber="505">type</typeStatus>
locality – Lotheni Natural Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C378652AE954FFFBF29FF98EFAFAF94F" pageId="10" pageNumber="505" type="distribution">
<paragraph id="8BDD36A1E954FFFBF29FF98EFC79F987" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FF98EFEA7F9FA" bold="true" box="[199,353,1554,1580]" pageId="10" pageNumber="505">Distribution.</emphasis>
<taxonomicName id="4C624D22E954FFFBF33FF98FFD49F9FB" authority="Janak" authorityName="Janak" authorityYear="2024" box="[359,655,1555,1581]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="lothenicus" status="sp. nov.">
<emphasis id="B916EAB3E954FFFBF33FF98FFD83F9FB" box="[359,581,1555,1581]" italics="true" pageId="10" pageNumber="505">Octavius lothenicus</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E954FFFBF0C2F98FFD32F9FB" bold="true" box="[666,756,1555,1581]" pageId="10" pageNumber="505">
<taxonomicNameLabel id="A22557C8E954FFFBF0C2F98FFD32F9FB" box="[666,756,1555,1581]" pageId="10" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently recorded only from the Lotheni Natural Reserve in Drakensberg Mts.,
<collectingRegion id="49A6F843E954FFFBF3D2F9ABFD61F987" box="[394,679,1591,1617]" country="South Africa" name="KwaZulu-Natal" pageId="10" pageNumber="505">KwaZulu-Natal Province</collectingRegion>
,
<collectingCountry id="F3757631E954FFFBF0EAF9ABFC87F987" box="[690,833,1591,1617]" name="South Africa" pageId="10" pageNumber="505">South Africa</collectingCountry>
(
<figureCitation id="13592A24E954FFFBF108F9ABFC74F987" box="[848,946,1591,1617]" captionStart="FIGURES 113–121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113–121. Octavius undulatus Janák, sp. nov.; 113–116, 118–119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116–117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114–115; 116–117; 118–119; o: ommatidium. 120: Distribution of Octavius – subgroup A. 121: Distribution of Octavius – subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="10" pageNumber="505">Fig. 120</figureCitation>
).
</paragraph>
<paragraph id="8BDD36A1E954FFFBF29FF9C6FAFAF94F" blockId="10.[151,1437,438,1689]" pageId="10" pageNumber="505">
<emphasis id="B916EAB3E954FFFBF29FF9C6FE8DF9A2" bold="true" box="[199,331,1626,1652]" pageId="10" pageNumber="505">Bionomics.</emphasis>
All specimens were collected in siftings of forest litter in a patch of an indigenous forest at the elevation of about
<quantity id="4C9A9B44E954FFFBF330F9E2FE7BF94F" box="[360,445,1662,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="10" pageNumber="505" unit="m" value="1570.0">1570 m</quantity>
a.s.l. with the abundance of 11.6 specimens per kg of sifted material (
<figureCitation id="13592A24E954FFFBF695F9E3FAE9F94F" box="[1229,1327,1663,1689]" captionStart="FIGURES 128–133" captionStartId="25.[151,264,1651,1675]" captionTargetBox="[154,1434,182,1627]" captionTargetId="figure-15@25.[154,1434,181,1627]" captionTargetPageId="25" captionText="FIGURES 128–133. 128: Cathedral Peak, Masongwane river banks. 129: Injisuthi. 130: Monkʼs Cowl, Hlatikulu forest. 131: Lotheni. 132: Kamberg, locality 1. 133: Kamberg locality 2." figureDoi="http://doi.org/10.5281/zenodo.11064306" httpUri="https://zenodo.org/record/11064306/files/figure.png" pageId="10" pageNumber="505">Fig. 131</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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