268 lines
28 KiB
XML
268 lines
28 KiB
XML
<document id="40A72F0F9E00E807711FED4D8FB366F5" ID-CLB-Dataset="29271" ID-DOI="10.11646/zootaxa.4496.1.11" ID-GBIF-Dataset="688e03b4-7031-4dfc-96f3-d5d354e4f8ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446737" ID-ZooBank="AC1033D3-3D56-4ADC-9764-F628F75457A3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747053050" checkinUser="plazi" docAuthor="Szederjesi, Tímea, Dányi, László, Kaydan, Mehmet Bora & Csuzdi, Csaba" docDate="2018" docId="FD1387B1FFFBED4CFF55FF146CA8F97B" docLanguage="en" docName="zootaxa.4496.1.11.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Healyella zicsii Szederjesi & Dányi & Kaydan & Csuzdi 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="168" masterDocId="012AFFC9FFF3ED44FFC2FD536934FFC0" masterDocTitle="Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili)" masterLastPageNumber="172" masterPageNumber="160" pageNumber="168" updateTime="1698565628082" updateUser="ExternalLinkService">
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<mods:titleInfo id="B7EE9439C1A2A3502E296C1FF0D31913">
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<mods:title id="DEED5AF2733D5132591A8A89DEB9086A">Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili)</mods:title>
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<mods:name id="FA5E077D00D387CD0410C603CEA125C3" type="personal">
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<mods:namePart id="E6FE00575047AFAAACA71BD2FFAA697D">Szederjesi, Tímea</mods:namePart>
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<mods:namePart id="B226E8A8E3FF2EBC57340E41A400C7FB">Dányi, László</mods:namePart>
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<mods:namePart id="A069228A8ED053AEEEE2203F63739AD9">Kaydan, Mehmet Bora</mods:namePart>
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</mods:name>
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<mods:name id="853D6A42A804957B8BC89DE07A51F2F6" type="personal">
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<mods:roleTerm id="22C87875E06BA7A91E1CF7155271DF30">Author</mods:roleTerm>
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<mods:namePart id="FB23EC4248BA07E2A561C4DBAE7F8B3A">Csuzdi, Csaba</mods:namePart>
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<mods:title id="0A6F0DDA8DD2E4CB09F6BE10C8774D82">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="293529C2FCECD8FF975CC87FCF0BF236">
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<mods:date id="2386AEE4D885DF7E8C55D876F57804C8">2018</mods:date>
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<mods:number id="2795EC450FB760DD98890008D6A76B66">2018-10-05</mods:number>
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<mods:number id="301C1D3C738A002947427907C008EA37">4496</mods:number>
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<mods:identifier id="4769A00A66DA1DE8D900DD1CDF5599F2" type="DOI">10.11646/zootaxa.4496.1.11</mods:identifier>
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<mods:identifier id="CF483A24DB2B1C83AA603C46081F2288" type="GBIF-Dataset">688e03b4-7031-4dfc-96f3-d5d354e4f8ee</mods:identifier>
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<treatment id="FD1387B1FFFBED4CFF55FF146CA8F97B" ID-DOI="http://doi.org/10.5281/zenodo.5980404" ID-GBIF-Taxon="148639721" ID-Zenodo-Dep="5980404" LSID="urn:lsid:plazi:treatment:FD1387B1FFFBED4CFF55FF146CA8F97B" httpUri="http://treatment.plazi.org/id/FD1387B1FFFBED4CFF55FF146CA8F97B" lastPageNumber="168" pageId="8" pageNumber="168">
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<subSubSection id="3DA0652CFFFBED4CFF55FF146833FD43" pageId="8" pageNumber="168" type="nomenclature">
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<paragraph id="750536A7FFFBED4CFF55FF146B82FDA2" blockId="8.[151,694,583,643]" box="[151,694,583,610]" pageId="8" pageNumber="168">
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<heading id="2E4D81CBFFFBED4CFF55FF146B82FDA2" bold="true" box="[151,694,583,610]" fontSize="11" level="1" pageId="8" pageNumber="168" reason="1">
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<emphasis id="47CEEAB5FFFBED4CFF55FF146B82FDA2" bold="true" box="[151,694,583,610]" pageId="8" pageNumber="168">
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<taxonomicName id="B2BA4D24FFFBED4CFF55FF146B65FDA2" authority="Szederjesi & Csuzdi" authorityName="Szederjesi & Dányi & Kaydan & Csuzdi" authorityYear="2018" box="[151,593,583,610]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="zicsii" status="sp. nov.">
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<emphasis id="47CEEAB5FFFBED4CFF55FF14687BFDA1" bold="true" box="[151,335,583,609]" italics="true" pageId="8" pageNumber="168">Healyella zicsii</emphasis>
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Szederjesi & Csuzdi
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</taxonomicName>
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<taxonomicNameLabel id="5CFD57CEFFFBED4CFD9BFF1B6B82FDA2" box="[601,694,584,610]" pageId="8" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="750536A7FFFBED4CFF55FF396833FD43" blockId="8.[151,694,583,643]" box="[151,263,618,643]" pageId="8" pageNumber="168">
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(
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<figureCitation id="ED812A22FFFBED4CFF5CFF3969CBFD43" box="[158,255,618,643]" captionStart="FIGURE 4" captionStartId="9.[151,250,1320,1342]" captionTargetBox="[340,1247,483,1293]" captionTargetId="figure@9.[340,1247,483,1299]" captionTargetPageId="9" captionText="FIGURE 4. Healyella zicsii sp. nov. (A, C) and Healyella baloghi (Zicsi, 1981) (B). A. Setal ratio of Healyella zicsii sp. nov. B. Setal ratio of Healyella baloghi. C. Ventrolateral view of the forebody of He. zicsii sp. nov." httpUri="https://zenodo.org/record/1446745/files/figure.png" pageId="8" pageNumber="168">Figure 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="3DA0652CFFFBED4CFF55FFE16A5BFC9B" pageId="8" pageNumber="168" type="materials_examined">
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<paragraph id="750536A7FFFBED4CFF55FFE16AD2FD2F" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
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<materialsCitation id="C5D23CFAFFFBED4CFF55FFE16AD2FD2F" ID-GBIF-Occurrence="1914265194" collectionCode="HNHM" collectorName="L. Danyi & Kaydan" elevation="103" latitude="41.529865" location="Derbent" longLatPrecision="1" longitude="32.492783" pageId="8" pageNumber="168" specimenCount="1" specimenCount-adult="1" stateProvince="Bartin" typeStatus="holotype">
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<emphasis id="47CEEAB5FFFBED4CFF55FFE1683FFD0B" bold="true" box="[151,267,690,715]" pageId="8" pageNumber="168">
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<typeStatus id="AA018805FFFBED4CFF55FFE16832FD0B" box="[151,262,690,715]" pageId="8" pageNumber="168" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="13ABAE62FFFBED4CFED8FFE16845FD0B" box="[282,369,690,715]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="168">HNHM</collectionCode>
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/17262
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<specimenCount id="63BCFD2EFFFBED4CFE0FFFE06B17FD0B" box="[461,547,690,715]" pageId="8" pageNumber="168" type="adult">1 adult</specimenCount>
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ex.,
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<emphasis id="47CEEAB5FFFBED4CFDA8FFE06B89FD0B" box="[618,701,691,715]" italics="true" pageId="8" pageNumber="168">No. 20</emphasis>
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,
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<location id="7065607CFFFBED4CFD16FFE16A05FD0B" LSID="urn:lsid:plazi:treatment:FD1387B1FFFBED4CFF55FF146CA8F97B:7065607CFFFBED4CFD16FFE16A05FD0B" box="[724,817,690,715]" latitude="41.529865" longLatPrecision="1" longitude="32.492783" name="Derbent" pageId="8" pageNumber="168" stateProvince="Bartin">Derbent</location>
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,
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<collectingRegion id="B77EF845FFFBED4CFC87FFE06ABAFD0A" box="[837,910,691,714]" country="Turkey" name="Bartin" pageId="8" pageNumber="168">Bartın</collectingRegion>
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, SE of the village, disturbed beech forest,
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<geoCoordinate id="108E5060FFFBED4CFF55FF846812FD2F" box="[151,294,727,751]" direction="north" orientation="latitude" pageId="8" pageNumber="168" precision="1" value="41.529865">N41°31.792'</geoCoordinate>
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<geoCoordinate id="108E5060FFFBED4CFEE9FF85688EFD2F" box="[299,442,726,751]" direction="east" orientation="longitude" pageId="8" pageNumber="168" precision="1" value="32.492783">E32°29.567'</geoCoordinate>
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,
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<elevation id="FE97D194FFFBED4CFE06FF846B39FD2E" box="[452,525,727,751]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="168" unit="m" value="103.0">
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<quantity id="B2429B42FFFBED4CFE06FF846B39FD2E" box="[452,525,727,751]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="168" unit="m" value="103.0">103 m</quantity>
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||
</elevation>
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, leg.
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<collectorName id="D84F5371FFFBED4CFD8BFF846B9BFD2F" box="[585,687,726,751]" pageId="8" pageNumber="168">L. Dányi</collectorName>
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||
, M.B.
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<collectorName id="D84F5371FFFBED4CFD39FF856A61FD2F" box="[763,853,726,751]" pageId="8" pageNumber="168">Kaydan</collectorName>
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||
, 22.0 4.2015.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="750536A7FFFBED4CFF05FFA96A5BFC9B" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
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||
<materialsCitation id="C5D23CFAFFFBED4CFF05FFA96C05FCF7" ID-GBIF-Occurrence="1914265199" collectionCode="HNHM" collectorName="L. Danyi & Kaydan" elevation="656" latitude="41.40497" location="Incecay" longLatPrecision="1" longitude="32.71932" pageId="8" pageNumber="168" specimenCount="1" specimenCount-adult="1" stateProvince="Karabuk" typeStatus="paratype">
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<emphasis id="47CEEAB5FFFBED4CFF05FFA96872FCD3" bold="true" box="[199,326,762,787]" pageId="8" pageNumber="168">
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<typeStatus id="AA018805FFFBED4CFF05FFA96875FCD3" box="[199,321,762,787]" pageId="8" pageNumber="168" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="13ABAE62FFFBED4CFE90FFA9689DFCD3" box="[338,425,762,787]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="168">HNHM</collectionCode>
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/17056
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<specimenCount id="63BCFD2EFFFBED4CFDC0FFA86B61FCD3" box="[514,597,762,787]" pageId="8" pageNumber="168" type="adult">1 adult</specimenCount>
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||
ex.,
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<emphasis id="47CEEAB5FFFBED4CFD54FFA86BD2FCD3" box="[662,742,763,787]" italics="true" pageId="8" pageNumber="168">No. 23</emphasis>
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,
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<location id="7065607CFFFBED4CFD3BFFA86A65FCD3" LSID="urn:lsid:plazi:treatment:FD1387B1FFFBED4CFF55FF146CA8F97B:7065607CFFFBED4CFD3BFFA86A65FCD3" box="[761,849,763,787]" latitude="41.40497" longLatPrecision="1" longitude="32.71932" name="Incecay" pageId="8" pageNumber="168" stateProvince="Karabuk">Inceçay</location>
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,
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<collectingRegion id="B77EF845FFFBED4CFCA0FFA96AF1FCD3" box="[866,965,762,787]" country="Turkey" name="Karabuk" pageId="8" pageNumber="168">Karabük</collectingRegion>
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, NW of the village, stream in a mixed
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<taxonomicName id="B2BA4D24FFFBED4CFF55FE4D69CFFCF6" box="[151,251,798,822]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="8" pageNumber="168" phylum="Tracheophyta" rank="genus">
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<emphasis id="47CEEAB5FFFBED4CFF55FE4D69CFFCF6" box="[151,251,798,822]" italics="true" pageId="8" pageNumber="168">Platanus</emphasis>
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</taxonomicName>
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and pine forest,
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<geoCoordinate id="108E5060FFFBED4CFE07FE4C6B67FCF7" box="[453,595,799,823]" direction="north" orientation="latitude" pageId="8" pageNumber="168" precision="1" value="41.40497">N41°24.298'</geoCoordinate>
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<geoCoordinate id="108E5060FFFBED4CFD9FFE4D6BDFFCF7" box="[605,747,798,823]" direction="east" orientation="longitude" pageId="8" pageNumber="168" precision="1" value="32.71932">E32°43.159'</geoCoordinate>
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,
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<elevation id="FE97D194FFFBED4CFD3BFE4D6A72FCF6" box="[761,838,798,823]" metricMagnitude="2" metricUnit="m" metricValue="6.56" pageId="8" pageNumber="168" unit="m" value="656.0">
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||
<quantity id="B2429B42FFFBED4CFD3BFE4D6A72FCF6" box="[761,838,798,823]" metricMagnitude="2" metricUnit="m" metricValue="6.56" pageId="8" pageNumber="168" unit="m" value="656.0">656 m</quantity>
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||
</elevation>
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, leg.
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<collectorName id="D84F5371FFFBED4CFC4BFE4C6AC7FCF7" box="[905,1011,798,823]" pageId="8" pageNumber="168">L. Dányi</collectorName>
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||
, M.B.
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<collectorName id="D84F5371FFFBED4CFB84FE4D6D94FCF7" box="[1094,1184,798,823]" pageId="8" pageNumber="168">Kaydan</collectorName>
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, 22.0 4.2015
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</materialsCitation>
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. HNHM/ 17079 1 adult ex., locality and date same as that of the Holotype.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3DA0652CFFFBED4CFF05FE3568AFFC63" pageId="8" pageNumber="168" type="etymology">
|
||
<paragraph id="750536A7FFFBED4CFF05FE3568AFFC63" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
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||
<emphasis id="47CEEAB5FFFBED4CFF05FE356893FCBF" bold="true" box="[199,423,870,895]" pageId="8" pageNumber="168">Derivatio nominis.</emphasis>
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The new species is named in honour of the late Professor András Zicsi, the renowned earthworm taxonomist.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3DA0652CFFFBED4CFF05FEFD68ADFAEF" pageId="8" pageNumber="168" type="diagnosis">
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||
<paragraph id="750536A7FFFBED4CFF05FEFD6A61FBF3" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
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||
<emphasis id="47CEEAB5FFFBED4CFF05FEFD6875FC07" bold="true" box="[199,321,942,967]" pageId="8" pageNumber="168">Diagnosis.</emphasis>
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Length
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||
<quantity id="B2429B42FFFBED4CFE66FEFC6B05FC06" box="[420,561,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="9.65" metricValueMax="10.9" metricValueMin="8.4" pageId="8" pageNumber="168" unit="mm" value="96.5" valueMax="109.0" valueMin="84.0">84–109 mm</quantity>
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||
, diameter
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||
<quantity id="B2429B42FFFBED4CFD6EFEFD6A17FC06" box="[684,803,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="8" pageNumber="168" unit="mm" value="6.75" valueMax="7.5" valueMin="6.0">6–7.5 mm</quantity>
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, setae distant. Colour pale, pigmentation lacking. First dorsal pore in 5/6. Clitellum on 26–31, tubercles lacking, clitellar bands present. Male pores on 15. Nephridial pores aligned above
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<emphasis id="47CEEAB5FFFBED4CFEBDFEA568B9FBCE" box="[383,397,1014,1038]" italics="true" pageId="8" pageNumber="168">b</emphasis>
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. Two pairs of vesicles in 11, 12. Spermathecae lacking. Calciferous glands in 10–12. Hearts in segments 7–11, nephridial bladders simple, sausage-shaped.
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</paragraph>
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||
<paragraph id="750536A7FFFBED4CFF05F96D68ADFAEF" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
|
||
<emphasis id="47CEEAB5FFFBED4CFF05F96D688FFB97" bold="true" box="[199,443,1086,1111]" pageId="8" pageNumber="168">External characters.</emphasis>
|
||
<typeStatus id="AA018805FFFBED4CFE03F96D6B1CFB96" box="[449,552,1086,1110]" pageId="8" pageNumber="168" type="holotype">
|
||
<emphasis id="47CEEAB5FFFBED4CFE03F96D6B1CFB96" box="[449,552,1086,1110]" italics="true" pageId="8" pageNumber="168">Holotype</emphasis>
|
||
</typeStatus>
|
||
<quantity id="B2429B42FFFBED4CFDEDF96C6B49FB96" box="[559,637,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="168" unit="mm" value="84.0">84 mm</quantity>
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||
long and
|
||
<quantity id="B2429B42FFFBED4CFD29F96D6A18FB96" box="[747,812,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="168" unit="mm" value="6.0">6 mm</quantity>
|
||
wide. Number of segments 162.
|
||
<typeStatus id="AA018805FFFBED4CFB5FF96C6C3BFB96" box="[1181,1295,1087,1110]" pageId="8" pageNumber="168" type="paratype">
|
||
<emphasis id="47CEEAB5FFFBED4CFB5FF96C6C3BFB96" box="[1181,1295,1087,1110]" italics="true" pageId="8" pageNumber="168">Paratypes</emphasis>
|
||
</typeStatus>
|
||
<quantity id="B2429B42FFFBED4CFAD7F96C6CA8FB96" box="[1301,1436,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="9.8" metricValueMax="10.9" metricValueMin="8.7" pageId="8" pageNumber="168" unit="mm" value="98.0" valueMax="109.0" valueMin="87.0">87–109 mm</quantity>
|
||
long and
|
||
<quantity id="B2429B42FFFBED4CFECCF93168BCFBBA" box="[270,392,1122,1147]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="8" pageNumber="168" unit="mm" value="7.25" valueMax="7.5" valueMin="7.0">7–7.5 mm</quantity>
|
||
wide. Number of segments 157–176. Colour greyish preserved, pigmentation lacking. Prostomium prolobous. First dorsal pore at intersegmental furrow 5/6. Setae distant. Setal arrangement behind clitellum:
|
||
<emphasis id="47CEEAB5FFFBED4CFECFF9F8681DFB02" box="[269,297,1195,1218]" italics="true" pageId="8" pageNumber="168">aa</emphasis>
|
||
:
|
||
<emphasis id="47CEEAB5FFFBED4CFEF3F9F86879FB03" box="[305,333,1195,1219]" italics="true" pageId="8" pageNumber="168">ab</emphasis>
|
||
:
|
||
<emphasis id="47CEEAB5FFFBED4CFE96F9F8685BFB03" box="[340,367,1195,1219]" italics="true" pageId="8" pageNumber="168">bc</emphasis>
|
||
:
|
||
<emphasis id="47CEEAB5FFFBED4CFEB5F9F868A5FB03" box="[375,401,1195,1219]" italics="true" pageId="8" pageNumber="168">cd</emphasis>
|
||
:
|
||
<emphasis id="47CEEAB5FFFBED4CFE5BF9F86881FB03" box="[409,437,1195,1219]" italics="true" pageId="8" pageNumber="168">dd</emphasis>
|
||
= 2.3:1.6:1.1:1:4 (
|
||
<figureCitation id="ED812A22FFFBED4CFD50F9F96BC4FB03" box="[658,752,1194,1219]" captionStart="FIGURE 4" captionStartId="9.[151,250,1320,1342]" captionTargetBox="[340,1247,483,1293]" captionTargetId="figure@9.[340,1247,483,1299]" captionTargetPageId="9" captionText="FIGURE 4. Healyella zicsii sp. nov. (A, C) and Healyella baloghi (Zicsi, 1981) (B). A. Setal ratio of Healyella zicsii sp. nov. B. Setal ratio of Healyella baloghi. C. Ventrolateral view of the forebody of He. zicsii sp. nov." httpUri="https://zenodo.org/record/1446745/files/figure.png" pageId="8" pageNumber="168">Fig. 4A</figureCitation>
|
||
). Male pores great slits facing ventrad on segment 15 just above setal line
|
||
<emphasis id="47CEEAB5FFFBED4CFE89F99C686DFB26" box="[331,345,1231,1254]" italics="true" pageId="8" pageNumber="168">a</emphasis>
|
||
, located on the rear of the segment. Nephridial pores aligned above setal line
|
||
<emphasis id="47CEEAB5FFFBED4CFB7EF99D6DFEFB26" box="[1212,1226,1230,1254]" italics="true" pageId="8" pageNumber="168">b</emphasis>
|
||
. Clitellum annular on 26–31. Tubercula pubertatis lacking but a pair of darker clitellar bands stretches ventrolaterally on both sides of the clitellum (
|
||
<figureCitation id="ED812A22FFFBED4CFEF1F84568B8FAEF" box="[307,396,1302,1327]" captionStart="FIGURE 4" captionStartId="9.[151,250,1320,1342]" captionTargetBox="[340,1247,483,1293]" captionTargetId="figure@9.[340,1247,483,1299]" captionTargetPageId="9" captionText="FIGURE 4. Healyella zicsii sp. nov. (A, C) and Healyella baloghi (Zicsi, 1981) (B). A. Setal ratio of Healyella zicsii sp. nov. B. Setal ratio of Healyella baloghi. C. Ventrolateral view of the forebody of He. zicsii sp. nov." httpUri="https://zenodo.org/record/1446745/files/figure.png" pageId="8" pageNumber="168">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DA0652CFFFBED4CFF05F8696AFBFA23" pageId="8" pageNumber="168" type="description">
|
||
<paragraph id="750536A7FFFBED4CFF05F8696AFBFA23" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
|
||
<emphasis id="47CEEAB5FFFBED4CFF05F869689AFA93" bold="true" box="[199,430,1338,1363]" pageId="8" pageNumber="168">Internal characters</emphasis>
|
||
(based on holotype and one paratype). Septa 6/7–
|
||
<date id="01041067FFFBED4CFC1DF8696D4CFA93" box="[991,1144,1338,1363]" pageId="8" pageNumber="168" value="2015-07-08" valueMax="2015-07-14" valueMin="2015-07-08">7/8 and 14/15</date>
|
||
slightly thickened,
|
||
<date id="01041067FFFBED4CFA96F8686987FAB7" pageId="8" pageNumber="168" value="2010-08-09" valueMax="2010-08-09" valueMin="2010-08-09">8/9–9/ 10</date>
|
||
thickened,
|
||
<date id="01041067FFFBED4CFEF6F80C68F4FAB7" box="[308,448,1375,1399]" pageId="8" pageNumber="168" value="2014-10-11" valueMax="2014-10-13" valueMin="2014-10-11">10/11–13/14</date>
|
||
strongly thickened. Testes and funnels paired in 10–11. Two pairs of seminal vesicles in 11, and 12. Spermathecae lacking. Calciferous glands in 10–12 with small diverticula in 10. Last heart in 11, with a pair of extraoesophageal vessels in 12. Nephridial bladders simple, sausage-shaped. Crop in 15–16, gizzard in 17– 18. Typhlosolis large, bilobate. Longitudinal musculature of pinnate type.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DA0652CFFFBED4CFF05F8BD6CA8F97B" pageId="8" pageNumber="168" type="discussion">
|
||
<paragraph id="750536A7FFFBED4CFF05F8BD6CA8F97B" blockId="8.[151,1437,690,1723]" pageId="8" pageNumber="168">
|
||
<emphasis id="47CEEAB5FFFBED4CFF05F8BD6800F9C7" bold="true" box="[199,308,1518,1543]" pageId="8" pageNumber="168">Remarks</emphasis>
|
||
. There are three
|
||
<taxonomicName id="B2BA4D24FFFBED4CFE3BF8BD6B52F9C6" authorityName="Omodeo & Rota" authorityYear="1989" box="[505,614,1518,1542]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="genus">
|
||
<emphasis id="47CEEAB5FFFBED4CFE3BF8BD6B52F9C6" box="[505,614,1518,1542]" italics="true" pageId="8" pageNumber="168">Healyella</emphasis>
|
||
</taxonomicName>
|
||
species with clitellum on 26–31:
|
||
<emphasis id="47CEEAB5FFFBED4CFC29F8BC6D27F9C6" box="[1003,1043,1519,1542]" italics="true" pageId="8" pageNumber="168">He.</emphasis>
|
||
<taxonomicName id="B2BA4D24FFFBED4CFBDEF8BD6C3BF9C7" authority="(Zicsi, 1981)" baseAuthorityName="Zicsi" baseAuthorityYear="1981" box="[1052,1295,1518,1543]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="baloghi">
|
||
<emphasis id="47CEEAB5FFFBED4CFBDEF8BD6D45F9C6" box="[1052,1137,1518,1542]" italics="true" pageId="8" pageNumber="168">baloghi</emphasis>
|
||
(
|
||
<bibRefCitation id="112B4B56FFFBED4CFB40F8BD6C32F9C7" box="[1154,1286,1518,1543]" pageId="8" pageNumber="168" refId="ref10414" refString="Zicsi, A. (1981) Probleme der Lumbriciden-Systematik sowie die Revision zweier Gattungen (Oligochaeta). Acta Zoologica Hungarica, 27, 431 - 442." type="journal article">Zicsi, 1981</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="47CEEAB5FFFBED4CFADEF8BC6CAFF9C6" box="[1308,1435,1519,1542]" italics="true" pageId="8" pageNumber="168">He. mariae</emphasis>
|
||
<bibRefCitation id="112B4B56FFFBED4CFF55FB4168A3F9EB" box="[151,407,1554,1579]" pageId="8" pageNumber="168" refId="ref9572" refString="Omodeo, P. & Rota, E. (1989) Earthworms of Turkey. Bolletino di Zoologia, 56, 167 - 199. https: // doi. org / 10.1080 / 11250008909355639" type="journal article">Omodeo & Rota, 1989</bibRefCitation>
|
||
and
|
||
<emphasis id="47CEEAB5FFFBED4CFE0CFB406B1BF9EA" box="[462,559,1555,1578]" italics="true" pageId="8" pageNumber="168">He. naja</emphasis>
|
||
Omodeo & Rota, 1981. The latter two are small-sized purple-red worms with well-developed sexual ridges above the male pores.
|
||
<emphasis id="47CEEAB5FFFBED4CFD28FB646A60F98E" box="[746,852,1591,1614]" italics="true" pageId="8" pageNumber="168">
|
||
He.
|
||
<taxonomicName id="B2BA4D24FFFBED4CFCD9FB646A60F98E" authorityName="Szederjesi & Dányi & Kaydan & Csuzdi" authorityYear="2018" box="[795,852,1591,1614]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="zicsii">zicsii</taxonomicName>
|
||
</emphasis>
|
||
is greyish without pigmentation, twice as long and wide as
|
||
<emphasis id="47CEEAB5FFFBED4CFF3AFB086868F9B2" box="[248,348,1627,1650]" italics="true" pageId="8" pageNumber="168">He. naja</emphasis>
|
||
and
|
||
<emphasis id="47CEEAB5FFFBED4CFE5AFB086B2DF9B2" box="[408,537,1627,1650]" italics="true" pageId="8" pageNumber="168">He. mariae</emphasis>
|
||
and lacks sexual ridges on the male pores. The new species differs from
|
||
<emphasis id="47CEEAB5FFFBED4CFAB6FB0869D8F956" italics="true" pageId="8" pageNumber="168">
|
||
He.
|
||
<taxonomicName id="B2BA4D24FFFBED4CFF55FB2D69D8F956" baseAuthorityName="Zicsi" baseAuthorityYear="1981" box="[151,236,1662,1686]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="baloghi">baloghi</taxonomicName>
|
||
</emphasis>
|
||
in the setal arrangement which is distant in
|
||
<emphasis id="47CEEAB5FFFBED4CFD35FB2C6A57F956" box="[759,867,1663,1686]" italics="true" pageId="8" pageNumber="168">
|
||
He.
|
||
<taxonomicName id="B2BA4D24FFFBED4CFCEBFB2C6A57F956" authorityName="Szederjesi & Dányi & Kaydan & Csuzdi" authorityYear="2018" box="[809,867,1663,1686]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="zicsii">zicsii</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="ED812A22FFFBED4CFCB7FB2D6AFAF957" box="[885,974,1662,1687]" captionStart="FIGURE 4" captionStartId="9.[151,250,1320,1342]" captionTargetBox="[340,1247,483,1293]" captionTargetId="figure@9.[340,1247,483,1299]" captionTargetPageId="9" captionText="FIGURE 4. Healyella zicsii sp. nov. (A, C) and Healyella baloghi (Zicsi, 1981) (B). A. Setal ratio of Healyella zicsii sp. nov. B. Setal ratio of Healyella baloghi. C. Ventrolateral view of the forebody of He. zicsii sp. nov." httpUri="https://zenodo.org/record/1446745/files/figure.png" pageId="8" pageNumber="168">Fig. 4B</figureCitation>
|
||
) but quite paired in
|
||
<emphasis id="47CEEAB5FFFBED4CFB03FB2C6C7CF956" box="[1217,1352,1662,1686]" italics="true" pageId="8" pageNumber="168">
|
||
He.
|
||
<taxonomicName id="B2BA4D24FFFBED4CFB31FB2D6C7CF956" baseAuthorityName="Zicsi" baseAuthorityYear="1981" box="[1267,1352,1662,1686]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="baloghi">baloghi</taxonomicName>
|
||
</emphasis>
|
||
, in the position of the male pore (between
|
||
<emphasis id="47CEEAB5FFFBED4CFDE2FBF06B08F97B" box="[544,572,1699,1723]" italics="true" pageId="8" pageNumber="168">ab</emphasis>
|
||
vs. between
|
||
<emphasis id="47CEEAB5FFFBED4CFD0EFBF06BD3F97B" box="[716,743,1699,1723]" italics="true" pageId="8" pageNumber="168">bc</emphasis>
|
||
) and in the presence of clitellar bands (absent in
|
||
<emphasis id="47CEEAB5FFFBED4CFACAFBF06CBFF97B" box="[1288,1419,1699,1723]" italics="true" pageId="8" pageNumber="168">
|
||
He.
|
||
<taxonomicName id="B2BA4D24FFFBED4CFAF4FBF06CBFF97B" baseAuthorityName="Zicsi" baseAuthorityYear="1981" box="[1334,1419,1699,1723]" class="Clitellata" family="Lumbricidae" genus="Healyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="168" phylum="Annelida" rank="species" species="baloghi">baloghi</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |