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<document id="354A4EEF699DE4A5C8CFEE746F539153" ID-CLB-Dataset="5455" ID-DOI="10.5852/ejt.2021.758.1431" ID-GBIF-Dataset="f01782d1-107a-4884-8319-4f9c53d8cdd1" ID-ISSN="2118-9773" ID-Zenodo-Dep="5101636" ID-ZooBank="5D21C06C-EE8D-43EC-B607-EDB9BF0B91F8" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626264912167" checkinUser="felipe" docAuthor="Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis &amp; Praz, Christophe J." docDate="2021" docId="F12A87E8FFA5FFDFFE52FBEDFD6BF9A6" docLanguage="en" docName="ejt-758_147-193.pdf" docOrigin="European Journal of Taxonomy 758" docSource="http://dx.doi.org/10.5852/ejt.2021.758.1431" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Taeniandrena) beaumonti Benoist 1961, stat. rev." docType="treatment" docVersion="8" lastPageNumber="162" masterDocId="0D13FF90FFA9FFD0FFE7FFE5FFABFF8F" masterDocTitle="Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae)" masterLastPageNumber="193" masterPageNumber="147" pageNumber="159" updateTime="1699036717860" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BDF975C275D4DECE36EA68B163DF8AEC">Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="1BA09E5280124CC8504791887BBC791D">Wood, Thomas J.</mods:namePart>
<mods:nameIdentifier id="86C7CFACE5017A33008CB79D348CECA4" type="LSID">670C3E36-1D28-4FCA-887C-91D6116E6F9C</mods:nameIdentifier>
<mods:affiliation id="53AAEF0986F3EDC3ECC6FEAEA41D3037">Laboratory of Zoology, University of Mons, Avenue du Champs de Mars 6, 7000 Mons, Belgium.</mods:affiliation>
<mods:nameIdentifier id="C7F48058CB373A956803FBA1857927B3" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:roleTerm id="3A0686B765C9736119616638D9AD3C14">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3EB45DB45277FB990D4564670D310DCD">Ghisbain, Guillaume</mods:namePart>
<mods:nameIdentifier id="C8B014FA5D1272B4165DC043B0233FC3" type="LSID">FD9CD87B-EAAB-4037-BFDE-58AE0E7349B7</mods:nameIdentifier>
<mods:affiliation id="34710667449DC400EEC3675E7D2A3DB2">Laboratory of Zoology, University of Mons, Avenue du Champs de Mars 6, 7000 Mons, Belgium.</mods:affiliation>
<mods:nameIdentifier id="F03AD1F2A5E8CD95F95096C6A606D3FB" type="email">guillaume.ghisbain@umons.ac.be</mods:nameIdentifier>
</mods:name>
<mods:name id="DAFAB27E9296884922ADA8AFBC4E96FD" type="personal">
<mods:role id="E877173C94D31E3B9DAC51736DA4F8E0">
<mods:roleTerm id="799529AD7476ECBBCC5731FE9BBDC5C3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26B4267113C476A7AA5D880D2E0A0A79">Michez, Denis</mods:namePart>
<mods:nameIdentifier id="B186357DA31C1C8B38C5AEDE37F2FE50" type="LSID">8B04585A-FE00-4D9A-AFD6-1BD2A1584CFA</mods:nameIdentifier>
<mods:affiliation id="72CDBC623C2D0819BE171538B3DB6493">Laboratory of Zoology, University of Mons, Avenue du Champs de Mars 6, 7000 Mons, Belgium.</mods:affiliation>
<mods:nameIdentifier id="DFE0FBBDAE9318CA295747EB207EFEAD" type="email">denis.michez@umons.ac.be</mods:nameIdentifier>
</mods:name>
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<mods:role id="D8890C10F2CAE92056E1BF4F2F61CA33">
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</mods:role>
<mods:namePart id="AF7A31A0679BA741323BEFA21CEBA81C">Praz, Christophe J.</mods:namePart>
<mods:nameIdentifier id="EDDE64F35C624B2EEC3B0537DFAD2D59" type="LSID">4435BC3F-1647-4D9C-BA92-74775C2C704E</mods:nameIdentifier>
<mods:affiliation id="852859F7C9319E3227460E371D3F2E1E">University of Neuchâtel, Institute of Biology, Rue Emile-Argand 11, 2000 Neuchâtel, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="D8DB276052E2251DD2AB50263ED5150C" type="email">christophe.praz@unine.ch</mods:nameIdentifier>
</mods:name>
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<mods:title id="87AEDD059A7020FFF03F514FD26DC606">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D15912582199A72544B6188AE4C4D5D2">
<mods:date id="949B2F9842966D3C6A2CDBB96C9BEC04">2021</mods:date>
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<mods:number id="57D81C2D96B6F556FF9B8C54385C70E1">2021-07-12</mods:number>
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<mods:number id="8994F3820FC4E949DF1FEA47BE0F0E9D">758</mods:number>
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<mods:url id="8CAB3307DFE81A2E21696A84409E0F9A">http://dx.doi.org/10.5852/ejt.2021.758.1431</mods:url>
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<mods:identifier id="3403F0F13D46A2051E8CD58B452ADCC7" type="DOI">10.5852/ejt.2021.758.1431</mods:identifier>
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<treatment id="F12A87E8FFA5FFDFFE52FBEDFD6BF9A6" ID-DOI="http://doi.org/10.5281/zenodo.5103113" ID-GBIF-Taxon="183542964" ID-Zenodo-Dep="5103113" LSID="urn:lsid:plazi:treatment:F12A87E8FFA5FFDFFE52FBEDFD6BF9A6" httpUri="http://treatment.plazi.org/id/F12A87E8FFA5FFDFFE52FBEDFD6BF9A6" lastPageId="15" lastPageNumber="162" pageId="12" pageNumber="159">
<subSubSection id="31996575FFA5FFDCFE52FBEDFBD4FBAC" box="[437,1151,1032,1059]" pageId="12" pageNumber="159" type="nomenclature">
<paragraph id="793C36FEFFA5FFDCFE52FBEDFBD4FBAC" blockId="12.[437,1151,1032,1095]" box="[437,1151,1032,1059]" pageId="12" pageNumber="159">
<heading id="22748192FFA5FFDCFE52FBEDFBD4FBAC" box="[437,1151,1032,1059]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="159" reason="2">
<taxonomicName id="BE834D7DFFA5FFDCFE52FBEDFBBEFBAD" ID-CoL="DR54" authority="Benoist, 1961" authorityName="Benoist" authorityYear="1961" box="[437,1045,1032,1059]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="159" phylum="Arthropoda" rank="species" species="beaumonti" status="stat. nov." subGenus="Taeniandrena">
<emphasis id="4BF7EAECFFA5FFDCFE52FBEDFCC0FBAC" bold="true" box="[437,875,1032,1059]" pageId="12" pageNumber="159">
<emphasis id="4BF7EAECFFA5FFDCFE52FBEDFDB4FBAC" bold="true" box="[437,543,1032,1059]" italics="true" pageId="12" pageNumber="159">Andrena</emphasis>
(
<emphasis id="4BF7EAECFFA5FFDCFDD7FBEDFD73FBAC" bold="true" box="[560,728,1032,1059]" italics="true" pageId="12" pageNumber="159">Taeniandrena</emphasis>
)
<emphasis id="4BF7EAECFFA5FFDCFD0DFBEDFCC0FBAC" bold="true" box="[746,875,1032,1059]" italics="true" pageId="12" pageNumber="159">beaumonti</emphasis>
</emphasis>
<bibRefCitation id="1D124B0FFFA5FFDCFC95FBEDFBBEFBAD" author="Benoist R." box="[882,1045,1032,1058]" pageId="12" pageNumber="159" pagination="85 - 95" refId="ref21766" refString="Benoist R. 1961. Hymenopteres recoltes par une mission Suisse au Maroc (1947) Apidae, genre Andrena. Bulletin de la Societe des Sciences naturelles et physiques du Maroc 41: 85 - 95." type="journal article" year="1961">Benoist, 1961</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA5FFDCFBFBFBECFBD4FBAC" box="[1052,1151,1033,1059]" pageId="12" pageNumber="159">stat. rev.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="31996575FFA5FFDCFDB3FBC9FC74FBC8" box="[596,991,1068,1095]" pageId="12" pageNumber="159" type="description">
<paragraph id="793C36FEFFA5FFDCFDB3FBC9FC74FBC8" blockId="12.[437,1151,1032,1095]" box="[596,991,1068,1095]" pageId="12" pageNumber="159">
<figureCitation id="E1B82A7BFFA5FFDCFDB3FBC9FC99FBC9" box="[596,818,1068,1095]" captionStart="Figs 20–27. 20, 22, 24, 26" captionStartId="13.[189,241,1926,1952]" captionTargetBox="[254,1333,268,1886]" captionTargetId="figure-71@13.[254,1333,268,1886]" captionTargetPageId="13" captionText="Figs 20–27. 20, 22, 24, 26. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 20. Female profile. 22. Female face. 24. Female scutum. 26. Female terga. — 21, 23, 25, 27. Andrena beaumonti stat. rev. (TJWC). 21. Female profile. 23. Female face. 25. Female scutum. 27. Female terga. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101650" httpUri="https://zenodo.org/record/5101650/files/figure.png" pageId="12" pageNumber="159">Figs 21, 23, 25, 27</figureCitation>
,
<figureCitation id="E1B82A7BFFA5FFDCFCD9FBC9FC74FBC8" box="[830,991,1068,1095]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="12" pageNumber="159">29, 31, 33, 35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="31996575FFA5FFDCFF5AFB8BFAA5FB06" box="[189,1294,1134,1161]" pageId="12" pageNumber="159" type="reference_group">
<paragraph id="793C36FEFFA5FFDCFF5AFB8BFAA5FB06" blockId="12.[189,1294,1134,1161]" box="[189,1294,1134,1161]" pageId="12" pageNumber="159">
<treatmentCitationGroup id="599311D0FFA5FFDCFF5AFB8BFAA5FB06" box="[189,1294,1134,1161]" pageId="12" pageNumber="159">
<treatmentCitation id="F82210EFFFA5FFDCFF5AFB8BFDD5FB06" author="Benoist R." box="[189,638,1134,1161]" page="91" pageId="12" pageNumber="159" year="1961">
<taxonomicName id="BE834D7DFFA5FFDCFF5AFB8BFDD5FB06" ID-CoL="62BS5" authority="Beaumonti Benoist, 1961: 91" authorityName="Beaumonti Benoist" authorityPageNumber="91" authorityYear="1961" box="[189,638,1134,1161]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="4BF7EAECFFA5FFDCFF5AFB8BFE0CFB07" box="[189,423,1134,1160]" italics="true" pageId="12" pageNumber="159">Andrena Beaumonti</emphasis>
<bibRefCitation id="1D124B0FFFA5FFDCFE48FB8BFDD5FB06" author="Benoist R." box="[431,638,1134,1161]" pageId="12" pageNumber="159" pagination="85 - 95" refId="ref21766" refString="Benoist R. 1961. Hymenopteres recoltes par une mission Suisse au Maroc (1947) Apidae, genre Andrena. Bulletin de la Societe des Sciences naturelles et physiques du Maroc 41: 85 - 95." type="journal article" year="1961">Benoist, 1961: 91</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<collectingCountry id="0194766EFFA5FFDCFD69FB8AFD56FB06" box="[654,765,1135,1161]" name="Morocco" pageId="12" pageNumber="159">Morocco</collectingCountry>
, between Asni and Arroud,
<date id="0D3D103EFFA5FFDCFBA5FB8AFB70FB06" box="[1090,1243,1135,1161]" pageId="12" pageNumber="159" value="1947-06-18">18 Jun. 1947</date>
,
<specimenCount id="6F85FD77FFA5FFDCFB00FB8AFB57FB06" box="[1255,1276,1135,1161]" pageId="12" pageNumber="159" type="female">♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="31996575FFA5FFDCFF5AFB55FCD3FACC" pageId="12" pageNumber="159" type="materials_examined">
<paragraph id="793C36FEFFA5FFDCFF5AFB55FE01FB45" blockId="12.[189,426,1200,1226]" box="[189,426,1200,1226]" pageId="12" pageNumber="159">
<heading id="22748192FFA5FFDCFF5AFB55FE01FB45" bold="true" box="[189,426,1200,1226]" fontSize="11" level="3" pageId="12" pageNumber="159" reason="3">
<emphasis id="4BF7EAECFFA5FFDCFF5AFB55FE01FB45" bold="true" box="[189,426,1200,1226]" pageId="12" pageNumber="159">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="793C36FEFFA5FFDCFF5AFB05FCD3FACC" blockId="12.[189,1398,1248,1349]" pageId="12" pageNumber="159">
<materialsCitation id="C9EB3CA3FFA5FFDCFF5AFB05FB74FB72" ID-GBIF-Occurrence="3321923315" box="[189,1247,1248,1277]" collectingDate="1987-06-29" collectionCode="OOLM" collectorName="M. Schwarz" country="Morocco" elevation="1900" location="MOROCCO" pageId="12" pageNumber="159" specimenCount="1" specimenCount-female="1">
<collectingCountry id="0194766EFFA5FFDCFF5AFB05FEFAFB75" box="[189,337,1248,1274]" name="Morocco" pageId="12" pageNumber="159">MOROCCO</collectingCountry>
•
<specimenCount id="6F85FD77FFA5FFDCFE8AFB05FE37FB74" box="[365,412,1248,1275]" pageId="12" pageNumber="159" type="female">1 ♀</specimenCount>
; Tizi-n-Test, S;
<elevation id="F2AED1CDFFA5FFDCFDB9FB05FD51FB75" box="[606,762,1248,1275]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="159" unit="m" value="1900.0">
<quantity id="BE7B9B1BFFA5FFDCFDB9FB05FD12FB74" box="[606,697,1248,1275]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="159" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
;
<date id="0D3D103EFFA5FFDCFCEEFB05FC0CFB75" box="[777,935,1248,1275]" pageId="12" pageNumber="159" value="1987-06-29">
<collectingDate id="1D79E9D6FFA5FFDCFCEEFB05FC0CFB75" box="[777,935,1248,1275]" pageId="12" pageNumber="159" value="1987-06-29">29 Jun. 1987</collectingDate>
</date>
;
<collectorName id="D4765328FFA5FFDCFC53FB04FBEFFB75" box="[948,1092,1248,1275]" pageId="12" pageNumber="159">M. Schwarz</collectorName>
leg.;
<collectionCode id="1F92AE3BFFA5FFDCFB6EFB04FB74FB72" box="[1161,1247,1249,1277]" pageId="12" pageNumber="159">OÖLM</collectionCode>
</materialsCitation>
•
<materialsCitation id="C9EB3CA3FFA5FFDCFB1CFB05FD20FA91" ID-GBIF-Occurrence="3321923342" collectingDate="1987-06-29" collectionCode="TJWC" collectorName="Tizi-n-Test, S &amp; M. Schwarz" country="Morocco" elevation="1900" location="MOROCCO" pageId="12" pageNumber="159" specimenCount="1" specimenCount-female="1">
<specimenCount id="6F85FD77FFA5FFDCFB1CFB05FA81FB74" box="[1275,1322,1248,1275]" pageId="12" pageNumber="159" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="1F92AE3BFFA5FFDCFDD9FAE1FD20FA91" box="[574,651,1284,1310]" pageId="12" pageNumber="159">TJWC</collectionCode>
</materialsCitation>
•
<materialsCitation id="C9EB3CA3FFA5FFDCFD42FAE1FCDFFACC" ID-GBIF-Occurrence="3321923466" collectingDate="2015-05-08" collectionCode="OOLM" collectorName="K. Denes" country="Morocco" elevation="2700" location="Oukaimeden" pageId="12" pageNumber="159" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="6F85FD77FFA5FFDCFD42FAE1FD78FA90" box="[677,723,1284,1311]" pageId="12" pageNumber="159" type="male">1 ♂</specimenCount>
,
<specimenCount id="6F85FD77FFA5FFDCFD07FAE1FCA5FA90" box="[736,782,1284,1311]" pageId="12" pageNumber="159" type="female">1 ♀</specimenCount>
;
<location id="7C5C6025FFA5FFDCFCFAFAE1FC1CFA91" LSID="urn:lsid:plazi:treatment:F12A87E8FFA5FFDFFE52FBEDFD6BF9A6:7C5C6025FFA5FFDCFCFAFAE1FC1CFA91" box="[797,951,1284,1310]" country="Morocco" name="Oukaimeden" pageId="12" pageNumber="159">Oukaimeden</location>
;
<elevation id="F2AED1CDFFA5FFDCFC24FAE1FBF7FA91" box="[963,1116,1284,1311]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="159" unit="m" value="2700.0">
<quantity id="BE7B9B1BFFA5FFDCFC24FAE1FBB7FA90" box="[963,1052,1284,1311]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="159" unit="m" value="2700.0">2700 m</quantity>
a.s.l.
</elevation>
;
<date id="0D3D103EFFA5FFDCFB8DFAE1FB57FA90" box="[1130,1276,1284,1311]" pageId="12" pageNumber="159" value="2015-05-08">
<collectingDate id="1D79E9D6FFA5FFDCFB8DFAE1FB57FA90" box="[1130,1276,1284,1311]" pageId="12" pageNumber="159" value="2015-05-08">8 May 2015</collectingDate>
</date>
;
<collectorName id="D4765328FFA5FFDCFAEEFAE0FADDFA91" box="[1289,1398,1284,1311]" pageId="12" pageNumber="159">K. Deneš</collectorName>
leg.;
<collectionCode id="1F92AE3BFFA5FFDCFF1FFACCFEE5FACA" box="[248,334,1321,1349]" pageId="12" pageNumber="159">OÖLM</collectionCode>
(illustrated
<figureCitation id="E1B82A7BFFA5FFDCFE3BFACDFD11FACD" box="[476,698,1320,1347]" captionStart="Figs 20–27. 20, 22, 24, 26" captionStartId="13.[189,241,1926,1952]" captionTargetBox="[254,1333,268,1886]" captionTargetId="figure-71@13.[254,1333,268,1886]" captionTargetPageId="13" captionText="Figs 20–27. 20, 22, 24, 26. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 20. Female profile. 22. Female face. 24. Female scutum. 26. Female terga. — 21, 23, 25, 27. Andrena beaumonti stat. rev. (TJWC). 21. Female profile. 23. Female face. 25. Female scutum. 27. Female terga. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101650" httpUri="https://zenodo.org/record/5101650/files/figure.png" pageId="12" pageNumber="159">Figs 21, 23, 25, 27</figureCitation>
,
<figureCitation id="E1B82A7BFFA5FFDCFD21FACDFCC0FACC" box="[710,875,1320,1347]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="12" pageNumber="159">29, 31, 33, 35</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31996575FFA5FFDFFF5AFA8FFF55FE78" lastPageId="15" lastPageNumber="162" pageId="12" pageNumber="159" type="description">
<paragraph id="793C36FEFFA5FFDCFF5AFA8FFEE5FA0B" blockId="12.[189,334,1386,1412]" box="[189,334,1386,1412]" pageId="12" pageNumber="159">
<heading id="22748192FFA5FFDCFF5AFA8FFEE5FA0B" bold="true" box="[189,334,1386,1412]" fontSize="11" level="3" pageId="12" pageNumber="159" reason="3">
<emphasis id="4BF7EAECFFA5FFDCFF5AFA8FFEE5FA0B" bold="true" box="[189,334,1386,1412]" pageId="12" pageNumber="159">Description</emphasis>
</heading>
</paragraph>
<paragraph id="793C36FEFFA5FFDCFF0DFA7FFE81FA3B" blockId="12.[189,729,1434,1496]" box="[234,298,1434,1460]" pageId="12" pageNumber="159">
<emphasis id="4BF7EAECFFA5FFDCFF0DFA7FFE81FA3B" bold="true" box="[234,298,1434,1460]" pageId="12" pageNumber="159">Male</emphasis>
</paragraph>
<paragraph id="793C36FEFFA5FFDCFF5AFA5BFD72FA57" blockId="12.[189,729,1434,1496]" box="[189,729,1470,1496]" pageId="12" pageNumber="159">
MEASUREMENTS. Body length
<quantity id="BE7B9B1BFFA5FFDCFDF4FA5BFDCFFA57" box="[531,612,1470,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="159" unit="mm" value="11.0">11 mm</quantity>
(
<figureCitation id="E1B82A7BFFA5FFDCFD93FA5BFD67FA57" box="[628,716,1470,1496]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="12" pageNumber="159">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="793C36FEFFA5FFDCFF5AF9E5FCD4F909" blockId="12.[189,1399,1536,1671]" pageId="12" pageNumber="159">
HEAD. Dark, slightly as wide as long. Clypeus flattened, densely punctate, punctures separated by 0.5 puncture diameters, underlying surface shiny, without microsculpture. Gena, face, vertex, and scape covered with long whitish hairs that equal or exceed scape in length. Antennae dark, A4–13 slightly lightened grey below, A3 1.15 times as long as A4 (
<figureCitation id="E1B82A7BFFA5FFDCFCFDF989FCDAF909" box="[794,881,1644,1670]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="12" pageNumber="159">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="793C36FEFFA5FFDFFF5AF94AFF55FE78" blockId="12.[188,1400,1710,2025]" lastBlockId="15.[188,1400,265,503]" lastPageId="15" lastPageNumber="162" pageId="12" pageNumber="159">
MESOSOMA. Scutum shagreened and dull over majority of disc, becoming weaker centrally and posteriorly, here weakly shining, surface densely but shallowly punctured, punctures apically separated by 0.5 puncture diameters, centrally becoming sparser, here separated by 1–2 puncture diameters. Scutellum more uniformly shiny and densely punctate, punctures separated by 1 puncture diameter. Episternum and propodeum weakly reticulate, underlying surface weakly shining, propodeal triangle basally comparatively more strongly reticulate, this dispersing apically. All parts of mesosoma with long brownish (scutum and scutellum) to whitish (episternum and propodeum) hairs that exceed length of scape. Legs dark, apex of hind tibiae and entirety of hind basitarsi orange, pubescence whitish. Wings hyaline, venation orange, nervulus interstitial. METASOMA. Terga dark, apical margins impressed, strongly impressed on T3–5, covered in short, thick, and complete white hair bands (
<figureCitation id="E1B82A7BFFA6FFDFFDD6FEC9FD23FEC9" box="[561,648,300,326]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="15" pageNumber="162">Fig. 33</figureCitation>
). Terga densely punctate, punctures separated by 0.5–1 puncture diameters, underlying surface shagreened and dull basally, becoming weaker apically, T4–5 weakly shining. S8 columnar, parallel sided, apically slightly notched. Genitalia simple, gonocoxa with internal margins diverging apically, laterally with weak shagreenation, apical corners rounded (
<figureCitation id="E1B82A7BFFA6FFDFFB20FE73FA8BFE3E" box="[1223,1312,406,433]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="15" pageNumber="162">Fig. 35</figureCitation>
). Penis valve relatively narrow, apically tapering, gonostyli with broad bases, apical blades clearly longer than wide.
</paragraph>
</subSubSection>
<caption id="2DFC6676FFA4FFDDFF5AF863FA86F868" ID-DOI="http://doi.org/10.5281/zenodo.5101650" ID-Zenodo-Dep="5101650" httpUri="https://zenodo.org/record/5101650/files/figure.png" pageId="13" pageNumber="160" startId="13.[189,241,1926,1952]" targetBox="[254,1333,268,1886]" targetPageId="13">
<paragraph id="793C36FEFFA4FFDDFF5AF863FA86F868" blockId="13.[189,1399,1926,2024]" pageId="13" pageNumber="160">
<emphasis id="4BF7EAECFFA4FFDDFF5AF863FDBFF82E" bold="true" box="[189,532,1926,1953]" pageId="13" pageNumber="160">Figs 20–27. 20, 22, 24, 26.</emphasis>
<taxonomicName id="BE834D7DFFA4FFDDFDC7F862FC09F82E" authority="Wood &amp; Praz" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" box="[544,930,1927,1953]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="160" phylum="Arthropoda" rank="species" species="benoisti" status="sp. nov.">
<emphasis id="4BF7EAECFFA4FFDDFDC7F862FD46F82E" box="[544,749,1927,1953]" italics="true" pageId="13" pageNumber="160">Andrena benoisti</emphasis>
Wood &amp; Praz
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA4FFDDFC48F862FBA5F82E" box="[943,1038,1927,1953]" pageId="13" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
(TJWC).
<emphasis id="4BF7EAECFFA4FFDDFB69F862FB00F82E" bold="true" box="[1166,1195,1927,1953]" pageId="13" pageNumber="160">20</emphasis>
. Female profile.
<emphasis id="4BF7EAECFFA4FFDDFF5AF84FFF71F84B" bold="true" box="[189,218,1962,1988]" pageId="13" pageNumber="160">22</emphasis>
. Female face.
<emphasis id="4BF7EAECFFA4FFDDFE6EF84FFE0DF84B" bold="true" box="[393,422,1962,1988]" pageId="13" pageNumber="160">24</emphasis>
. Female scutum.
<emphasis id="4BF7EAECFFA4FFDDFD9FF84FFD3EF84B" bold="true" box="[632,661,1962,1988]" pageId="13" pageNumber="160">26</emphasis>
. Female terga. —
<emphasis id="4BF7EAECFFA4FFDDFC95F84FFBBEF84B" bold="true" box="[882,1045,1962,1988]" pageId="13" pageNumber="160">21, 23, 25, 27</emphasis>
.
<taxonomicName id="BE834D7DFFA4FFDDFBC3F84FFAA7F84B" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[1060,1292,1962,1988]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="160" phylum="Arthropoda" rank="species" species="beaumonti" status="stat. nov.">
<emphasis id="4BF7EAECFFA4FFDDFBC3F84FFAA7F84B" box="[1060,1292,1962,1988]" italics="true" pageId="13" pageNumber="160">Andrena beaumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA4FFDDFAF4F84EFADCF84A" box="[1299,1399,1963,1989]" pageId="13" pageNumber="160">stat. rev.</taxonomicNameLabel>
(TJWC).
<emphasis id="4BF7EAECFFA4FFDDFECBF828FEE2F868" bold="true" box="[300,329,1997,2023]" pageId="13" pageNumber="160">21</emphasis>
. Female profile.
<emphasis id="4BF7EAECFFA4FFDDFDF6F828FD85F868" bold="true" box="[529,558,1997,2023]" pageId="13" pageNumber="160">23</emphasis>
. Female face.
<emphasis id="4BF7EAECFFA4FFDDFD3DF828FD5CF868" bold="true" box="[730,759,1997,2023]" pageId="13" pageNumber="160">25</emphasis>
. Female scutum.
<emphasis id="4BF7EAECFFA4FFDDFC20F828FC4FF868" bold="true" box="[967,996,1997,2023]" pageId="13" pageNumber="160">27</emphasis>
. Female terga. Not to scale.
</paragraph>
</caption>
<caption id="2DFC6676FFA7FFDEFF5AF862FB35F868" ID-DOI="http://doi.org/10.5281/zenodo.5101654" ID-Zenodo-Dep="5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="14" pageNumber="161" startId="14.[189,241,1927,1953]" targetBox="[253,1334,264,1890]" targetPageId="14">
<paragraph id="793C36FEFFA7FFDEFF5AF862FB35F868" blockId="14.[189,1399,1927,2024]" pageId="14" pageNumber="161">
<emphasis id="4BF7EAECFFA7FFDEFF5AF862FE5AF82E" bold="true" box="[189,497,1927,1953]" pageId="14" pageNumber="161">Figs 28–35. 28, 30, 32, 34</emphasis>
.
<taxonomicName id="BE834D7DFFA7FFDEFDE7F862FCC6F82E" authority="Wood &amp; Praz" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" box="[512,877,1927,1953]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="161" phylum="Arthropoda" rank="species" species="benoisti" status="sp. nov.">
<emphasis id="4BF7EAECFFA7FFDEFDE7F862FD63F82E" box="[512,712,1927,1953]" italics="true" pageId="14" pageNumber="161">Andrena benoisti</emphasis>
Wood &amp; Praz
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA7FFDEFC93F862FC65F82E" box="[884,974,1927,1953]" pageId="14" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
(TJWC).
<emphasis id="4BF7EAECFFA7FFDEFBA3F862FBCAF82E" bold="true" box="[1092,1121,1927,1953]" pageId="14" pageNumber="161">28</emphasis>
. Male profile.
<emphasis id="4BF7EAECFFA7FFDEFAE9F862FA80F82E" bold="true" box="[1294,1323,1927,1953]" pageId="14" pageNumber="161">30</emphasis>
. Male antennae.
<emphasis id="4BF7EAECFFA7FFDEFED2F84FFEF9F84B" bold="true" box="[309,338,1962,1988]" pageId="14" pageNumber="161">32</emphasis>
. Male terga.
<emphasis id="4BF7EAECFFA7FFDEFE15F84FFDA5F84B" bold="true" box="[498,526,1962,1988]" pageId="14" pageNumber="161">34</emphasis>
. Male genitalia. —
<emphasis id="4BF7EAECFFA7FFDEFD18F84FFC09F84B" bold="true" box="[767,930,1962,1988]" pageId="14" pageNumber="161">29, 31, 33, 35</emphasis>
.
<taxonomicName id="BE834D7DFFA7FFDEFC55F84FFB30F84B" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[946,1179,1962,1988]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="161" phylum="Arthropoda" rank="species" species="beaumonti" status="stat. nov.">
<emphasis id="4BF7EAECFFA7FFDEFC55F84FFB30F84B" box="[946,1179,1962,1988]" italics="true" pageId="14" pageNumber="161">Andrena beaumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA7FFDEFB44F84EFAACF84A" box="[1187,1287,1963,1989]" pageId="14" pageNumber="161">stat. rev.</taxonomicNameLabel>
(TJWC).
<emphasis id="4BF7EAECFFA7FFDEFF59F828FF71F868" bold="true" box="[190,218,1997,2023]" pageId="14" pageNumber="161">29</emphasis>
. Male profile.
<emphasis id="4BF7EAECFFA7FFDEFE6FF828FE0EF868" bold="true" box="[392,421,1997,2023]" pageId="14" pageNumber="161">31</emphasis>
. Male antennae.
<emphasis id="4BF7EAECFFA7FFDEFD89F828FD20F868" bold="true" box="[622,651,1997,2023]" pageId="14" pageNumber="161">33</emphasis>
. Male terga.
<emphasis id="4BF7EAECFFA7FFDEFCC0F828FCEFF868" bold="true" box="[807,836,1997,2023]" pageId="14" pageNumber="161">35</emphasis>
. Male genitalia. Not to scale.
</paragraph>
</caption>
<subSubSection id="31996575FFA6FFDFFF5AFDFAFCA7FB6B" pageId="15" pageNumber="162" type="discussion">
<paragraph id="793C36FEFFA6FFDFFF5AFDFAFE84FDB6" blockId="15.[189,303,543,569]" box="[189,303,543,569]" pageId="15" pageNumber="162">
<heading id="22748192FFA6FFDFFF5AFDFAFE84FDB6" bold="true" box="[189,303,543,569]" fontSize="11" level="3" pageId="15" pageNumber="162" reason="3">
<emphasis id="4BF7EAECFFA6FFDFFF5AFDFAFE84FDB6" bold="true" box="[189,303,543,569]" pageId="15" pageNumber="162">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="793C36FEFFA6FFDFFF5AFDABFCA7FB6B" blockId="15.[189,1400,589,1253]" pageId="15" pageNumber="162">
In his original description,
<bibRefCitation id="1D124B0FFFA6FFDFFDEBFDA8FD6BFDE7" author="Benoist R." box="[524,704,589,616]" pageId="15" pageNumber="162" pagination="85 - 95" refId="ref21766" refString="Benoist R. 1961. Hymenopteres recoltes par une mission Suisse au Maroc (1947) Apidae, genre Andrena. Bulletin de la Societe des Sciences naturelles et physiques du Maroc 41: 85 - 95." type="journal article" year="1961">Benoist (1961)</bibRefCitation>
drew a comparison between
<taxonomicName id="BE834D7DFFA6FFDFFBCAFDABFB65FDE8" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[1069,1230,589,616]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="beaumonti" status="stat. nov.">
<emphasis id="4BF7EAECFFA6FFDFFBCAFDABFB65FDE8" box="[1069,1230,589,616]" italics="true" pageId="15" pageNumber="162">A. beaumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA6FFDFFB3EFDABFAEAFDE7" box="[1241,1345,590,616]" pageId="15" pageNumber="162">stat. rev.</taxonomicNameLabel>
and
<taxonomicName id="BE834D7DFFA6FFDFFF5AFD94FD98FD04" authority="Panzer, 1799" authorityName="Panzer" authorityYear="1799" box="[189,563,625,651]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="4BF7EAECFFA6FFDFFF5AFD94FE2CFD04" box="[189,391,625,651]" italics="true" pageId="15" pageNumber="162">Andrena flavipes</emphasis>
Panzer, 1799
</taxonomicName>
, presumably because of the dense hair bands that help characterise this species.
<bibRefCitation id="1D124B0FFFA6FFDFFEBEFD71FDB0FD20" author="Warncke K." box="[345,539,660,687]" pageId="15" pageNumber="162" pagination="171 - 318" refId="ref23847" refString="Warncke K. 1967. Beitrag zur Klarung palaarktischer Andrena - Arten. Eos 43: 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
noted that this association was clearly incorrect at a subgeneric level, associating the bee with members of the subgenus
<taxonomicName id="BE834D7DFFA6FFDFFCFBFD52FC16FD5E" authorityName="Hedicke" authorityYear="1933" box="[796,957,695,721]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="subGenus" subGenus="Taeniandrena">
<emphasis id="4BF7EAECFFA6FFDFFCFBFD52FC16FD5E" box="[796,957,695,721]" italics="true" pageId="15" pageNumber="162">Taeniandrena</emphasis>
</taxonomicName>
instead. On the basis of its large and dense puncturing, its golden hairs flanking the pygidial plate, and the orange metatarsi of the second pair of legs,
<bibRefCitation id="1D124B0FFFA6FFDFFEFDFD1BFE7CFC96" author="Warncke K." box="[282,471,766,793]" pageId="15" pageNumber="162" pagination="171 - 318" refId="ref23847" refString="Warncke K. 1967. Beitrag zur Klarung palaarktischer Andrena - Arten. Eos 43: 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
associated this taxon with
<taxonomicName id="BE834D7DFFA6FFDFFCF4FD1BFC70FC97" authorityName="" baseAuthorityName="Kirby" baseAuthorityYear="1802" box="[787,987,766,792]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="wilkella">
<emphasis id="4BF7EAECFFA6FFDFFCF4FD1BFC70FC97" box="[787,987,766,792]" italics="true" pageId="15" pageNumber="162">Andrena wilkella</emphasis>
</taxonomicName>
, but as a subspecies because of the unbroken hair bands on T3+4. He then went on to report
<taxonomicName id="BE834D7DFFA6FFDFFC8CFCC7FBC5FCB4" authorityName="Benoist" authorityYear="1961" box="[875,1134,801,828]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="wilkella" subSpecies="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFC8CFCC7FBC5FCB4" box="[875,1134,801,828]" italics="true" pageId="15" pageNumber="162">A. wilkella beaumonti</emphasis>
</taxonomicName>
from
<collectingCountry id="0194766EFFA6FFDFFB5FFCC4FB57FCB4" box="[1208,1276,801,827]" name="Georgia" pageId="15" pageNumber="162">Iberia</collectingCountry>
(
<bibRefCitation id="1D124B0FFFA6FFDFFAEAFCC4FF57FCD0" author="Warncke K." pageId="15" pageNumber="162" pagination="119 - 223" refId="ref23991" refString="Warncke K. 1976. Die Bienengattung Andrena F., 1775, in Iberien (Hym. Apidae). Teil B. Eos 50: 119 - 223." type="journal article" year="1976">Warncke 1976</bibRefCitation>
; see also
<bibRefCitation id="1D124B0FFFA6FFDFFE90FCA0FDCDFCD0" author="Ortiz-Sanchez F. J." box="[375,614,837,863]" pageId="15" pageNumber="162" pagination="265 - 281" refId="ref23069" refString="Ortiz-Sanchez F. J. 2011. Lista actualizada de las especies de abejas de Espana (Hymenoptera: Apoidea: Apiformes). Boletin de la Sociedad Entomologica Aragonesa 49: 265 - 281." type="journal article" year="2011">Ortiz-Sánchez 2011</bibRefCitation>
,
<bibRefCitation id="1D124B0FFFA6FFDFFD91FCA0FD1EFCD0" author="Ortiz-Sanchez F. J." box="[630,693,837,863]" pageId="15" pageNumber="162" pagination="14" refId="ref23105" refString="Ortiz-Sanchez F. J. 2020. Checklist de Fauna Iberica. Serie Anthophila (Insecta: Hymenoptera: Apoidea) en la peninsula iberica e islas Baleares (edicion 2020). In: Ramos M. A. &amp; Sanchez Ruiz M. (eds) Documentos Fauna Iberica, 14. Museo Nacional de Ciencias Naturales, CSIC. Madrid." type="book chapter" year="2020">2020</bibRefCitation>
), giving it a distribution of
<collectingCountry id="0194766EFFA6FFDFFBEAFCA0FBD1FCD0" box="[1037,1146,837,863]" name="Morocco" pageId="15" pageNumber="162">Morocco</collectingCountry>
,
<collectingCountry id="0194766EFFA6FFDFFB6DFCA0FB7BFCD0" box="[1162,1232,837,863]" name="Spain" pageId="15" pageNumber="162">Spain</collectingCountry>
, and
<collectingCountry id="0194766EFFA6FFDFFAF2FCA0FAD3FCD0" box="[1301,1400,837,863]" name="Portugal" pageId="15" pageNumber="162">Portugal</collectingCountry>
(see also
<bibRefCitation id="1D124B0FFFA6FFDFFEC8FC8DFD82FC0D" author="Lhomme P. &amp; Michez D. &amp; Christmann S. &amp; Scheuchl E. &amp; El Abdouni I. &amp; Hamroud L. &amp; Ihsane O. &amp; Sentil A. &amp; Smaili M. C. &amp; Schwarz M. &amp; Dathe H. H. &amp; Straka J. &amp; Pauly A. &amp; Schmid-Egger C. &amp; Patiny S. &amp; Terzo M. &amp; Muller A. &amp; Praz C. &amp; Risch S. &amp; Kasparek M. &amp; Kuhlmann M. &amp; Wood T. J. &amp; Bogusch P. &amp; Ascher J. S. &amp; Rasmont P." box="[303,553,872,899]" pageId="15" pageNumber="162" pagination="1 - 159" refId="ref22569" refString="Lhomme P., Michez D., Christmann S., Scheuchl E., El Abdouni I., Hamroud L., Ihsane O., Sentil A., Smaili M. C., Schwarz M., Dathe H. H., Straka J., Pauly A., Schmid-Egger C., Patiny S., Terzo M., Muller A., Praz C., Risch S., Kasparek M., Kuhlmann M., Wood T. J., Bogusch P., Ascher J. S. &amp; Rasmont P. 2020. The wild bees (Hymenoptera: Apoidea) of Morocco. Zootaxa 4892: 1 - 159. https: // doi. org / 10.11646 / zootaxa. 4892.1.1" type="journal article" year="2020">
Lhomme
<emphasis id="4BF7EAECFFA6FFDFFE42FC8CFE73FC0D" box="[421,472,872,898]" italics="true" pageId="15" pageNumber="162">et al</emphasis>
. 2020
</bibRefCitation>
). However, genetic barcoding shows that
<taxonomicName id="BE834D7DFFA6FFDFFBC5FC8CFB6BFC0D" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[1058,1216,872,899]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFBC5FC8CFB6BFC0D" box="[1058,1216,872,899]" italics="true" pageId="15" pageNumber="162">A. beaumonti</emphasis>
</taxonomicName>
is distinct, and Moroccan material is also morphologically different from Warncke’s concept of
<taxonomicName id="BE834D7DFFA6FFDFFB92FC69FADCFC29" authorityName="Benoist" authorityYear="1961" box="[1141,1399,908,934]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="wilkella" subSpecies="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFB92FC69FADCFC29" box="[1141,1399,908,934]" italics="true" pageId="15" pageNumber="162">A. wilkella beaumonti</emphasis>
</taxonomicName>
in
<collectingCountry id="0194766EFFA6FFDFFF38FC4AFE8DFC46" box="[223,294,943,969]" name="Georgia" pageId="15" pageNumber="162">Iberia</collectingCountry>
, most obviously by lacking a shiny spot on the scutum that is well differentiated from the surrounding shagreenation. It can also be separated by the finer hairs on the scutum, the generally paler pubescence, and by the thicker and more complete hair bands on terga 2–4 (
<figureCitation id="E1B82A7BFFA6FFDFFBD8FC13FB62FB9F" box="[1087,1225,1014,1040]" captionStart="Figs 20–27. 20, 22, 24, 26" captionStartId="13.[189,241,1926,1952]" captionTargetBox="[254,1333,268,1886]" captionTargetId="figure-71@13.[254,1333,268,1886]" captionTargetPageId="13" captionText="Figs 20–27. 20, 22, 24, 26. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 20. Female profile. 22. Female face. 24. Female scutum. 26. Female terga. — 21, 23, 25, 27. Andrena beaumonti stat. rev. (TJWC). 21. Female profile. 23. Female face. 25. Female scutum. 27. Female terga. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101650" httpUri="https://zenodo.org/record/5101650/files/figure.png" pageId="15" pageNumber="162">Figs 20–27</figureCitation>
), meaning that
<taxonomicName id="BE834D7DFFA6FFDFFF59FBFCFD2EFBBC" authority="sensu Warncke" authorityName="Warncke" box="[190,645,1049,1076]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="subSpecies" species="wilkella" subSpecies="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFF59FBFCFE6CFBBC" box="[190,455,1049,1075]" italics="true" pageId="15" pageNumber="162">A. wilkella beaumonti</emphasis>
sensu Warncke
</taxonomicName>
in
<collectingCountry id="0194766EFFA6FFDFFD54FBFCFD5CFBBC" box="[691,759,1049,1075]" name="Georgia" pageId="15" pageNumber="162">Iberia</collectingCountry>
is actually undescribed (see below). The previously undescribed male of
<taxonomicName id="BE834D7DFFA6FFDFFE25FBD8FDCFFBD9" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[450,612,1084,1111]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFE25FBD8FDCFFBD9" box="[450,612,1084,1111]" italics="true" pageId="15" pageNumber="162">A. beaumonti</emphasis>
</taxonomicName>
can also be distinguished using the same characters (colour of pubescence, nature of tergal hair bands) and also by the length of A4 which is 1.15 times as long as A3, whereas in
<taxonomicName id="BE834D7DFFA6FFDFFE91FB66FD31FB11" authority="Wood &amp; Praz" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" box="[374,666,1155,1182]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="benoisti" status="sp. nov.">
<emphasis id="4BF7EAECFFA6FFDFFE91FB66FE5FFB12" box="[374,500,1155,1181]" italics="true" pageId="15" pageNumber="162">A. benoisti</emphasis>
Wood &amp; Praz
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA6FFDFFD45FB61FD56FB11" box="[674,765,1156,1182]" pageId="15" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
(see below) it is 1.3 times longer (
<figureCitation id="E1B82A7BFFA6FFDFFB72FB66FAB5FB12" box="[1173,1310,1155,1181]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="15" pageNumber="162">Figs 30–31</figureCitation>
). There are also slight differences in the genitalia, with a comparatively narrower penis valve and more clearly separated gonocoxa (
<figureCitation id="E1B82A7BFFA6FFDFFE52FB2FFD97FB6A" box="[437,572,1226,1253]" captionStart="Figs 28–35. 28, 30, 32, 34" captionStartId="14.[189,241,1927,1953]" captionTargetBox="[253,1334,264,1890]" captionTargetId="figure-76@14.[253,1334,264,1890]" captionTargetPageId="14" captionText="Figs 28–35. 28, 30, 32, 34. Andrena benoisti Wood &amp; Praz sp. nov. (TJWC). 28. Male profile. 30. Male antennae. 32. Male terga. 34. Male genitalia. — 29, 31, 33, 35. Andrena beaumonti stat. rev. (TJWC). 29. Male profile. 31. Male antennae. 33. Male terga. 35. Male genitalia. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5101654" httpUri="https://zenodo.org/record/5101654/files/figure.png" pageId="15" pageNumber="162">Figs 34–35</figureCitation>
) in
<taxonomicName id="BE834D7DFFA6FFDFFD8EFB2FFCAEFB6B" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[617,773,1226,1252]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFD8EFB2FFCAEFB6B" box="[617,773,1226,1252]" italics="true" pageId="15" pageNumber="162">A. beaumonti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="31996575FFA6FFDFFF5AFAEEFD6BF9A6" pageId="15" pageNumber="162" type="distribution">
<paragraph id="793C36FEFFA6FFDFFF5AFAEEFEFDFAAA" blockId="15.[189,342,1291,1317]" box="[189,342,1291,1317]" pageId="15" pageNumber="162">
<heading id="22748192FFA6FFDFFF5AFAEEFEFDFAAA" bold="true" box="[189,342,1291,1317]" fontSize="11" level="3" pageId="15" pageNumber="162" reason="3">
<emphasis id="4BF7EAECFFA6FFDFFF5AFAEEFEFDFAAA" bold="true" box="[189,342,1291,1317]" pageId="15" pageNumber="162">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="793C36FEFFA6FFDFFF5AFADEFD6BF9A6" blockId="15.[188,1399,1338,1577]" pageId="15" pageNumber="162">
<collectingCountry id="0194766EFFA6FFDFFF5AFADEFE81FADA" box="[189,298,1339,1365]" name="Morocco" pageId="15" pageNumber="162">Morocco</collectingCountry>
, from the High Atlas Mountains in the area south of Marrakesh. The village of Asni is found at a moderate elevation of
<quantity id="BE7B9B1BFFA6FFDFFE14FABBFDE5FAF7" box="[499,590,1374,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="162" unit="m" value="1200.0">1200 m</quantity>
. It is not clear exactly where Arroud is because of changes in spelling practices, but it probably refers to the village Aroumd some
<quantity id="BE7B9B1BFFA6FFDFFC95FA64FC11FA14" box="[882,954,1409,1436]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="162" unit="km" value="25.0">25 km</quantity>
SSE of Asni. Contemporary sampling locations of
<taxonomicName id="BE834D7DFFA6FFDFFEB7FA40FE46FA31" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[336,493,1444,1471]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="beaumonti" status="stat. nov.">
<emphasis id="4BF7EAECFFA6FFDFFEB7FA40FE46FA31" box="[336,493,1444,1471]" italics="true" pageId="15" pageNumber="162">A. beaumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50C45797FFA6FFDFFE12FA40FDF3FA30" box="[501,600,1445,1471]" pageId="15" pageNumber="162">stat. rev.</taxonomicNameLabel>
are relatively close to Asni itself. Tizi-n-Test is some
<quantity id="BE7B9B1BFFA6FFDFFB3FFA41FA89FA31" box="[1240,1314,1444,1470]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="162" unit="km" value="60.0">60 km</quantity>
SW of Asni, and Oukaimeden is even closer, just
<quantity id="BE7B9B1BFFA6FFDFFD48FA2DFD52FA6E" box="[687,761,1479,1506]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="162" unit="km" value="10.0">10 km</quantity>
to the SE. The site of Oukaimeden is at a much higher elevation of
<quantity id="BE7B9B1BFFA6FFDFFEB7FA0EFE00F98A" box="[336,427,1515,1541]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="162" unit="m" value="3200.0">3200 m</quantity>
, and the cluster of records in this region suggests that
<taxonomicName id="BE834D7DFFA6FFDFFBCBFA0EFB63F98A" authorityName="Wood &amp; Ghisbain &amp; Michez &amp; Praz" authorityYear="2021" baseAuthorityName="Benoist" baseAuthorityYear="1961" box="[1068,1224,1515,1541]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="162" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="4BF7EAECFFA6FFDFFBCBFA0EFB63F98A" box="[1068,1224,1515,1541]" italics="true" pageId="15" pageNumber="162">A. beaumonti</emphasis>
</taxonomicName>
is restricted to high-elevation sites flying in May and June.
</paragraph>
</subSubSection>
</treatment>
</document>
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