treatments-xml/data/EA/4B/BD/EA4BBD773A6155598CA183393CFC82E5.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="EA4BBD773A6155598CA183393CFC82E5" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina hudsoni Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="95FBA44C-42AA-4F8F-8919-244FD403A912" docUuidSource="ZooBank" docVersion="5" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1668149974149" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Surbakti, Suriani</mods:namePart>
<mods:affiliation>Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sumoked, Bob</mods:namePart>
<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1026</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>67</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180680991" LSID="urn:lsid:zoobank.org:act:95FBA44C-42AA-4F8F-8919-244FD403A912" httpUri="http://treatment.plazi.org/id/EA4BBD773A6155598CA183393CFC82E5" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/95FBA44C-42AA-4F8F-8919-244FD403A912" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina hudsoni" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="hudsoni" status="sp. nov.">Exocelina hudsoni Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">Figures 9</figureCitation>
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">, 10</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138336" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" elevation="1880" location="Cyclops Mts" municipality="Jayapura Regency" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
, 
<collectingMunicipality>Jayapura Regency</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:EA4BBD773A6155598CA183393CFC82E5:DAC920CDE626B54E2BA2DAC0823B60D4" country="Indonesia" municipality="Jayapura Regency" name="Cyclops Mts" stateProvince="Papua Barat">Cyclops Mts</location>
, 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.88" unit="m" value="1880.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.88" unit="m" value="1880.0">1880 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3062138318" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB" country="Indonesia" elevation="1880" location="Cyclops Mountains" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:EA4BBD773A6155598CA183393CFC82E5:9B2FB605BAF0BE4C446044D5029D9B92" country="Indonesia" name="Cyclops Mountains" stateProvince="Papua Barat">Cyclops Mountains</location>
, below summit, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.88" unit="m" value="1880.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.88" unit="m" value="1880.0">1880m</elevation>
</quantity>
, ii.201, Sentani Naturalist Club (Pap70)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138340" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount type="female">7 females</specimenCount>
with the same label as the 
<typeStatus>holotype</typeStatus>
(
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
, 
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small to medium-sized: TL-H 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.75" metricValueMin="3.4" unit="mm" value="3.575" valueMax="3.75" valueMin="3.4">3.4-3.75 mm</quantity>
, TL 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.975" metricValueMax="4.2" metricValueMin="3.75" unit="mm" value="3.975" valueMax="4.2" valueMin="3.75">3.75-4.2 mm</quantity>
, MW 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.9249999999999998" metricValueMax="2.05" metricValueMin="1.8" unit="mm" value="1.9249999999999998" valueMax="2.05" valueMin="1.8">1.8-2.05 mm</quantity>
(
<typeStatus>holotype</typeStatus>
: TL-H 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.4" unit="mm" value="3.4">3.4 mm</quantity>
, TL 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
, 
<collectionCode country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.8" unit="mm" value="1.8">1.8 mm</quantity>
), with oblong-oval habitus (Fig. 
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">9</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally piceous, with paler head and pronotum (Fig. 
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">9</figureCitation>
). Head piceous in posterior half and dark brown in anterior half; pronotum piceous on disc, sometimes narrowly, and brown to dark brown on sides, yellowish red to reddish brown at anterior angles; elytra piceous, with brown to dark brown sutural lines; head appendages and proximal part of legs yellowish brown, legs distally reddish brown.
</paragraph>
<caption doi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45" start="Figures 9, 10" startId="F4">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figures 9, 10.</emphasis>
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina hudsoni" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="hudsoni">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina hudsoni</emphasis>
</taxonomicName>
sp. nov., 
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">9</emphasis>
habitus and colouration 
<specimenCount type="male">
<emphasis bold="true" pageId="0" pageNumber="45">10</emphasis>
male
</specimenCount>
structures 
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
right protarsomeres 4 and 
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
median lobe in ventral view 
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
median lobe in lateral view 
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
right paramere in external view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Submatt dorsally, with inconspicuous elytral punctation and strongly impressed dorsal microreticulation. Head with sparse central punctation (spaces between punctures 1-4 times size of punctures), denser towards eyes; diameter of punctures smaller than diameter of cells of microreticulation; punctation relatively shallow. Pronotum with distinctly finer, sparser, and more evenly distributed punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Dorsal surface with strongly impressed microreticulation, microreticulation weaker on elytra and stronger on pronotum and head. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and very weak transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and almost invisible punctation.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antennomeres 4-10 slightly but distinctly enlarged (Fig. 
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">9</figureCitation>
). Pro- and mesotarsomeres 1-3 not dilated, narrow. Protarsomere 4 cylindrical, narrow, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of eleven and posterior row of five short setae (Fig. 
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">10A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with apex broad, curved downwards, and pointed at tip; in ventral view, with distinct submedian constriction, distal part narrower than proximal one, apex asymmetrical, narrowly concave (Fig. 
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">10B, C</figureCitation>
). Paramere with strong notch on dorsal side, with median notch tip sharply pointed, and subdistal part large and elongate; subdistal setae long, dense, curved at apex, and of two different 
<typeStatus>types</typeStatus>
: upper setae thinner, more hair-like and lower setae thick and flattened; proximal setae hair-like, more inconspicuous than subdistal ones (Fig. 
<figureCitation captionStart="Figures 9, 10" captionStartId="F4" captionText="Figures 9, 10. Exocelina hudsoni sp. nov., holotype 9 habitus and colouration 10 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures9-10" httpUri="https://binary.pensoft.net/fig/524448" pageId="0" pageNumber="45">10D</figureCitation>
). Abdominal ventrite 6 broadly rounded, with nine lateral striae on each side.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Antennomeres 4-10 stout. Pro- and mesotarsi not modified. Abdominal ventrite 6 without lateral striae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="affinities">
<paragraph pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="45">
The species evidently belongs to the 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. The species is very similar to 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis">
<emphasis italics="true" pageId="0" pageNumber="45">E. brahminensis</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis italics="true" pageId="0" pageNumber="45">E. apistefti</emphasis>
</taxonomicName>
sp. nov. in general structure of male genitalia, especially in the sharply pointed median notch tip of paramere, but differs from them in submatt dorsal surface due to stronger microreticulation, presence of pronotal bead, enlarged antennomeres 4-10, shape of the median lobe, and setation of the paramere.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138317" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Cyclops Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
. 
<collectingMunicipality>This</collectingMunicipality>
species is known only from the 
<typeStatus>type</typeStatus>
locality in 
<location LSID="urn:lsid:plazi:treatment:EA4BBD773A6155598CA183393CFC82E5:7573AE5A69604AA9135ECAAC8A344734" country="Indonesia" municipality="This" name="Cyclops Mountains" stateProvince="Papua Barat">Cyclops Mountains</location>
(Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">The specimens were collected from small puddles at low spot of a small ravine.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">
This species is named after Hudson Wild, a most dedicated naturalist and community worker in 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
. The name is a noun in the genitive case.
</paragraph>
</subSubSection>
</treatment>
</document>