209 lines
27 KiB
XML
209 lines
27 KiB
XML
<document id="B8CF1082941B10A33C4830EFC8878BDD" ID-DOI="10.11646/zootaxa.3647.2.5" ID-GBIF-Dataset="04304da3-4175-4127-a605-ec7e6637c927" ID-ISSN="1175-5326" ID-Zenodo-Dep="216203" ID-ZooBank="43984C9D-C80D-4D1C-9737-4CB8879640C9" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460605199099" checkinUser="plazi" docAuthor="Giupponi, Alessandro P. L. & Kury, Adriano B." docDate="2013" docId="D84C307CFFE9FF86FF2516B9F6237271" docLanguage="en" docName="zt03647p342.pdf" docOrigin="Zootaxa 3647 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterophrynus boterorum Giupponi & Kury, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="334" masterDocId="24754804FFEDFF83FFB2105CF34A751E" masterDocTitle="Two new species of Heterophrynus Pocock, 1894 from Colombia with distribution notes and a new synonymy (Arachnida: Amblypygi: Phrynidae)" masterLastPageNumber="342" masterPageNumber="329" pageNumber="333" updateTime="1698327867736" updateUser="plazi">
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<mods:titleInfo id="4B555071E856D303EF380E4C75ED24DE">
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<mods:title id="F74315D96EEC14C7694809FFF031E6F6">Two new species of Heterophrynus Pocock, 1894 from Colombia with distribution notes and a new synonymy (Arachnida: Amblypygi: Phrynidae)</mods:title>
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<mods:name id="3D927CD32CE22118FFF6B088F93BDCDB" type="personal">
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<mods:roleTerm id="B8304958F03551ECA63300D3ADDB2FE1">Author</mods:roleTerm>
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<mods:namePart id="E9B52FD8ACC4FD29E5E039BB9DC64270">Giupponi, Alessandro P. L.</mods:namePart>
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<mods:name id="D3A8DFCBB76A3318E2B5E72D1957BCBE" type="personal">
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<mods:roleTerm id="1DC432C4C8342C15EA1006121D523076">Author</mods:roleTerm>
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<mods:namePart id="B360D387A0BB8CBE6A4232D88E3D8A63">Kury, Adriano B.</mods:namePart>
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<mods:typeOfResource id="C0F6AFA99AC71C73FA7B05B5DCBB36DE">text</mods:typeOfResource>
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<mods:title id="3BB24789E008328EFA4839C59719E8D5">Zootaxa</mods:title>
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<mods:part id="4C61119B0DD65391F7FD9C658D7934A7">
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<mods:date id="C711E94B623661C96B38A0B17670374C">2013</mods:date>
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<mods:number id="9CE28A1A2F230117078FEFDBD37B4555">3647</mods:number>
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<mods:detail id="1025C6116227111804B0BA5DDB5DD754" type="issue">
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<mods:number id="E56DF3729514B4E22DF825D5C9ACBE7F">2</mods:number>
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<mods:classification id="8F4BF4B3D12E6EAAF0E3A3D40A10048B">journal article</mods:classification>
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<mods:identifier id="9ADAB4C8FB7FA6C27D06C5E209A54AC0" type="DOI">10.11646/zootaxa.3647.2.5</mods:identifier>
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<treatment id="D84C307CFFE9FF86FF2516B9F6237271" ID-DOI="http://doi.org/10.5281/zenodo.6156696" ID-GBIF-Taxon="119567691" ID-Zenodo-Dep="6156696" LSID="urn:lsid:plazi:treatment:D84C307CFFE9FF86FF2516B9F6237271" httpUri="http://treatment.plazi.org/id/D84C307CFFE9FF86FF2516B9F6237271" lastPageId="5" lastPageNumber="334" pageId="4" pageNumber="333">
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<subSubSection id="18FFD2E1FFE9FF87FF2516B9F17273E1" box="[151,568,1765,1791]" pageId="4" pageNumber="333" type="nomenclature">
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<paragraph id="505A816AFFE9FF87FF2516B9F17273E1" blockId="4.[151,568,1765,1791]" box="[151,568,1765,1791]" pageId="4" pageNumber="333">
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<heading id="0B123606FFE9FF87FF2516B9F17273E1" bold="true" box="[151,568,1765,1791]" fontSize="11" level="1" pageId="4" pageNumber="333" reason="1">
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<emphasis id="62915D78FFE9FF87FF2516B9F17273E1" bold="true" box="[151,568,1765,1791]" pageId="4" pageNumber="333">
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<taxonomicName id="97E5FAE9FFE9FF87FF2516B9F29E73E1" ID-CoL="3LBZG" box="[151,468,1765,1791]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="boterorum" status="sp. nov.">
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<emphasis id="62915D78FFE9FF87FF2516B9F29E73E1" bold="true" box="[151,468,1765,1791]" italics="true" pageId="4" pageNumber="333">Heterophrynus boterorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="79A2E003FFE9FF87FE6916B9F17273E1" box="[475,568,1765,1791]" pageId="4" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="18FFD2E1FFE9FF87FF251773F0FF7275" pageId="4" pageNumber="333" type="etymology">
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<paragraph id="505A816AFFE9FF87FF251773F0FF7275" blockId="4.[151,1436,1839,1972]" pageId="4" pageNumber="333">
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<emphasis id="62915D78FFE9FF87FF251773F2547259" bold="true" box="[151,286,1839,1863]" pageId="4" pageNumber="333">Etymology.</emphasis>
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Species name is a tribute to our Colombian friends and colleagues Ricardo Botero Trujillo and Juan Pablo Botero, who aided in many ways in the discovery of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="18FFD2E1FFE9FF87FF75172BF77F72AD" pageId="4" pageNumber="333" type="materials_examined">
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<paragraph id="505A816AFFE9FF87FF75172BF77F72AD" blockId="4.[151,1436,1839,1972]" pageId="4" pageNumber="333">
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<emphasis id="62915D78FFE9FF87FF75172BF23E7291" bold="true" box="[199,372,1910,1935]" pageId="4" pageNumber="333">
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<typeStatus id="8F5E3FC8FFE9FF87FF75172BF2487291" box="[199,258,1911,1935]" pageId="4" pageNumber="333">Type</typeStatus>
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material.
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</emphasis>
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1 3
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<typeStatus id="8F5E3FC8FFE9FF87FE18172BF1467291" box="[426,524,1911,1935]" pageId="4" pageNumber="333" type="holotype">holotype</typeStatus>
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and 1 3
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<typeStatus id="8F5E3FC8FFE9FF87FDC31724F19C728E" box="[625,726,1912,1936]" pageId="4" pageNumber="333" type="paratype">paratype</typeStatus>
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:
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<collectingCountry id="28F2C1FAFFE9FF87FD52172BF0307291" box="[736,890,1911,1935]" name="Colombia" pageId="4" pageNumber="333">COLOMBIA</collectingCountry>
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: Tolima, Ibague,
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<date id="245BA7AAFFE9FF87FB8B172BF7E87291" box="[1081,1186,1911,1935]" pageId="4" pageNumber="333" value="2006-08">viii-2006</date>
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, Col., A. Giupponi, R. Botero e J. P. Botero (ICN). Other
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<typeStatus id="8F5E3FC8FFE9FF87FDA917C0F1CC72AA" box="[539,646,1948,1972]" pageId="4" pageNumber="333" type="paratype">paratypes</typeStatus>
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same data 1 3 and 1 Ƥ MNRJ 0 9124.
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</paragraph>
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</subSubSection>
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<subSubSection id="18FFD2E1FFE8FF86FF7510CBF0A07402" pageId="5" pageNumber="334" type="diagnosis">
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<paragraph id="505A816AFFE8FF86FF7510CBF0A07402" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
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<emphasis id="62915D78FFE8FF86FF7510CBF20B75AE" bold="true" box="[199,321,151,176]" pageId="5" pageNumber="334">Diagnosis.</emphasis>
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Tibia I with 35 pseudoarticles and tarsus I with 79-80 pseudoarticles. Male gonopod: tegument of
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<emphasis id="62915D78FFE8FF86FF2510E0F2EE75CA" box="[151,420,188,212]" italics="true" pageId="5" pageNumber="334">lobus lateralis secundus</emphasis>
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with densely clustered small acuminate denticles and length almost equal to
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<emphasis id="62915D78FFE8FF86FB4E10E0F3AF75E6" italics="true" pageId="5" pageNumber="334">lobus lateralis primus</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FF4A1083F20075E9" box="[248,330,223,247]" captionStart="FIGURE 5" captionStartId="7.[151,250,1864,1887]" captionTargetBox="[151,1435,387,1838]" captionTargetId="figure@7.[151,1436,387,1839]" captionTargetPageId="7" captionText="FIGURE 5. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype, genitalia and surroundings: a. Male gonopod, general ventral view; b. Same, detail of left and right lobes; c. Same, detail of left distal part of lateral lobe (LoL 2), spinytongue-like with only one layer covered with denticles; d. Same, detail of basal part of Lol 2; e. Male gonopod, general dorsal view; f. Same, detail of right dorsal and lateral lobes." httpUri="https://zenodo.org/record/216208/files/figure.png" pageId="5" pageNumber="334">Figs. 5</figureCitation>
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b–c). Female gonopod: sclerotized claw bifid. Distalmost (third) inner tooth of basal article of chelicera bifid. Small body size for the genus (Total body length: 23.0 mm).
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</paragraph>
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</subSubSection>
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<subSubSection id="18FFD2E1FFE8FF86FF75117BF6237271" pageId="5" pageNumber="334" type="description">
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<paragraph id="505A816AFFE8FF86FF75117BF2C77709" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
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<emphasis id="62915D78FFE8FF86FF75117BF104745E" bold="true" box="[199,590,295,320]" pageId="5" pageNumber="334">
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Description of female (
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<typeStatus id="8F5E3FC8FFE8FF86FE6F117BF10D745E" box="[477,583,295,320]" pageId="5" pageNumber="334" type="holotype">holotype</typeStatus>
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)
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</emphasis>
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:
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<emphasis id="62915D78FFE8FF86FDD2117BF199745E" bold="true" box="[608,723,295,320]" pageId="5" pageNumber="334">Carapace</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FD57117BF0667421" box="[741,812,295,319]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Fig. 4</figureCitation>
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a): flattened, wider than long; prominent eyes, median eye tubercle dark-brown, without apical setae, lateral eye tubercle smaller, with around half its size. Margin of carapace entirely with sinuous lines and coarse granules, anterior margin in a very smooth, practically straight curve. Overall densely covered by small granules mostly anteriorly, bundles of bigger and more sparse granules departing from the fovea towards the border. Frontal process well-developed, with greatly widened base and abruptly getting thin in distal half, ending in a very acute hook, not exceeding front line of carapace, this way not visible in dorsal view.
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</paragraph>
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<paragraph id="505A816AFFE8FF86FF75127FF60177B9" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
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<emphasis id="62915D78FFE8FF86FF75127FF2647722" bold="true" box="[199,302,547,572]" pageId="5" pageNumber="334">Sternum</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FEF01278F2C17722" box="[322,395,548,572]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Fig. 4</figureCitation>
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b): Tri-segmented, typical of
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<taxonomicName id="97E5FAE9FFE8FF86FD571278F0107722" box="[741,858,548,572]" class="Arachnida" family="Phrynidae" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="334" phylum="Arthropoda" rank="family">Phrynidae</taxonomicName>
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. Tritosternum extremely long projecting between pedipalp coxae, reaching almost half of their length. Covered with bristles on its entire length, from apex to base, with an increase in the density at base. Mesosternum convex, reduced to a small tubercle, ranging from slightly oblong to sharply oblong, with six to eight bristles. Metasternum is a small convex mound with two bristles
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</paragraph>
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<paragraph id="505A816AFFE8FF86FF7512EFF61277D2" blockId="5.[151,1436,151,1904]" box="[199,1368,691,716]" pageId="5" pageNumber="334">
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<emphasis id="62915D78FFE8FF86FF7512EFF27677D2" bold="true" box="[199,316,691,716]" pageId="5" pageNumber="334">Abdomen</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FEF912E8F2D877D2" box="[331,402,692,716]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Figs.4</figureCitation>
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a–b): oblong, with almost indistinguishable punctuations, thinner than in the carapace.
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</paragraph>
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<paragraph id="505A816AFFE8FF86FF75128BF0217642" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
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<emphasis id="62915D78FFE8FF86FF75128BF27077EE" bold="true" box="[199,314,727,752]" pageId="5" pageNumber="334">Chelicera</emphasis>
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: Cheliceral furrow (
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<figureCitation id="C8DE9DEFFFE8FF86FD90128BF12577F1" box="[546,623,727,751]" captionStart="FIGURE 6" captionStartId="8.[151,250,1599,1622]" captionTargetBox="[241,1345,463,1562]" captionTargetId="figure@8.[241,1346,463,1563]" captionTargetPageId="8" captionText="FIGURE 6. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Right chelicera, inner lateral view; b. Same, outer lateral view; c. Leg IV, basitibia and distitibia showing trichobothria, dorsal view; d. Female gonopod, general dorsal view. Scale bars = 1 mm (Figs. a – b); 5 mm (Fig. c); 0.5 mm (Fig. d)." httpUri="https://zenodo.org/record/216210/files/figure.png" pageId="5" pageNumber="334">Figs. 6</figureCitation>
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a–b) with 3 internal teeth, the distal one bifid, the first cusp bigger than the second one II<I=IIIa>IIIb. The third tooth (middle) much smaller than the others. With 3 external teeth, the distal one bifid, the first cusp bigger than the second one II>I=IIIa>IIIb. Claw with five denticles, with the two distalmost much reduced, the second smaller than first and third.
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</paragraph>
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<paragraph id="505A816AFFE8FF86FF75133BF13B7031" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
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<emphasis id="62915D78FFE8FF86FF75133BF27B769E" bold="true" box="[199,305,871,896]" pageId="5" pageNumber="334">Pedipalp</emphasis>
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:
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<emphasis id="62915D78FFE8FF86FEF21334F28E769E" bold="true" box="[320,452,872,896]" pageId="5" pageNumber="334">Trochanter</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FE65133BF16C7661" box="[471,550,871,895]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Figs. 4</figureCitation>
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c–d): with well-developed retro-ventral apophysis, armed with four spines, the one adjoining the retro-ventral apophysis is the largest, but not more than twice the size of others, which are subequal. The biggest spine is of comparable size to the fourth ventral spine of pedipalpal femur.
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<emphasis id="62915D78FFE8FF86FB6713ECF66C76D6" bold="true" box="[1237,1318,944,968]" pageId="5" pageNumber="334">Femur</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FA8613F3F6CB76D9" box="[1332,1409,943,967]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Figs. 4</figureCitation>
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c– d): three dorsal spines in the main series (1>2>3), the first one three times bigger than the last one; prominent geminate spines before the first spine, the dorsalmost subequal to spine 2 and the other subequal to spine 3; four ventral spines (1>2>3>4) of approximately the same length of the dorsal ones; one spine placed proximally to the first one, which is located more interiorly than the main series of spines, with a similar length of the third one, but less robust; last spine ½ the length of the first one.
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<emphasis id="62915D78FFE8FF86FD42143FF0657162" bold="true" box="[752,815,1123,1148]" pageId="5" pageNumber="334">Tibia</emphasis>
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(
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<figureCitation id="C8DE9DEFFFE8FF86FCF01438F0D87162" box="[834,914,1124,1148]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Figs. 4</figureCitation>
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c–d): six spines, main series with four spines (I=II=III>IV); two accessory spine anterior to spine I, with its length similar to spine IV; five ventral spines (III>IV>II>I>V),
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<emphasis id="62915D78FFE8FF86FED814F7F2AD71DA" bold="true" box="[362,487,1195,1220]" pageId="5" pageNumber="334">Basitarsus</emphasis>
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(
|
||
<figureCitation id="C8DE9DEFFFE8FF86FDB214F0F11271DA" box="[512,600,1196,1220]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Figs. 4</figureCitation>
|
||
c–d): two dorsal spines and two ventral spines dorsal distal>dorsal basal=ventral distal>ventral basal. The distal dorsal spine subequal to the claw.
|
||
<emphasis id="62915D78FFE8FF86FB971493F72E71F6" bold="true" box="[1061,1124,1231,1256]" pageId="5" pageNumber="334">Claw</emphasis>
|
||
(
|
||
<figureCitation id="C8DE9DEFFFE8FF86FBC61493F78F71F9" box="[1140,1221,1231,1255]" captionStart="FIGURE 4" captionStartId="6.[151,250,1619,1642]" captionTargetBox="[151,1435,195,1598]" captionTargetId="figure@6.[151,1436,195,1598]" captionTargetPageId="6" captionText="FIGURE 4. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Habitus, dorsal view; b. Same, ventral view; c. Right pedipalp, dorsal view; d. Right pedipalp, ventral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/216207/files/figure.png" pageId="5" pageNumber="334">Figs. 4</figureCitation>
|
||
c–d): long, with an acute, curved tip. The males do not exhibit considerable secondary sexual dimorphic features, and the pedipalps of males and females are of comparable size.
|
||
</paragraph>
|
||
<paragraph id="505A816AFFE8FF86FF751560F15470FA" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
|
||
<emphasis id="62915D78FFE8FF86FF751560F242704A" bold="true" box="[199,264,1340,1364]" pageId="5" pageNumber="334">Legs:</emphasis>
|
||
All densely setose. Ventral corner of the prolateral face of femora II–IV projecting in a distinct spiniform process.
|
||
<emphasis id="62915D78FFE8FF86FF4B153CF2D77066" bold="true" box="[249,413,1376,1400]" pageId="5" pageNumber="334">Femur length</emphasis>
|
||
I>III>IV>II. Tibia I with 35 pseudoarticles and tarsus (basitarsus+distitarsus) I with 79-80 pseudoarticles in both legs.
|
||
<emphasis id="62915D78FFE8FF86FE7E15D8F16F7082" bold="true" box="[460,549,1412,1436]" pageId="5" pageNumber="334">Leg IV:</emphasis>
|
||
(
|
||
<figureCitation id="C8DE9DEFFFE8FF86FD8715D8F1327082" box="[565,632,1412,1436]" captionStart="FIGURE 6" captionStartId="8.[151,250,1599,1622]" captionTargetBox="[241,1345,463,1562]" captionTargetId="figure@8.[241,1346,463,1563]" captionTargetPageId="8" captionText="FIGURE 6. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Right chelicera, inner lateral view; b. Same, outer lateral view; c. Leg IV, basitibia and distitibia showing trichobothria, dorsal view; d. Female gonopod, general dorsal view. Scale bars = 1 mm (Figs. a – b); 5 mm (Fig. c); 0.5 mm (Fig. d)." httpUri="https://zenodo.org/record/216210/files/figure.png" pageId="5" pageNumber="334">Fig. 6</figureCitation>
|
||
c):
|
||
<emphasis id="62915D78FFE8FF86FD2815DFF0477082" bold="true" box="[666,781,1411,1436]" pageId="5" pageNumber="334">Basitibia:</emphasis>
|
||
4 pseudoarticles, one trichobothrium at the first and one at the last pseudoarticle.
|
||
<emphasis id="62915D78FFE8FF86FE0415FBF16770DE" bold="true" box="[438,557,1447,1472]" pageId="5" pageNumber="334">Distitibia:</emphasis>
|
||
16 trichobothria.
|
||
<emphasis id="62915D78FFE8FF86FCA915FBF71C70DE" bold="true" box="[795,1110,1447,1472]" pageId="5" pageNumber="334">Basitibia-distitibia length</emphasis>
|
||
BT1>DT>BT3=BT4>BT2.
|
||
<emphasis id="62915D78FFE8FF86FF251597F26B70FA" bold="true" box="[151,289,1483,1508]" pageId="5" pageNumber="334">Distitarsus:</emphasis>
|
||
with 3 pseudoarticles.
|
||
</paragraph>
|
||
<paragraph id="505A816AFFE8FF86FF7515ACF01B736A" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
|
||
<emphasis id="62915D78FFE8FF86FF7515ACF23C7316" bold="true" box="[199,374,1520,1544]" pageId="5" pageNumber="334">Measurements</emphasis>
|
||
.
|
||
<emphasis id="62915D78FFE8FF86FE3415ACF2A27316" bold="true" box="[390,488,1520,1544]" pageId="5" pageNumber="334">Females</emphasis>
|
||
(n=1): Total body length: 23.0, Cephalothorax: Length: 8.0 mm, Width: 10,5 mm. Abdomen: 13.0 mm. Pedipalp: Femur 9.0 mm, Tibia 10.0 mm, Tarsus 5.0 mm, Tarsal claw
|
||
<quantity id="971D2C8FFFE8FF86FB131648F7B17332" box="[1185,1275,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="334" unit="mm" value="3.5">3.5 mm</quantity>
|
||
.
|
||
<emphasis id="62915D78FFE8FF86FAB41648F6047332" bold="true" box="[1286,1358,1556,1580]" pageId="5" pageNumber="334">Males</emphasis>
|
||
(n=1): Total body length: 24.2, Cephalothorax: Length: 10.0 mm, Width: 11,0 mm. Abdomen: 17.0 mm. Pedipalp: Femur 11.0 mm, Tibia
|
||
<quantity id="971D2C8FFFE8FF86FEFA1600F2E4736A" box="[328,430,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="334" unit="mm" value="14.5">14.5 mm</quantity>
|
||
, Tarsus
|
||
<quantity id="971D2C8FFFE8FF86FDB81600F12B736A" box="[522,609,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="334" unit="mm" value="5.1">5.1 mm</quantity>
|
||
, Tarsal claw 4.0 mm.
|
||
</paragraph>
|
||
<paragraph id="505A816AFFE8FF86FF751623F28473A2" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
|
||
<emphasis id="62915D78FFE8FF86FF751623F2217386" bold="true" box="[199,363,1663,1688]" pageId="5" pageNumber="334">Color pattern</emphasis>
|
||
(in alcohol): Chelicerae, pedipalps, legs and carapace reddish-brown. Coxae and ventral surface of abdomen lighter colored.
|
||
</paragraph>
|
||
<paragraph id="505A816AFFE8FF86FF75169BF7827239" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
|
||
<emphasis id="62915D78FFE8FF86FF75169BF27F73FE" bold="true" box="[199,309,1735,1760]" pageId="5" pageNumber="334">Genitalia</emphasis>
|
||
:
|
||
<emphasis id="62915D78FFE8FF86FEF71694F2CB73FE" bold="true" box="[325,385,1736,1760]" pageId="5" pageNumber="334">Male</emphasis>
|
||
(
|
||
<figureCitation id="C8DE9DEFFFE8FF86FE21169BF2AB73C1" box="[403,481,1735,1759]" captionStart="FIGURE 5" captionStartId="7.[151,250,1864,1887]" captionTargetBox="[151,1435,387,1838]" captionTargetId="figure@7.[151,1436,387,1839]" captionTargetPageId="7" captionText="FIGURE 5. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype, genitalia and surroundings: a. Male gonopod, general ventral view; b. Same, detail of left and right lobes; c. Same, detail of left distal part of lateral lobe (LoL 2), spinytongue-like with only one layer covered with denticles; d. Same, detail of basal part of Lol 2; e. Male gonopod, general dorsal view; f. Same, detail of right dorsal and lateral lobes." httpUri="https://zenodo.org/record/216208/files/figure.png" pageId="5" pageNumber="334">Figs. 5</figureCitation>
|
||
a–f). PI longer than LaM. LoL1 and LoL2 subequal in length, both densely covered with minute denticles (but larger than in
|
||
<taxonomicName id="97E5FAE9FFE8FF86FDD416B1F198721A" box="[614,722,1772,1796]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="silviae">
|
||
<emphasis id="62915D78FFE8FF86FDD416B1F198721A" box="[614,722,1772,1796]" italics="true" pageId="5" pageNumber="334">H. silviae</emphasis>
|
||
</taxonomicName>
|
||
). In dorsal view the LoD is subequal in size to LoL1.
|
||
<emphasis id="62915D78FFE8FF86FA8C16B0F6D6721A" bold="true" box="[1342,1436,1772,1796]" pageId="5" pageNumber="334">Female.</emphasis>
|
||
(
|
||
<figureCitation id="C8DE9DEFFFE8FF86FF2D1753F3A87239" box="[159,226,1807,1831]" captionStart="FIGURE 6" captionStartId="8.[151,250,1599,1622]" captionTargetBox="[241,1345,463,1562]" captionTargetId="figure@8.[241,1346,463,1563]" captionTargetPageId="8" captionText="FIGURE 6. Heterophrynus boterorum sp. nov., male (MNRJ 09124) paratype: a. Right chelicera, inner lateral view; b. Same, outer lateral view; c. Leg IV, basitibia and distitibia showing trichobothria, dorsal view; d. Female gonopod, general dorsal view. Scale bars = 1 mm (Figs. a – b); 5 mm (Fig. c); 0.5 mm (Fig. d)." httpUri="https://zenodo.org/record/216210/files/figure.png" pageId="5" pageNumber="334">Fig. 6</figureCitation>
|
||
d) Gonopod claws bifid. Inner branch probably arising from basal third of claw complex.
|
||
</paragraph>
|
||
<paragraph id="505A816AFFE8FF86FF751768F6237271" blockId="5.[151,1436,151,1904]" pageId="5" pageNumber="334">
|
||
<emphasis id="62915D78FFE8FF86FF751768F2C37252" bold="true" box="[199,393,1843,1868]" pageId="5" pageNumber="334">Natural history.</emphasis>
|
||
The specimens described were collected on rocks and ravines in a fragment of Andean montane secondary forest in a very disturbed area surrounded by coffee plantations and rural residential areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |