120 lines
14 KiB
XML
120 lines
14 KiB
XML
<document id="E4D02AD9A284D3D7AD8A5C6D608DE682" ID-CLB-Dataset="30423" ID-DOI="10.11646/zootaxa.4399.2.7" ID-GBIF-Dataset="6c5127b3-a849-4ea8-a965-8e915f63a551" ID-ISSN="1175-5326" ID-Zenodo-Dep="1207158" ID-ZooBank="073E1E48-50BE-42DE-B21A-FA9043786E23" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521995446897" checkinUser="plazi" docAuthor="Couri, Márcia S. & Pont, Adrian C." docDate="2018" docId="CD02BC11FFD6E565FF5CD47AFB2157B4" docLanguage="en" docName="zootaxa.4399.2.7.pdf.imf" docOrigin="Zootaxa 4399 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina naivashensis Embden 1951, stat. nov." docType="treatment" docVersion="7" lastPageNumber="240" masterDocId="313BC469FFD1E562FFCBD567FFB0542C" masterDocTitle="The male terminalia of some African species of Helina Robineau-Desvoidy, 1830 (Diptera, Muscidae)" masterLastPageNumber="247" masterPageNumber="233" pageNumber="240" updateTime="1698533436902" updateUser="ExternalLinkService">
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<mods:titleInfo id="10A8FA153165503C0FDBECC5095C6368">
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<mods:title id="CE1C75677DEFA29AD56DD18DCF761FE1">The male terminalia of some African species of Helina Robineau-Desvoidy, 1830 (Diptera, Muscidae)</mods:title>
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<mods:namePart id="14FD3D5BFF66A0C1A5F58CE58511C490">Couri, Márcia S.</mods:namePart>
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<mods:namePart id="825F3026A8A71E345C164CFAAC3490CF">Pont, Adrian C.</mods:namePart>
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<mods:typeOfResource id="0191278C8D5C20845FE33157F3EAF098">text</mods:typeOfResource>
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<mods:title id="CCD796827047CE1D49FEFB40E2CFC69B">Zootaxa</mods:title>
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<mods:part id="C51CC67C9FD5DCF0E1671E5F6A035825">
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<mods:date id="509944D19C10477B3BB9E32D97A11AD0">2018</mods:date>
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<mods:number id="EAD7BD8CC7099F53528D54BDD1CF58CF">2018-03-21</mods:number>
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<mods:number id="EE1E03EACA42EDF3AFADDBA6F967397B">4399</mods:number>
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<mods:number id="3520D9EAD961394F886A9D3ECC320539">2</mods:number>
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<mods:identifier id="D7276C34AB587233D0D9D241213B9665" type="DOI">10.11646/zootaxa.4399.2.7</mods:identifier>
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<treatment id="CD02BC11FFD6E565FF5CD47AFB2157B4" ID-DOI="http://doi.org/10.5281/zenodo.6489499" ID-GBIF-Taxon="142611014" ID-Zenodo-Dep="6489499" LSID="urn:lsid:plazi:treatment:CD02BC11FFD6E565FF5CD47AFB2157B4" httpUri="http://treatment.plazi.org/id/CD02BC11FFD6E565FF5CD47AFB2157B4" lastPageNumber="240" pageId="7" pageNumber="240">
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<subSubSection id="0DB15E8CFFD6E565FF5CD47AFD04551B" box="[151,692,285,311]" pageId="7" pageNumber="240" type="nomenclature">
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<paragraph id="45140D07FFD6E565FF5CD47AFD04551B" blockId="7.[151,692,285,344]" box="[151,692,285,311]" pageId="7" pageNumber="240">
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<heading id="1E5CBA6BFFD6E565FF5CD47AFD04551B" bold="true" box="[151,692,285,311]" fontSize="11" level="1" pageId="7" pageNumber="240" reason="1">
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<emphasis id="77DFD115FFD6E565FF5CD47AFD04551B" bold="true" box="[151,692,285,311]" pageId="7" pageNumber="240">
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<taxonomicName id="82AB7684FFD6E565FF5CD47AFD8D551B" ID-CoL="6LJV5" authority="Embden, 1951" authorityName="Embden" authorityYear="1951" box="[151,573,285,311]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="240" phylum="Arthropoda" rank="species" species="naivashensis" status="stat. nov.">
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<emphasis id="77DFD115FFD6E565FF5CD47AFE3C551B" bold="true" box="[151,396,285,311]" italics="true" pageId="7" pageNumber="240">Helina naivashensis</emphasis>
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<bibRefCitation id="213A70F6FFD6E565FE58D47AFD87551B" box="[403,567,285,311]" pageId="7" pageNumber="240" refId="ref4484" refString="Emden, F. I. van (1951) Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoninae. In: Ruwenzori ExPedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 325 - 710." type="book chapter">Emden, 1951</bibRefCitation>
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,
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</taxonomicName>
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<taxonomicNameLabel id="6CEC6C6EFFD6E565FD8FD47AFD04551B" box="[580,692,285,311]" pageId="7" pageNumber="240">stat. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DB15E8CFFD6E565FF5CD458FE975574" box="[151,295,319,344]" pageId="7" pageNumber="240" type="description">
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<paragraph id="45140D07FFD6E565FF5CD458FE975574" blockId="7.[151,692,285,344]" box="[151,295,319,344]" pageId="7" pageNumber="240">
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(
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<figureCitation id="DD901182FFD6E565FF54D458FEAF5574" box="[159,287,319,344]" captionStart="FIGURES 28–34" captionStartId="9.[151,264,1908,1931]" captionTargetBox="[185,1375,203,1864]" captionTargetId="figure@9.[185,1409,193,1901]" captionTargetPageId="9" captionText="FIGURES 28–34. Helina mollis (Stein, 1906), male: (28) sternite 5, dorsal view; (29) cercal plate and surstylus, dorsal view; (30) cercal plate and surstylus, lateral view; (31) aedeagal complex, lateral view. Helina naivashensis Emden, 1951, stat. nov., paratype male:(32) sternite 5, dorsalview; (33) cercal plate, surstylus and aedeagal complex, lateral view; (34) cercal plate and surstylus, dorsal view; Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207170/files/figure.png" pageId="7" pageNumber="240">Figs 32–34</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0DB15E8CFFD6E565FF5CD4E0FE1C55E9" pageId="7" pageNumber="240" type="materials_examined">
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<paragraph id="45140D07FFD6E565FF5CD4E0FE1C55E9" blockId="7.[151,1437,391,920]" pageId="7" pageNumber="240">
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<emphasis id="77DFD115FFD6E565FF5CD4E0FE73558C" bold="true" box="[151,451,391,416]" pageId="7" pageNumber="240">
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Examined
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<typeStatus id="9A10B3A5FFD6E565FED2D4E0FEFC558C" box="[281,332,391,416]" pageId="7" pageNumber="240">type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="9A10B3A5FFD6E565FE00D4E0FD83558C" box="[459,563,391,416]" pageId="7" pageNumber="240" type="holotype">Holotype</typeStatus>
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male seen; right wing damaged at tip. One
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<typeStatus id="9A10B3A5FFD6E565FBE3D4EFFB38558C" box="[1064,1160,392,416]" pageId="7" pageNumber="240" type="paratype">paratype</typeStatus>
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from
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<collectingCountry id="3DBC4D97FFD6E565FB1BD4EFFAA9558C" box="[1232,1305,392,416]" name="Kenya" pageId="7" pageNumber="240">Kenya</collectingCountry>
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(Naivasha) dissected and illustrated.
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</paragraph>
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</subSubSection>
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<subSubSection id="0DB15E8CFFD6E565FF0CD4A8FD3E56EC" pageId="7" pageNumber="240" type="diagnosis">
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<paragraph id="45140D07FFD6E565FF0CD4A8FB43561C" blockId="7.[151,1437,391,920]" pageId="7" pageNumber="240">
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<emphasis id="77DFD115FFD6E565FF0CD4A8FEF155C4" bold="true" box="[199,321,463,488]" pageId="7" pageNumber="240">Diagnosis.</emphasis>
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Fore tarsus with claws and pulvilli very long in male, and apical tarsomere dilated at apex; hind femur with numerous very long, setulose anteroventral and posteroventral hairs; hind tibia with long anteroventral and ventral hairs, one long posterior, and long erect posterior to posteroventral hairs on apical third.
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</paragraph>
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<paragraph id="45140D07FFD6E565FF0CD75BFD3E56EC" blockId="7.[151,1437,391,920]" pageId="7" pageNumber="240">
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<emphasis id="77DFD115FFD6E565FF0CD75BFECC5679" box="[199,380,572,597]" italics="true" pageId="7" pageNumber="240">Male terminalia</emphasis>
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. Sternite 5 longer than wide, with medium-sized and long setae on disc and longer setae at tips of lobes (
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<figureCitation id="DD901182FFD6E565FECED738FEEB5654" box="[261,347,607,632]" captionStart="FIGURES 28–34" captionStartId="9.[151,264,1908,1931]" captionTargetBox="[185,1375,203,1864]" captionTargetId="figure@9.[185,1409,193,1901]" captionTargetPageId="9" captionText="FIGURES 28–34. Helina mollis (Stein, 1906), male: (28) sternite 5, dorsal view; (29) cercal plate and surstylus, dorsal view; (30) cercal plate and surstylus, lateral view; (31) aedeagal complex, lateral view. Helina naivashensis Emden, 1951, stat. nov., paratype male:(32) sternite 5, dorsalview; (33) cercal plate, surstylus and aedeagal complex, lateral view; (34) cercal plate and surstylus, dorsal view; Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207170/files/figure.png" pageId="7" pageNumber="240">Fig. 32</figureCitation>
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); cercal plate long, with 3 pairs of short stout setae at apex and two pairs a little above them; surstylus longer than cercal plate, sinuous in apical portion (
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<figureCitation id="DD901182FFD6E565FC8ED7E3FC2A56B0" box="[837,922,644,669]" captionStart="FIGURES 28–34" captionStartId="9.[151,264,1908,1931]" captionTargetBox="[185,1375,203,1864]" captionTargetId="figure@9.[185,1409,193,1901]" captionTargetPageId="9" captionText="FIGURES 28–34. Helina mollis (Stein, 1906), male: (28) sternite 5, dorsal view; (29) cercal plate and surstylus, dorsal view; (30) cercal plate and surstylus, lateral view; (31) aedeagal complex, lateral view. Helina naivashensis Emden, 1951, stat. nov., paratype male:(32) sternite 5, dorsalview; (33) cercal plate, surstylus and aedeagal complex, lateral view; (34) cercal plate and surstylus, dorsal view; Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207170/files/figure.png" pageId="7" pageNumber="240">Fig. 34</figureCitation>
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); aedeagal complex with phallapodeme long; distiphallus short and membranous (
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<figureCitation id="DD901182FFD6E565FDE6D7C0FD3156EC" box="[557,641,679,704]" captionStart="FIGURES 28–34" captionStartId="9.[151,264,1908,1931]" captionTargetBox="[185,1375,203,1864]" captionTargetId="figure@9.[185,1409,193,1901]" captionTargetPageId="9" captionText="FIGURES 28–34. Helina mollis (Stein, 1906), male: (28) sternite 5, dorsal view; (29) cercal plate and surstylus, dorsal view; (30) cercal plate and surstylus, lateral view; (31) aedeagal complex, lateral view. Helina naivashensis Emden, 1951, stat. nov., paratype male:(32) sternite 5, dorsalview; (33) cercal plate, surstylus and aedeagal complex, lateral view; (34) cercal plate and surstylus, dorsal view; Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207170/files/figure.png" pageId="7" pageNumber="240">Fig. 33</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0DB15E8CFFD6E565FF0CD7ABFB2157B4" pageId="7" pageNumber="240" type="discussion">
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<paragraph id="45140D07FFD6E565FF0CD7ABFB2157B4" blockId="7.[151,1437,391,920]" pageId="7" pageNumber="240">
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<emphasis id="77DFD115FFD6E565FF0CD7ABFEA156C9" bold="true" box="[199,273,716,741]" pageId="7" pageNumber="240">Notes.</emphasis>
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This species was originally described as a subspecies of
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<emphasis id="77DFD115FFD6E565FC12D7ABFB0356C9" box="[985,1203,716,741]" italics="true" pageId="7" pageNumber="240">Helina quadruplex</emphasis>
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(
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<bibRefCitation id="213A70F6FFD6E565FB01D7ABFAE656C8" box="[1226,1366,716,741]" pageId="7" pageNumber="240" refId="ref5056" refString="Stein, P. (1913) Neue afrikanische Anthomyiden. Annales historico-naturales Musei nationalis Hungarici, 11, 457 - 583." type="journal article">Stein, 1913</bibRefCitation>
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). The terminalia of the two subspecies were dissected and, although their external morphology is similar, they differ in the arrangement of setae on the cercal plate, the enlargement of the hypandrium in lateral view, and the length of the distiphallus. According to
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<bibRefCitation id="213A70F6FFD6E565FE3FD650FD65577C" box="[500,725,823,848]" pageId="7" pageNumber="240" refId="ref4484" refString="Emden, F. I. van (1951) Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoninae. In: Ruwenzori ExPedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 325 - 710." type="book chapter">Emden (1951: 597)</bibRefCitation>
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, the females of both taxa are indistinguishable, but the males differ considerably in the chaetotaxy of the hind leg (
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<bibRefCitation id="213A70F6FFD6E565FCEFD63BFC4E5759" box="[804,1022,860,885]" pageId="7" pageNumber="240" refId="ref4484" refString="Emden, F. I. van (1951) Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoninae. In: Ruwenzori ExPedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 325 - 710." type="book chapter">Emden 1951: 597</bibRefCitation>
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). The status of
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<emphasis id="77DFD115FFD6E565FB08D63BFE9657B4" italics="true" pageId="7" pageNumber="240">Helina quadruplex naivashensis</emphasis>
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is therefore revised and the subspecies is elevated to species rank (
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<emphasis id="77DFD115FFD6E565FBDDD618FB3157B4" bold="true" box="[1046,1153,895,920]" pageId="7" pageNumber="240">stat. nov.</emphasis>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |