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<document ID-ISSN="0003-0090" approvalRequired="248" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="67" approvalRequired_for_textStreams="120" approvalRequired_for_treatments="60" checkinTime="1630255760165" checkinUser="felipe" docAuthor="Conrad, J. L." docDate="2008" docId="BF23879DD136FFDFFF53AB8F4B60D47D" docLanguage="en" docName="B310.pdf" docOrigin="Bulletin of the American Museum of Natural History 2008 (310)" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.1:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="1" docTitle="Teiidae Gray 1827" docType="treatment" docVersion="2" lastPageNumber="102" masterDocId="431AFFE5D153FFBAFFA0AE674F56D039" masterDocTitle="Phylogeny And Systematics Of Squamata (Reptilia) Based On Morphology" masterLastPageNumber="182" masterPageNumber="1" pageNumber="102" updateTime="1630877461833" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Phylogeny And Systematics Of Squamata (Reptilia) Based On Morphology</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Conrad, J. L.</mods:namePart>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2008</mods:date>
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<mods:number>2008-06-03</mods:number>
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<mods:number>2008</mods:number>
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<mods:number>310</mods:number>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<heading box="[243,469,1512,1534]" centered="true" fontSize="9" level="2" pageId="101" pageNumber="102" reason="2">
<taxonomicName authority="Gray, 1827" authorityName="Gray" authorityYear="1827" box="[243,469,1512,1534]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">
Teiidae
<bibRefCitation author="Gray, J. E." box="[338,469,1512,1533]" pageId="101" pageNumber="102" pagination="54 - 58" refId="ref90087" refString="Gray, J. E. 1827. A synopsis of the genera of saurian reptiles, in which some new genera are indicated and the others reviewed by actual examination. Annals of Philosophy, n. 5. 2: 54 - 58." type="journal article" year="1827">Gray, 1827</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="101.[232,480,1512,1565]" box="[232,480,1542,1565]" pageId="101" pageNumber="102">
<heading box="[232,480,1542,1565]" centered="true" fontSize="10" level="2" pageId="101" pageNumber="102" reason="4">(figs. 54C, 55C, 56B)</heading>
</paragraph>
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<subSubSection pageId="101" pageNumber="102" type="description">
<paragraph blockId="101.[93,620,1587,1759]" pageId="101" pageNumber="102">
DEFINITION:
<taxonomicName authorityName="Hensel" authorityYear="1868" box="[285,422,1589,1610]" class="Reptilia" family="Teiidae" genus="Teius" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="species" species="teyou">
<emphasis box="[285,422,1589,1610]" italics="true" pageId="101" pageNumber="102">Teius teyou</emphasis>
</taxonomicName>
,
<taxonomicName class="Reptilia" family="Teiidae" genus="Tupinambis" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="species" species="teguixin">
<emphasis italics="true" pageId="101" pageNumber="102">Tupinambis teguixin</emphasis>
</taxonomicName>
, and all descendants of their last common ancestor.
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="102" type="diagnosis">
<paragraph blockId="101.[93,620,1587,1759]" lastBlockId="101.[667,1193,213,1092]" pageId="101" pageNumber="102">
DIAGNOSIS:
<taxonomicName authorityName="Gray" authorityYear="1827" box="[272,358,1678,1700]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Teiidae</taxonomicName>
share five unambiguous synapomorphies to the exclusion of polyglyphanodontids: 8(1) presence of der- mal sculpturing on the maxilla, 100(0) presence of a dorsal process on the squamosal, 240(0) well-developed atlantal lateral processes, 243(3) cervical intercentra fused to the succeeding vertebrae, and 285(1) symphysial and tubercular portions of the pubis of subequal length.
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="102" type="discussion">
<paragraph blockId="101.[667,1193,213,1092]" pageId="101" pageNumber="102">
COMMENTS: Estes et al. defined
<taxonomicName authorityName="Gray" authorityYear="1827" box="[1075,1161,420,442]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Teiidae</taxonomicName>
as the last common ancestor of the
<taxonomicName authorityName="Presch" authorityYear="1970" box="[1057,1143,450,472]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="subFamily" subFamily="Teiinae">Teiinae</taxonomicName>
and
<taxonomicName authorityName="Presch" authorityYear="1974" box="[667,837,479,501]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="subFamily" subFamily="Tupinambinae">Tupinambinae</taxonomicName>
… and all organisms sharing a more recent common ancestor with these taxa than with any other extant organisms (Estes et al., 1988: 215). Polyglyphanodontids would be considered teiids under that definition. Indeed, polyglyphanodontids have been considered part of the
<taxonomicName authorityName="Gray" authorityYear="1827" box="[957,1043,657,679]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Teiidae</taxonomicName>
in the past (
<bibRefCitation author="Estes, R." box="[675,812,686,707]" pageId="101" pageNumber="102" refId="ref87654" refString="Estes, R. 1983. Sauria terrestria, Amphisbaenia. New York: Gustav Fischer Verlag, 249 pp." type="book" year="1983">Estes, 1983</bibRefCitation>
;
<bibRefCitation author="Presch, W." box="[827,979,686,708]" pageId="101" pageNumber="102" pagination="189 - 197" refId="ref97766" refString="Presch, W. 1983. The lizard family Teiidae: is it a monophyletic group? Zoological Journal of the Linnean Society 77: 189 - 197." type="journal article" year="1983">Presch, 1983</bibRefCitation>
;
<bibRefCitation author="Gao, K. - Q. &amp; M. A. Norell" pageId="101" pageNumber="102" pagination="1 - 118" refId="ref89476" refString="Gao, K. - Q., and M. A. Norell. 2000. Taxonomic composition, and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 249: 1 - 118." type="journal article" year="2000">Gao and Norell, 2000</bibRefCitation>
;
<bibRefCitation author="Nydam, R. L. &amp; R. L. Cifelli" box="[744,1057,716,738]" pageId="101" pageNumber="102" pagination="73 - 78" refId="ref96611" refString="Nydam, R. L., and R. L. Cifelli. 2005. New data on the dentition of the scincomorphan lizard Polyglyphanodon sternbergi. Acta Palaeontologica Polonica 50: 73 - 78." type="journal article" year="2005">Nydam and Cifelli, 2005</bibRefCitation>
). However,
<taxonomicName authorityName="Gilmore" authorityYear="1940" box="[667,863,745,768]" class="Reptilia" family="Teiidae" genus="Polyglyphanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="genus">
<emphasis box="[667,863,745,768]" italics="true" pageId="101" pageNumber="102">Polyglyphanodon</emphasis>
</taxonomicName>
-like teiioids have also often been considered to constitute up to four separate families from teiids;
<taxonomicName authorityName="Sulimski" authorityYear="1972" class="Reptilia" family="Adamisauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Adamisauridae</taxonomicName>
,
<taxonomicName authorityName="Langer" authorityYear="1998" box="[738,911,834,856]" class="Reptilia" family="Gilmoreteiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Gilmoreteiidae</taxonomicName>
(
<emphasis bold="true" box="[936,959,833,856]" pageId="101" pageNumber="102">5</emphasis>
<taxonomicName authorityName="Sulimski" authorityYear="1975" class="Reptilia" family="Macrocephalosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Macrocephalosauridae</taxonomicName>
;
<bibRefCitation author="Langer, M. C." box="[721,876,864,886]" pageId="101" pageNumber="102" pagination="13 - 18" refId="ref93187" refString="Langer, M. C. 1998. Gilmoreteiidae new family and Gilmoreteius new genus (Squamata Scincomorpha): replacement names for Macrocephalosauridae Sulimski, 1975, and Macrocephalosaurus Gilmore, 1943. Comunicacoes do Museu de Ciencias e Tecnologia da PUCRS 11: 13 - 18." type="journal article" year="1998">Langer, 1998</bibRefCitation>
),
<taxonomicName baseAuthorityName="Alifanov" baseAuthorityYear="2000" box="[894,1143,863,885]" class="Reptilia" family="Mongolochamopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Mongolochamopidae</taxonomicName>
(see Alifanov, 2000), and
<taxonomicName authorityName="Gilmore" authorityYear="1942" box="[938,1193,893,915]" class="Reptilia" family="Polyglyphanodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Polyglyphanodontidae</taxonomicName>
(see
<bibRefCitation author="Gilmore, C. W." box="[725,914,923,945]" pageId="101" pageNumber="102" pagination="229 - 265" refId="ref89934" refString="Gilmore, C. W. 1942 b. Osteology of Polyglyphanodon, an Upper Cretaceous lizard from Utah. Proceedings of the United States National Museum 92: 229 - 265." type="journal article" year="1942">Gilmore, 1942b</bibRefCitation>
;
<bibRefCitation author="Sulimski, A." box="[935,1112,923,945]" pageId="101" pageNumber="102" pagination="1 - 143" refId="ref100472" refString="Sulimski, A. 1972. Adamisaurus magnidentatus magnidentatus n. gen., n. sp. (Sauria) from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 27: 1 - 143." type="journal article" year="1972">Sulimski, 1972</bibRefCitation>
,
<bibRefCitation author="Sulimski, A." box="[1133,1189,922,944]" pageId="101" pageNumber="102" pagination="25 - 102" refId="ref100508" refString="Sulimski, A. 1975. Macrocephalosauridae, and Polyglyphanodontidae (Sauria) from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33: 25 - 102." type="journal article" year="1975">1975</bibRefCitation>
,
<bibRefCitation author="Sulimski, A." box="[667,723,952,973]" pageId="101" pageNumber="102" pagination="43 - 56" refId="ref100536" refString="Sulimski, A. 1978. New data on the genus Adamisaurus magnidentatus Sulimski 1972 (Sauria) from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 38: 43 - 56." type="journal article" year="1978">1978</bibRefCitation>
; Alifanov, 1993a, 2000;
<bibRefCitation author="Langer, M. C." box="[1022,1181,952,974]" pageId="101" pageNumber="102" pagination="13 - 18" refId="ref93187" refString="Langer, M. C. 1998. Gilmoreteiidae new family and Gilmoreteius new genus (Squamata Scincomorpha): replacement names for Macrocephalosauridae Sulimski, 1975, and Macrocephalosaurus Gilmore, 1943. Comunicacoes do Museu de Ciencias e Tecnologia da PUCRS 11: 13 - 18." type="journal article" year="1998">Langer, 1998</bibRefCitation>
). Because of this precedent and because polyglyphanodontids sensu lato (see usage below) do not fall within the crown group
<taxonomicName authorityName="Gray" authorityYear="1827" box="[1083,1169,1041,1063]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Teiidae</taxonomicName>
, I use the above definition for
<taxonomicName authorityName="Gray" authorityYear="1827" box="[987,1073,1070,1092]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="family">Teiidae</taxonomicName>
.
</paragraph>
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