296 lines
37 KiB
XML
296 lines
37 KiB
XML
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<mods:title id="A49ECD1AA455C899B4DDF4ABE6BC2379">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="23ECB13FA9F1D3D554E329A1035177D8">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="943D0FB6B8849D84FEA462B81C3FB133">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="98AFD6205A01C947759B244DB72CD1DF">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="B56F6B3BAE7A3B7DAF606F7074587412">Sole, Catherine L.</mods:namePart>
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<mods:date id="46028258B94544932522401B976D23DE">2018</mods:date>
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<subSubSection id="72C53E83FFBEFFFAFF0DFE0AFC920751" box="[151,832,506,532]" pageId="28" pageNumber="69" type="nomenclature">
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<taxonomicName id="FDDF168BFFBEFFFAFF0DFE0AFD710751" ID-CoL="6QSCZ" authority="Abdalla and Scholtz" authorityName="Abdalla & Deschodt & Scholtz & Sole" authorityYear="2018" box="[151,675,506,532]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="tortuosus" status="sp. nov.">
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<emphasis id="08ABB11AFFBEFFFAFF0DFE0AFE4F0751" bold="true" box="[151,413,506,532]" italics="true" pageId="28" pageNumber="69">Macroderes tortuosus</emphasis>
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Abdalla and Scholtz
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,
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<paragraph id="3A606D08FFBEFFFAFF0DFDEDFE7E0773" blockId="28.[151,832,506,566]" box="[151,428,541,566]" pageId="28" pageNumber="69">
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(
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<figureCitation id="A2E4718DFFBEFFFAFF05FDEDFF2E0773" box="[159,252,541,566]" captionStart="FIGURES 19–23" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 19–23. Habitus of Macroderes species. 19, M. oreatus Abdalla & Deschodt, new species; 20, M. cederbergensis Abdalla & Deschodt, new species; 21, M. soleiana Abdalla & Deschodt, new species; 22, M. tourtosus Abdalla & Scholtz, new species; 23, M. leipoldti Abdalla & Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="28" pageNumber="69">Figs. 22</figureCitation>
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,
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<figureCitation id="A2E4718DFFBEFFFAFE92FDEEFEF40773" box="[264,294,542,566]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov & Scholtz, 2005; 27, M. amplior Frolov & Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla & Deschodt, new species; 30, M. cornutus Frolov & Scholtz, 2005; 31, M. foveatus Frolov & Scholtz, 2005; 32, M. gifboomi Abdalla & Scholtz, new species; 33, M. leipoldti Abdalla & Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov & Scholtz, 2005; 36, M. oreatus Abdalla & Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla & Deschodt, new species; 39, M. tourtosus Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="28" pageNumber="69">39</figureCitation>
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,
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<figureCitation id="A2E4718DFFBEFFFAFEA8FDEEFE830773" box="[306,337,542,566]" captionStart="FIGURES 82–87" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 82–87. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla & Scholtz, new species; 83, M. cederbergensis Abdalla & Deschodt, new species; 84, M. gifboomi Abdalla & Scholtz, new species; 85, M. leipoldti Abdalla & Deschodt, new species; 86, M. oreatus Abdalla & Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="28" pageNumber="69">82</figureCitation>
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,
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<figureCitation id="A2E4718DFFBEFFFAFEC7FDEEFEA90773" box="[349,379,542,566]" captionStart="FIGURES 88–93" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 88–93. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (90–93) of Macroderes species. 88, M. soleiana Abdalla & Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="28" pageNumber="69">89</figureCitation>
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,
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<figureCitation id="A2E4718DFFBEFFFAFE1CFDEEFE710773" box="[390,419,542,566]" captionStart="FIGURES 94–99" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 94–99. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla & Deschodt, new species; 95, M. leipoldti Abdalla & Deschodt, new species; 96, M. oreatus Abdalla & Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla & Deschodt, new species; 99, M. tourtosus Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="28" pageNumber="69">99</figureCitation>
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)
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[printed] //
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<taxonomicName id="FDDF168BFFBEFFFAFF0DFCE4FE5F0669" authorityName="Abdalla & Deschodt & Scholtz & Sole" authorityYear="2018" box="[151,397,788,812]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="tortuosus">Macroderes tortuosus</taxonomicName>
|
||
/
|
||
</emphasis>
|
||
Abdalla & Scholtz / 2018 / [red label, printed]”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="8AB76755FFBEFFFAFC46FCE5FE830615" ID-GBIF-Occurrence="2028137838" collectingDate="2016-06-16" collectionCode="TMSA" collectorName="C. Scholtz" country="South Africa" elevation="377" latitude="-31.801449" location="Alt" longLatPrecision="1" longitude="18.918655" municipality="Gifberg" pageId="28" pageNumber="69" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
|
||
<typeStatus id="E564D3AAFFBEFFFAFC46FCE5FB9F0668" box="[988,1101,789,813]" pageId="28" pageNumber="69" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="2CD9A681FFBEFFFAFBC0FCE4FBAF0669" box="[1114,1149,788,812]" pageId="28" pageNumber="69" type="male">1♂</specimenCount>
|
||
and
|
||
<specimenCount id="2CD9A681FFBEFFFAFB2DFCE4FB220669" box="[1207,1264,788,812]" pageId="28" pageNumber="69" type="female">2♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="5CCEF5CDFFBEFFFAFA9AFCE4FA830669" box="[1280,1361,788,812]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="28" pageNumber="69">TMSA</collectionCode>
|
||
) same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8AB76755FFBEFFFAFEC6FCC8FB8506DD" ID-GBIF-Occurrence="2028137843" collectingDate="2016-07-21" collectionCode="TMSA" collectorName="C. Deschodt & W. P. Strumpher" country="South Africa" county="Gifberg" elevation="377" latitude="-31.801449" location="Alt" longLatPrecision="1" longitude="18.918655" municipality="Vanrhynsdorp" pageId="28" pageNumber="69" specimenCount="2" specimenCount-male="2" stateProvince="Western Cape" typeStatus="paratype">
|
||
<specimenCount id="2CD9A681FFBEFFFAFEC6FCC8FE49060A" box="[348,411,824,848]" pageId="28" pageNumber="69" type="male">2 ♂♂</specimenCount>
|
||
(
|
||
<collectionCode id="5CCEF5CDFFBEFFFAFE30FCC8FE2C0615" box="[426,510,824,848]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="28" pageNumber="69">TMSA</collectionCode>
|
||
): “
|
||
<collectingCountry id="42C82D98FFBEFFFAFD86FCC7FD7F0615" box="[540,685,823,848]" name="South Africa" pageId="28" pageNumber="69">South Africa</collectingCountry>
|
||
/
|
||
<collectingRegion id="F81BA3EAFFBEFFFAFD5EFCC8FC190615" box="[708,971,823,848]" country="South Africa" name="Western Cape" pageId="28" pageNumber="69">Western Cape Province</collectingRegion>
|
||
/
|
||
<collectingCounty id="D3011584FFBEFFFAFC7AFCC7FBE40615" box="[992,1078,823,848]" pageId="28" pageNumber="69">Gifberg</collectingCounty>
|
||
/
|
||
<quantity id="FD27C0EDFFBEFFFAFBD7FCC7FB460615" box="[1101,1172,823,848]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="28" pageNumber="69" unit="km" value="26.0">26 km</quantity>
|
||
SE of /
|
||
<collectingMunicipality id="DA04F772FFBEFFFAFB6AFCC7FA5F0615" box="[1264,1421,823,848]" pageId="28" pageNumber="69">Vanrhynsdorp</collectingMunicipality>
|
||
/
|
||
<geoCoordinate id="5FEB0BCFFFBEFFFAFF0DFCACFE910631" box="[151,323,860,885]" direction="south" orientation="latitude" pageId="28" pageNumber="69" precision="1" value="-31.801449">31° 48 '5.22 S</geoCoordinate>
|
||
<geoCoordinate id="5FEB0BCFFFBEFFFAFECAFCACFE2C0631" box="[336,510,860,885]" direction="east" orientation="longitude" pageId="28" pageNumber="69" precision="1" value="18.918655">18° 55' 7.16 E</geoCoordinate>
|
||
/
|
||
<location id="3F003BD3FFBEFFFAFD84FCACFD900630" LSID="urn:lsid:plazi:treatment:B276DC1EFFBEFFFAFF0DFE0AFF2302CD:3F003BD3FFBEFFFAFD84FCACFD900630" box="[542,578,860,885]" country="South Africa" county="Gifberg" latitude="-31.801449" longLatPrecision="1" longitude="18.918655" municipality="Vanrhynsdorp" name="Alt" pageId="28" pageNumber="69" stateProvince="Western Cape">Alt</location>
|
||
<quantity id="FD27C0EDFFBEFFFAFDD4FCACFD480631" box="[590,666,860,884]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="28" pageNumber="69" unit="m" value="377.0">
|
||
<elevation id="B1F28A3BFFBEFFFAFDD4FCACFD480631" box="[590,666,860,884]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="28" pageNumber="69" unit="m" value="377.0">377 m</elevation>
|
||
</quantity>
|
||
/
|
||
<date id="4E614BC8FFBEFFFAFD23FCACFCEB0630" box="[697,825,860,885]" pageId="28" pageNumber="69" value="2016-07-21">
|
||
<collectingDate id="5E25B220FFBEFFFAFD23FCACFCEB0630" box="[697,825,860,885]" pageId="28" pageNumber="69" value="2016-07-21">21.vii.2016</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="972A08DEFFBEFFFAFCC3FCACFC3B0630" box="[857,1001,860,885]" pageId="28" pageNumber="69">C. Deschodt</collectorName>
|
||
&
|
||
<collectorName id="972A08DEFFBEFFFAFB8CFCADFB180630" box="[1046,1226,860,885]" pageId="28" pageNumber="69">W.P. Strümpher</collectorName>
|
||
1 leg [printed] //
|
||
<typeStatus id="E564D3AAFFBEFFFAFF0DFC70FEF006DD" box="[151,290,896,920]" pageId="28" pageNumber="69" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<emphasis id="08ABB11AFFBEFFFAFEADFC70FDE906D2" box="[311,571,896,920]" italics="true" pageId="28" pageNumber="69">
|
||
<taxonomicName id="FDDF168BFFBEFFFAFEADFC70FDFE06DD" authorityName="Abdalla & Deschodt & Scholtz & Sole" authorityYear="2018" box="[311,556,896,920]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="tortuosus">Macroderes tortuosus</taxonomicName>
|
||
/
|
||
</emphasis>
|
||
Abdalla & Scholtz / 2018 / [red label, printed]”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3A606D08FFBEFFFAFF5DFC54FB6406F9" blockId="28.[151,1437,679,1928]" box="[199,1206,932,957]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DFC54FE1A06F8" bold="true" box="[199,456,932,957]" pageId="28" pageNumber="69">
|
||
<typeStatus id="E564D3AAFFBEFFFAFF5DFC54FEE606F8" box="[199,308,932,957]" pageId="28" pageNumber="69" type="holotype">Holotype</typeStatus>
|
||
description.
|
||
</emphasis>
|
||
<typeStatus id="E564D3AAFFBEFFFAFE54FC54FDEB06F8" box="[462,569,932,957]" pageId="28" pageNumber="69" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<specimenCount id="2CD9A681FFBEFFFAFDDFFC55FD8806F9" box="[581,602,933,956]" pageId="28" pageNumber="69" type="male">♂</specimenCount>
|
||
(
|
||
<figureCitation id="A2E4718DFFBEFFFAFDF3FC54FD6C06F9" box="[617,702,932,957]" captionStart="FIGURES 19–23" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 19–23. Habitus of Macroderes species. 19, M. oreatus Abdalla & Deschodt, new species; 20, M. cederbergensis Abdalla & Deschodt, new species; 21, M. soleiana Abdalla & Deschodt, new species; 22, M. tourtosus Abdalla & Scholtz, new species; 23, M. leipoldti Abdalla & Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="28" pageNumber="69">Fig. 22</figureCitation>
|
||
). Body length
|
||
<quantity id="FD27C0EDFFBEFFFAFCFBFC54FC1506F9" box="[865,967,932,957]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="28" pageNumber="69" unit="mm" value="10.6">10.6 mm</quantity>
|
||
, body width
|
||
<quantity id="FD27C0EDFFBEFFFAFBC3FC55FB6006F9" box="[1113,1202,933,956]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="28" pageNumber="69" unit="mm" value="7.7">7.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3A606D08FFBEFFFAFF5DFC37FAEE0108" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DFC37FEDF06A5" bold="true" box="[199,269,967,992]" pageId="28" pageNumber="69">Head.</emphasis>
|
||
Clypeus very broad, semi–circular, edges with narrow border and two teeth on anterior margin. Genae acute angled. Frons densely punctate with small round punctures. Frontoclypeal suture slightly curved without tubercle in the middle. Clypeogenal suture present but poorly defined towards the clypeal edge. Dorsal surface of clypeus densely punctate with more or less elongated punctures becoming rugose near the clypeal margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72C53E83FFBEFFFAFF5DFBA7FACF0300" pageId="28" pageNumber="69" type="description">
|
||
<paragraph id="3A606D08FFBEFFFAFF5DFBA7FA9D0061" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DFBA7FE950135" bold="true" box="[199,327,1111,1136]" pageId="28" pageNumber="69">Pronotum.</emphasis>
|
||
Form trapezoidal, shagreened, matte, excavated anterolaterally, lateral edges delimited by narrow borders, borders crenulated, punctate, base not bordered. Pronotum anterior angle obtusely angled, posterior rounded angled appears entire in dorsal view. Pronotal sides rugose with formation of distinct tubercles. Base lacking a longitudinal depression. Pronotal surface punctate with elongated, confluent, coarse punctures (
|
||
<figureCitation id="A2E4718DFFBEFFFAFAA3FB34FA450199" box="[1337,1431,1220,1245]" captionStart="FIGURES 82–87" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 82–87. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla & Scholtz, new species; 83, M. cederbergensis Abdalla & Deschodt, new species; 84, M. gifboomi Abdalla & Scholtz, new species; 85, M. leipoldti Abdalla & Deschodt, new species; 86, M. oreatus Abdalla & Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="28" pageNumber="69">Figs. 82</figureCitation>
|
||
,
|
||
<figureCitation id="A2E4718DFFBEFFFAFF0DFB18FF650045" box="[151,183,1256,1280]" captionStart="FIGURES 88–93" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 88–93. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (90–93) of Macroderes species. 88, M. soleiana Abdalla & Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="28" pageNumber="69">89</figureCitation>
|
||
). The confluent punctures flowing together forming twisted–like channels. Punctures with short yellow setae. Punctures becoming smaller towards the pronotum anterior edge. Punctures spaced by elevated matte areas.
|
||
</paragraph>
|
||
<paragraph id="3A606D08FFBEFFFAFF5DFADFFE5000F0" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DFADFFEC8000D" bold="true" box="[199,282,1327,1352]" pageId="28" pageNumber="69">Elytra.</emphasis>
|
||
Elytral intervals convex, strongly shagreened, matte, densely punctate with large roughly round punctures. Punctures associated with short yellow setae in the middle. Punctures separated by elevated matte areas. Striae with fine punctures spaced by 2–4 times their own diameter (
|
||
<figureCitation id="A2E4718DFFBEFFFAFC15FA87FC3600D5" box="[911,996,1399,1424]" captionStart="FIGURES 94–99" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 94–99. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla & Deschodt, new species; 95, M. leipoldti Abdalla & Deschodt, new species; 96, M. oreatus Abdalla & Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla & Deschodt, new species; 99, M. tourtosus Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="28" pageNumber="69">Fig. 99</figureCitation>
|
||
). Stria 9 short, close to stria 10, 2/3 the length of the elytron.
|
||
</paragraph>
|
||
<paragraph id="3A606D08FFBEFFFAFF5DFA4FFB37009D" blockId="28.[151,1437,679,1928]" box="[199,1253,1471,1497]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DFA4FFEE8009D" bold="true" box="[199,314,1471,1496]" pageId="28" pageNumber="69">Aedeagus</emphasis>
|
||
. Sclerite of internal sac curved basally and with very long lateral process, (
|
||
<figureCitation id="A2E4718DFFBEFFFAFB1FFA4FFB0B009D" box="[1157,1241,1471,1496]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov & Scholtz, 2005; 27, M. amplior Frolov & Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla & Deschodt, new species; 30, M. cornutus Frolov & Scholtz, 2005; 31, M. foveatus Frolov & Scholtz, 2005; 32, M. gifboomi Abdalla & Scholtz, new species; 33, M. leipoldti Abdalla & Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov & Scholtz, 2005; 36, M. oreatus Abdalla & Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla & Deschodt, new species; 39, M. tourtosus Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="28" pageNumber="69">Fig. 39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3A606D08FFBEFFFAFF5DFA14FF180365" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DFA14FE9F00B8" bold="true" box="[199,333,1508,1533]" pageId="28" pageNumber="69">Size range.</emphasis>
|
||
Males length: 10.0–
|
||
<quantity id="FD27C0EDFFBEFFFAFDA4FA14FD7400B9" box="[574,678,1508,1532]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="28" pageNumber="69" unit="mm" value="10.7">10.7 mm</quantity>
|
||
, width:
|
||
<quantity id="FD27C0EDFFBEFFFAFC9CFA14FC4600B9" box="[774,916,1508,1532]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="7.9" metricValueMin="7.4" pageId="28" pageNumber="69" unit="mm" value="7.65" valueMax="7.9" valueMin="7.4">7.4–7.9 mm</quantity>
|
||
; females length:
|
||
<quantity id="FD27C0EDFFBEFFFAFBC1FA14FB3A00B9" box="[1115,1256,1508,1532]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.8" metricValueMin="8.7" pageId="28" pageNumber="69" unit="mm" value="9.25" valueMax="9.8" valueMin="8.7">8.7–9.8 mm</quantity>
|
||
, width:
|
||
<quantity id="FD27C0EDFFBEFFFAFAD2FA14FF140365" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.2" metricValueMin="6.4" pageId="28" pageNumber="69" unit="mm" value="6.8" valueMax="7.2" valueMin="6.4">6.4–7.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3A606D08FFBEFFFAFF5DF9DCFACF0300" blockId="28.[151,1437,679,1928]" box="[199,1309,1580,1605]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DF9DCFEF70300" bold="true" box="[199,293,1580,1605]" pageId="28" pageNumber="69">Female.</emphasis>
|
||
Female differs from male by having pronotum less excavated anterolaterally than in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72C53E83FFBEFFFAFF5DF9BFFED903C9" pageId="28" pageNumber="69" type="etymology">
|
||
<paragraph id="3A606D08FFBEFFFAFF5DF9BFFED903C9" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DF9BFFE9C032D" bold="true" box="[199,334,1615,1640]" pageId="28" pageNumber="69">Etymology.</emphasis>
|
||
The name of this species is taken from the Latin adjective tortuosus (twisted) for many turns in the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72C53E83FFBEFFFAFF5DF967FCDA03BD" pageId="28" pageNumber="69" type="diagnosis">
|
||
<paragraph id="3A606D08FFBEFFFAFF5DF967FCDA03BD" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DF967FE0603F5" bold="true" box="[199,468,1687,1712]" pageId="28" pageNumber="69">Differential diagnosis.</emphasis>
|
||
This species is most similar to
|
||
<taxonomicName id="FDDF168BFFBEFFFAFCFEF969FC3B03F5" authorityName="Frolov & Scholtz" authorityYear="2005" box="[868,1001,1688,1712]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="amplior">
|
||
<emphasis id="08ABB11AFFBEFFFAFCFEF969FC3B03F5" box="[868,1001,1688,1712]" italics="true" pageId="28" pageNumber="69">M. amplior</emphasis>
|
||
</taxonomicName>
|
||
in having pronotum with elongated punctures (
|
||
<figureCitation id="A2E4718DFFBEFFFAFE8EF94CFEA00391" box="[276,370,1724,1749]" captionStart="FIGURES 82–87" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 82–87. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla & Scholtz, new species; 83, M. cederbergensis Abdalla & Deschodt, new species; 84, M. gifboomi Abdalla & Scholtz, new species; 85, M. leipoldti Abdalla & Deschodt, new species; 86, M. oreatus Abdalla & Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="28" pageNumber="69">Figs. 82</figureCitation>
|
||
,
|
||
<figureCitation id="A2E4718DFFBEFFFAFEE4F94CFE490391" box="[382,411,1724,1748]" captionStart="FIGURES 88–93" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 88–93. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (90–93) of Macroderes species. 88, M. soleiana Abdalla & Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla & Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="28" pageNumber="69">89</figureCitation>
|
||
) but it is different from the latter in having close punctures and the pronotum lateral edges punctuate and crenulation clearly visible in dorsal view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72C53E83FFBEFFFAFF5DF8F4FF2302CD" pageId="28" pageNumber="69" type="description">
|
||
<paragraph id="3A606D08FFBEFFFAFF5DF8F4FF2302CD" blockId="28.[151,1437,679,1928]" pageId="28" pageNumber="69">
|
||
<emphasis id="08ABB11AFFBEFFFAFF5DF8F4FE210258" bold="true" box="[199,499,1796,1821]" pageId="28" pageNumber="69">Habitat and distribution.</emphasis>
|
||
This species was collected from the base of the Gifberg (
|
||
<figureCitation id="A2E4718DFFBEFFFAFBECF8F4FB680259" box="[1142,1210,1796,1821]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="28" pageNumber="69">Fig. 2</figureCitation>
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) at the northern end of the Bokkeveld Escarpment Mountain Plateau just south of Vanrhynsdorp. These mountains consist of Bokkeveld Sandstone Fynbos (
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<bibRefCitation id="5E4E10F9FFBEFFFAFEE0F8BCFD100220" author="Mucina, L. & Rutherford, M. C." box="[378,706,1868,1893]" pageId="28" pageNumber="69" pagination="1 - 807" refId="ref21930" refString="Mucina, L. & Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina & Rutherford 2006</bibRefCitation>
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). The habitat is very rocky with shallow soil, poor in mineral content.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |