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<document id="960099B5D3ACF723BB2A9F3824C9C43B" ID-DOI="10.24349/acarologia/20194348" ID-ISSN="2107-7207" ID-Zenodo-Dep="5173801" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628526289822" checkinUser="carolina" docAuthor="Ermilov, Sergey G. &amp; Kalúz, Stanislav" docDate="2019" docId="822F006AAA332E0FFE1E6592FC05FA10" docLanguage="en" docName="Acarologia.59.4.456-474.pdf" docOrigin="Acarologia 59 (4)" docSource="http://dx.doi.org/10.24349/acarologia/20194348" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Leptotocepheus murphyi" docType="treatment" docVersion="4" lastPageNumber="473" masterDocId="7E167812AA382E1DFFF56038FFFBFFA6" masterDocTitle="A new species of Fissicepheus and a supplementary description of Leptotocepheus murphyi (Mahunka, 1989) (Acari, Oribatida, Otocepheidae) from Malaysia" masterLastPageNumber="474" masterPageNumber="456" pageNumber="466" updateTime="1699056533856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A9CEE27D9C3025E4095BDB2891B213A2">A new species of Fissicepheus and a supplementary description of Leptotocepheus murphyi (Mahunka, 1989) (Acari, Oribatida, Otocepheidae) from Malaysia</mods:title>
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<mods:namePart id="AF44D32363C8C92C2BF10080A0F707C0">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="121E05CEB3EFC423B21E1D7C636FC006">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="02387319606F98ED3904B852C0DBE4B1">Kalúz, Stanislav</mods:namePart>
<mods:affiliation id="69B0116530ADBE74D619856533870CAA">Institute of Zoology, Slovak Academy of Sciences, Bratislava, Slovakia.</mods:affiliation>
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<mods:title id="9005BF29A8759C781A681406A5A9D0AB">Acarologia</mods:title>
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<mods:date id="62E5C54F7629900D7DEC68ED4FB620A4">2019</mods:date>
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<mods:identifier id="7350529061E96207361045D35B4CA2E6" type="DOI">10.24349/acarologia/20194348</mods:identifier>
<mods:identifier id="62458F0CB9B5564EEC8E7D3C8DFF4985" type="ISSN">2107-7207</mods:identifier>
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<treatment id="822F006AAA332E0FFE1E6592FC05FA10" LSID="urn:lsid:plazi:treatment:822F006AAA332E0FFE1E6592FC05FA10" httpUri="http://treatment.plazi.org/id/822F006AAA332E0FFE1E6592FC05FA10" lastPageId="18" lastPageNumber="473" pageId="11" pageNumber="466">
<subSubSection id="429CE2F7AA332E16FE1E6592FBDDFA60" box="[491,1062,1450,1478]" pageId="11" pageNumber="466" type="nomenclature">
<paragraph id="0A39B17CAA332E16FE1E6592FBDDFA60" blockId="11.[491,1062,1450,1478]" box="[491,1062,1450,1478]" pageId="11" pageNumber="466">
<heading id="51710610AA332E16FE1E6592FBDDFA60" bold="true" box="[491,1062,1450,1478]" fontSize="12" level="2" pageId="11" pageNumber="466" reason="6">
<taxonomicName id="CD86CAFFAA332E16FE1E6592FBDDFA60" ID-CoL="8T8PR" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[491,1062,1450,1478]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="466" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA332E16FE1E6592FBDDFA60" bold="true" box="[491,1062,1450,1478]" pageId="11" pageNumber="466">
<emphasis id="38F26D6EAA332E16FE1E6592FCBAFA60" bold="true" box="[491,833,1450,1478]" italics="true" pageId="11" pageNumber="466">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA332E16FCA76592FBE4FA60" author="Mahunka S." box="[850,1055,1450,1478]" pageId="11" pageNumber="466" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="429CE2F7AA332E0FFE1E65D8FAAAFD18" lastPageId="18" lastPageNumber="473" pageId="11" pageNumber="466" type="description">
<paragraph id="0A39B17CAA332E16FE1E65D8FD7AFA5E" blockId="11.[491,1552,1504,2045]" box="[491,641,1504,1528]" pageId="11" pageNumber="466">
(
<figureCitation id="92BDADF9AA332E16FE0765D8FD81FA5E" box="[498,634,1504,1528]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStart-3="Figure 11" captionStart-4="Figure 12" captionStart-5="Figure 13" captionStartId-0="12.[119,188,1971,1992]" captionStartId-1="13.[119,188,1606,1627]" captionStartId-2="14.[119,188,1993,2014]" captionStartId-3="15.[119,188,1994,2015]" captionStartId-4="16.[119,188,1950,1971]" captionStartId-5="17.[119,188,1456,1477]" captionTargetBox-0="[118,1520,236,2127]" captionTargetBox-1="[118,1543,229,2127]" captionTargetBox-2="[95,1502,235,2040]" captionTargetBox-3="[95,1501,239,2041]" captionTargetBox-4="[193,1403,219,1926]" captionTargetBox-5="[118,1527,219,2127]" captionTargetId-0="figure-3@12.[118,1480,212,2138]" captionTargetId-1="figure-3@13.[118,1480,219,2145]" captionTargetId-2="figure-3@14.[175,1521,219,2122]" captionTargetId-3="figure-3@15.[193,1456,219,2006]" captionTargetId-4="figure-3@16.[193,1403,219,1931]" captionTargetId-5="figure-3@17.[118,1480,219,2145]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionTargetPageId-5="17" captionText-0="Figure 8 Leptotocepheus murphyi (Mahunka, 1989), adult: a – dorsal view (gnathosoma and legs omitted); b – ventral view (legs omitted). Scale bar 100 μm." captionText-1="Figure 9 Leptotocepheus murphyi (Mahunka, 1989), adult: a – prodorsal and notogastral condyles (unpair median notogastral condyle not developed), dorsal view; b – anterior part of body, lateral view (gnathosoma and legs omitted); c – posterior part of body, lateral view; d – subcapitulum, ventral view; e – palp, right, antiaxial view; f – anterior part of chelicera, left, paraxial view. Scale bar 50 μm (a), scale bar 100" captionText-2="Figure 10 Leptotocepheus murphyi (Mahunka, 1989), adult: a – leg I, right, antiaxial view; b – leg II, right, antiaxial view. Scale bar 50 μm." captionText-3="Figure 11 Leptotocepheus murphyi (Mahunka, 1989), adult: a – leg III, left, antiaxial view; b – leg IV, left, antiaxial view. Scale bar 50 μm." captionText-4="Figure 12 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral setah1, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d)." captionText-5="Figure 13 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – ventral view; b – anterior view; c – bothridial seta. Scale bar 200 μm (a), scale bar 100 μm (b), scale bar 10 μm (c)." figureDoi-0="http://doi.org/10.5281/zenodo.5173817" figureDoi-1="http://doi.org/10.5281/zenodo.5173819" figureDoi-2="http://doi.org/10.5281/zenodo.5173821" figureDoi-3="http://doi.org/10.5281/zenodo.5173823" figureDoi-4="http://doi.org/10.5281/zenodo.5173827" figureDoi-5="http://doi.org/10.5281/zenodo.5173829" httpUri-0="https://zenodo.org/record/5173817/files/figure.png" httpUri-1="https://zenodo.org/record/5173819/files/figure.png" httpUri-2="https://zenodo.org/record/5173821/files/figure.png" httpUri-3="https://zenodo.org/record/5173823/files/figure.png" httpUri-4="https://zenodo.org/record/5173827/files/figure.png" httpUri-5="https://zenodo.org/record/5173829/files/figure.png" pageId="11" pageNumber="466">Figures 8–13</figureCitation>
)
</paragraph>
<paragraph id="0A39B17CAA332E16FDE6663BFB1CF9C6" blockId="11.[491,1552,1504,2045]" pageId="11" pageNumber="466">
<emphasis id="38F26D6EAA332E16FDE6663BFCBCF9BD" bold="true" box="[531,839,1539,1563]" pageId="11" pageNumber="466">Supplementary description</emphasis>
— Measurements – Body length: 630–946 (
<specimenCount id="1C807AF5AA332E16FAEB663BFA48F9BD" box="[1310,1459,1539,1563]" pageId="11" pageNumber="466" type="generic">21 specimens</specimenCount>
:
<specimenCount id="1C807AF5AA332E16FA4B663BFDC5F99B" pageId="11" pageNumber="466" type="female">11 females</specimenCount>
and
<specimenCount id="1C807AF5AA332E16FD82661DFD22F99B" box="[631,729,1573,1597]" pageId="11" pageNumber="466" type="male">10 males</specimenCount>
); notogaster width: 232–398 (
<specimenCount id="1C807AF5AA332E16FBE3661DFB57F99B" box="[1046,1196,1573,1597]" pageId="11" pageNumber="466" type="generic">21 specimens</specimenCount>
). Length/width ratio 2.3–2.7. Females larger than males: 813–946 × 365–398 versus 630–730 × 232–282.
</paragraph>
<paragraph id="0A39B17CAA332E16FDE66653FA1DF903" blockId="11.[491,1552,1504,2045]" pageId="11" pageNumber="466">
Integument (
<figureCitation id="92BDADF9AA332E16FD546652FCECF924" box="[673,791,1642,1666]" captionStart="Figure 8" captionStartId="12.[119,188,1971,1992]" captionTargetBox="[118,1520,236,2127]" captionTargetId="figure-3@12.[118,1480,212,2138]" captionTargetPageId="12" captionText="Figure 8 Leptotocepheus murphyi (Mahunka, 1989), adult: a – dorsal view (gnathosoma and legs omitted); b – ventral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5173817" httpUri="https://zenodo.org/record/5173817/files/figure.png" pageId="11" pageNumber="466">Figs 8a, 8b</figureCitation>
,
<figureCitation id="92BDADF9AA332E16FCD66652FC9AF924" box="[803,865,1642,1666]" captionStart="Figure 9" captionStartId="13.[119,188,1606,1627]" captionTargetBox="[118,1543,229,2127]" captionTargetId="figure-3@13.[118,1480,219,2145]" captionTargetPageId="13" captionText="Figure 9 Leptotocepheus murphyi (Mahunka, 1989), adult: a – prodorsal and notogastral condyles (unpair median notogastral condyle not developed), dorsal view; b – anterior part of body, lateral view (gnathosoma and legs omitted); c – posterior part of body, lateral view; d – subcapitulum, ventral view; e – palp, right, antiaxial view; f – anterior part of chelicera, left, paraxial view. Scale bar 50 μm (a), scale bar 100" figureDoi="http://doi.org/10.5281/zenodo.5173819" httpUri="https://zenodo.org/record/5173819/files/figure.png" pageId="11" pageNumber="466">9b, 9c</figureCitation>
,
<figureCitation id="92BDADF9AA332E16FC986652FC72F924" box="[877,905,1642,1666]" captionStart="Figure 12" captionStartId="16.[119,188,1950,1971]" captionTargetBox="[193,1403,219,1926]" captionTargetId="figure-3@16.[193,1403,219,1931]" captionTargetPageId="16" captionText="Figure 12 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral setah1, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.5173827" httpUri="https://zenodo.org/record/5173827/files/figure.png" pageId="11" pageNumber="466">12</figureCitation>
a-d, 13a, 13b) – Body color brown, but legs, genital plates, subcapitular genae and anterior part of chelicerae dark brown. Body covered by layer
</paragraph>
<paragraph id="0A39B17CAA332E16FE1E6697FDC6F8F7" blockId="11.[491,1552,1504,2045]" pageId="11" pageNumber="466">of colorless, gel-like cerotegument. Body surface microgranulate. In addition, prodorsum, notogaster, anogenital region and anterior part of ventral plate foveolate (diameter of foveolae up to 16); region between costulae sometimes slightly rugose. Costulae partially foveolate. Lateral parts of body (between bothridia and acetabula I–III) without tubercles (their diameter up to 8).</paragraph>
<paragraph id="0A39B17CAA332E0FFDE66764FD3FFE86" blockId="11.[491,1552,1504,2045]" lastBlockId="18.[491,1541,229,1462]" lastPageId="18" lastPageNumber="473" pageId="11" pageNumber="466">
Prodorsum (
<figureCitation id="92BDADF9AA332E16FD696764FD16F8D2" box="[668,749,1884,1908]" captionStart="Figure 8" captionStartId="12.[119,188,1971,1992]" captionTargetBox="[118,1520,236,2127]" captionTargetId="figure-3@12.[118,1480,212,2138]" captionTargetPageId="12" captionText="Figure 8 Leptotocepheus murphyi (Mahunka, 1989), adult: a – dorsal view (gnathosoma and legs omitted); b – ventral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5173817" httpUri="https://zenodo.org/record/5173817/files/figure.png" pageId="11" pageNumber="466">Figs 8a</figureCitation>
,
<figureCitation id="92BDADF9AA332E16FD0D6764FCCEF8D2" box="[760,821,1884,1908]" captionStart="Figure 9" captionStartId="13.[119,188,1606,1627]" captionTargetBox="[118,1543,229,2127]" captionTargetId="figure-3@13.[118,1480,219,2145]" captionTargetPageId="13" captionText="Figure 9 Leptotocepheus murphyi (Mahunka, 1989), adult: a – prodorsal and notogastral condyles (unpair median notogastral condyle not developed), dorsal view; b – anterior part of body, lateral view (gnathosoma and legs omitted); c – posterior part of body, lateral view; d – subcapitulum, ventral view; e – palp, right, antiaxial view; f – anterior part of chelicera, left, paraxial view. Scale bar 50 μm (a), scale bar 100" figureDoi="http://doi.org/10.5281/zenodo.5173819" httpUri="https://zenodo.org/record/5173819/files/figure.png" pageId="11" pageNumber="466">9a, 9b</figureCitation>
,
<figureCitation id="92BDADF9AA332E16FCB56764FCA1F8D2" box="[832,858,1884,1908]" captionStart="Figure 12" captionStartId="16.[119,188,1950,1971]" captionTargetBox="[193,1403,219,1926]" captionTargetId="figure-3@16.[193,1403,219,1931]" captionTargetPageId="16" captionText="Figure 12 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral setah1, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.5173827" httpUri="https://zenodo.org/record/5173827/files/figure.png" pageId="11" pageNumber="466">12</figureCitation>
a-c, 13b, 13c) — Rostrum broadly rounded. Costulae well developed, long, reaching bothridia basally and lamellar setae distally. Tutorial ridges absent. One lateral carina between acetabulum I and insertion of rostral seta present on each side. Rostral (65–69) and lamellar (77–82) setae setiform, shortly ciliate on dorsal sides, curving and directed anteromedially. Interlamellar setae (98–110) thickened, barbed. Exobothridial setae Pedotecta I represented by large lamina, pedotecta II represented by small lamina. Discidia elongate triangular.
</paragraph>
<caption id="5EF9E1F4AA342E11FF82678BFEDCF879" ID-DOI="http://doi.org/10.5281/zenodo.5173817" ID-Zenodo-Dep="5173817" httpUri="https://zenodo.org/record/5173817/files/figure.png" pageId="12" pageNumber="467" startId="12.[119,188,1971,1992]" targetBox="[118,1520,236,2127]" targetPageId="12">
<paragraph id="0A39B17CAA342E11FF82678BFEDCF879" blockId="12.[119,1520,1970,2015]" pageId="12" pageNumber="467">
<emphasis id="38F26D6EAA342E11FF82678BFF34F86E" bold="true" box="[119,207,1971,1992]" pageId="12" pageNumber="467">Figure 8</emphasis>
<taxonomicName id="CD86CAFFAA342E11FF2E678AFD94F86E" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[219,623,1970,1992]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="467" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA342E11FF2E678AFE33F86E" box="[219,456,1970,1992]" italics="true" pageId="12" pageNumber="467">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA342E11FE23678AFD91F86E" author="Mahunka S." box="[470,618,1970,1992]" pageId="12" pageNumber="467" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</taxonomicName>
, adult: a – dorsal view (gnathosoma and legs omitted); b – ventral view (legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<caption id="5EF9E1F4AA352E10FF82667EFA28F92F" ID-DOI="http://doi.org/10.5281/zenodo.5173819" ID-Zenodo-Dep="5173819" httpUri="https://zenodo.org/record/5173819/files/figure.png" pageId="13" pageNumber="468" startId="13.[119,188,1606,1627]" targetBox="[118,1543,229,2127]" targetPageId="13">
<paragraph id="0A39B17CAA352E10FF82667EFA28F92F" blockId="13.[119,1534,1606,1695]" pageId="13" pageNumber="468">
<emphasis id="38F26D6EAA352E10FF82667EFF32F9FD" bold="true" box="[119,201,1606,1627]" pageId="13" pageNumber="468">Figure 9</emphasis>
<taxonomicName id="CD86CAFFAA352E10FF28667EFD89F9FA" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[221,626,1606,1628]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="468" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA352E10FF28667EFE31F9FA" box="[221,458,1606,1628]" italics="true" pageId="13" pageNumber="468">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA352E10FE2D667EFD97F9FA" author="Mahunka S." box="[472,620,1606,1628]" pageId="13" pageNumber="468" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</taxonomicName>
, adult: a – prodorsal and notogastral condyles (unpair median notogastral condyle not developed), dorsal view; b – anterior part of body, lateral view (gnathosoma and legs omitted); c – posterior part of body, lateral view; d – subcapitulum, ventral view; e – palp, right, antiaxial view; f – anterior part of chelicera, left, paraxial view. Scale bar 50 μm (a), scale bar 100
</paragraph>
</caption>
<caption id="5EF9E1F4AA362E13FF8267F1FA23F879" ID-DOI="http://doi.org/10.5281/zenodo.5173821" ID-Zenodo-Dep="5173821" box="[119,1496,1993,2015]" httpUri="https://zenodo.org/record/5173821/files/figure.png" pageId="14" pageNumber="469" startId="14.[119,188,1993,2014]" targetBox="[95,1502,235,2040]" targetPageId="14">
<paragraph id="0A39B17CAA362E13FF8267F1FA23F879" blockId="14.[119,1496,1993,2015]" box="[119,1496,1993,2015]" pageId="14" pageNumber="469">
<emphasis id="38F26D6EAA362E13FF8267F1FF21F878" bold="true" box="[119,218,1993,2014]" pageId="14" pageNumber="469">Figure 10</emphasis>
<taxonomicName id="CD86CAFFAA362E13FF1167F1FD8FF879" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[228,628,1993,2015]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="469" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA362E13FF1167F1FE2BF879" box="[228,464,1993,2015]" italics="true" pageId="14" pageNumber="469">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA362E13FE2867F1FD94F879" author="Mahunka S." box="[477,623,1993,2015]" pageId="14" pageNumber="469" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</taxonomicName>
, adult: a – leg I, right, antiaxial view; b – leg II, right, antiaxial view. Scale bar 50 μm.
</paragraph>
</caption>
<caption id="5EF9E1F4AA372E12FF8267F2FA46F879" ID-DOI="http://doi.org/10.5281/zenodo.5173823" ID-Zenodo-Dep="5173823" box="[119,1469,1994,2016]" httpUri="https://zenodo.org/record/5173823/files/figure.png" pageId="15" pageNumber="470" startId="15.[119,188,1994,2015]" targetBox="[95,1501,239,2041]" targetPageId="15">
<paragraph id="0A39B17CAA372E12FF8267F2FA46F879" blockId="15.[119,1469,1994,2016]" box="[119,1469,1994,2016]" pageId="15" pageNumber="470">
<emphasis id="38F26D6EAA372E12FF8267F2FF22F879" bold="true" box="[119,217,1994,2015]" pageId="15" pageNumber="470">Figure 11</emphasis>
<taxonomicName id="CD86CAFFAA372E12FF1667F2FD88F846" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[227,627,1994,2016]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="470" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA372E12FF1667F2FE34F846" box="[227,463,1994,2016]" italics="true" pageId="15" pageNumber="470">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA372E12FE2E67F2FD95F846" author="Mahunka S." box="[475,622,1994,2016]" pageId="15" pageNumber="470" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</taxonomicName>
, adult: a – leg III, left, antiaxial view; b – leg IV, left, antiaxial view. Scale bar 50 μm.
</paragraph>
</caption>
<caption id="5EF9E1F4AA282E0DFF8267A6FD80F847" ID-DOI="http://doi.org/10.5281/zenodo.5173827" ID-Zenodo-Dep="5173827" httpUri="https://zenodo.org/record/5173827/files/figure.png" pageId="16" pageNumber="471" startId="16.[119,188,1950,1971]" targetBox="[193,1403,219,1926]" targetPageId="16">
<paragraph id="0A39B17CAA282E0DFF8267A6FD80F847" blockId="16.[119,1494,1950,2017]" pageId="16" pageNumber="471">
<emphasis id="38F26D6EAA282E0DFF8267A6FF21F815" bold="true" box="[119,218,1950,1971]" pageId="16" pageNumber="471">Figure 12</emphasis>
<taxonomicName id="CD86CAFFAA282E0DFF1367A6FD83F812" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[230,632,1950,1972]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="471" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA282E0DFF1367A6FE29F812" box="[230,466,1950,1972]" italics="true" pageId="16" pageNumber="471">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA282E0DFE2A67A6FD89F812" author="Mahunka S." box="[479,626,1950,1972]" pageId="16" pageNumber="471" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</taxonomicName>
, adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral seta
<emphasis id="38F26D6EAA282E0DFC75678CFC77F86C" box="[896,908,1972,1994]" italics="true" pageId="16" pageNumber="471">h</emphasis>
<subScript id="9602B339AA282E0DFC7967F8FC6FF868" attach="left" box="[908,916,1984,1998]" fontSize="6" pageId="16" pageNumber="471">1</subScript>
, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d).
</paragraph>
</caption>
<caption id="5EF9E1F4AA292E0CFF826588FD7AFA7A" ID-DOI="http://doi.org/10.5281/zenodo.5173829" ID-Zenodo-Dep="5173829" httpUri="https://zenodo.org/record/5173829/files/figure.png" pageId="17" pageNumber="472" startId="17.[119,188,1456,1477]" targetBox="[118,1527,219,2127]" targetPageId="17">
<paragraph id="0A39B17CAA292E0CFF826588FD7AFA7A" blockId="17.[119,1492,1455,1500]" pageId="17" pageNumber="472">
<emphasis id="38F26D6EAA292E0CFF826588FF21FA63" bold="true" box="[119,218,1456,1477]" pageId="17" pageNumber="472">Figure 13</emphasis>
<taxonomicName id="CD86CAFFAA292E0CFF126597FD81FA63" authority="(Mahunka, 1989)" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[231,634,1455,1477]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="472" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA292E0CFF126597FE28FA63" box="[231,467,1455,1477]" italics="true" pageId="17" pageNumber="472">Leptotocepheus murphyi</emphasis>
(
<bibRefCitation id="6E17CC8DAA292E0CFE146597FD8FFA63" author="Mahunka S." box="[481,628,1455,1477]" pageId="17" pageNumber="472" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka, 1989</bibRefCitation>
)
</taxonomicName>
, adult, SEM photos: a – ventral view; b – anterior view; c – bothridial seta. Scale bar 200 μm (a), scale bar 100 μm (b), scale bar 10 μm (c).
</paragraph>
</caption>
<paragraph id="0A39B17CAA2A2E0FFDE66113FA1AFEE5" blockId="18.[491,1541,229,1462]" box="[531,1505,299,323]" pageId="18" pageNumber="473">
Anogenital region (
<figureCitation id="92BDADF9AA2A2E0FFD186113FCC4FEE5" box="[749,831,299,323]" captionStart="Figure 8" captionStartId="12.[119,188,1971,1992]" captionTargetBox="[118,1520,236,2127]" captionTargetId="figure-3@12.[118,1480,212,2138]" captionTargetPageId="12" captionText="Figure 8 Leptotocepheus murphyi (Mahunka, 1989), adult: a – dorsal view (gnathosoma and legs omitted); b – ventral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5173817" httpUri="https://zenodo.org/record/5173817/files/figure.png" pageId="18" pageNumber="473">Figs 8b</figureCitation>
,
<figureCitation id="92BDADF9AA2A2E0FFCBC6113FC7FFEE5" box="[841,900,299,323]" captionStart="Figure 9" captionStartId="13.[119,188,1606,1627]" captionTargetBox="[118,1543,229,2127]" captionTargetId="figure-3@13.[118,1480,219,2145]" captionTargetPageId="13" captionText="Figure 9 Leptotocepheus murphyi (Mahunka, 1989), adult: a – prodorsal and notogastral condyles (unpair median notogastral condyle not developed), dorsal view; b – anterior part of body, lateral view (gnathosoma and legs omitted); c – posterior part of body, lateral view; d – subcapitulum, ventral view; e – palp, right, antiaxial view; f – anterior part of chelicera, left, paraxial view. Scale bar 50 μm (a), scale bar 100" figureDoi="http://doi.org/10.5281/zenodo.5173819" httpUri="https://zenodo.org/record/5173819/files/figure.png" pageId="18" pageNumber="473">9b, 9c</figureCitation>
,
<figureCitation id="92BDADF9AA2A2E0FFC7B6113FC42FEE5" box="[910,953,299,323]" captionStart="Figure 12" captionStartId="16.[119,188,1950,1971]" captionTargetBox="[193,1403,219,1926]" captionTargetId="figure-3@16.[193,1403,219,1931]" captionTargetPageId="16" captionText="Figure 12 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral setah1, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.5173827" httpUri="https://zenodo.org/record/5173827/files/figure.png" pageId="18" pageNumber="473">12b</figureCitation>
,
<figureCitation id="92BDADF9AA2A2E0FFC316113FC13FEE5" box="[964,1000,299,323]" captionStart="Figure 13" captionStartId="17.[119,188,1456,1477]" captionTargetBox="[118,1527,219,2127]" captionTargetId="figure-3@17.[118,1480,219,2145]" captionTargetPageId="17" captionText="Figure 13 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – ventral view; b – anterior view; c – bothridial seta. Scale bar 200 μm (a), scale bar 100 μm (b), scale bar 10 μm (c)." figureDoi="http://doi.org/10.5281/zenodo.5173829" httpUri="https://zenodo.org/record/5173829/files/figure.png" pageId="18" pageNumber="473">13a</figureCitation>
) – Aggenital lyrifissures not visible. Four pairs
</paragraph>
<paragraph id="0A39B17CAA2A2E0FFE1E6175FADEFE6B" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
of genital (20–24) and one pair of aggenital (49–61) setae setiform, slightly barbed. Three pairs of adanal (57–59) and two pairs of anal (45–53) setae thickened, barbed. Adanal lyrifissures diagonal, slightly distanced from anal aperture, located between anal aperture and insertions of adanal setae
<emphasis id="38F26D6EAA2A2E0FFD86618DFD74FE6B" box="[627,655,437,461]" italics="true" pageId="18" pageNumber="473">ad</emphasis>
<subScript id="9602B339AA2A2E0FFD7A61F9FD6EFE74" attach="left" box="[655,661,449,466]" fontSize="7" pageId="18" pageNumber="473">3</subScript>
. Circumventral ridge present, but poorly visible (
<figureCitation id="92BDADF9AA2A2E0FFB46618DFAE3FE6B" box="[1203,1304,437,461]" captionStart="Figure 12" captionStartId="16.[119,188,1950,1971]" captionTargetBox="[193,1403,219,1926]" captionTargetId="figure-3@16.[193,1403,219,1931]" captionTargetPageId="16" captionText="Figure 12 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral setah1, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.5173827" httpUri="https://zenodo.org/record/5173827/files/figure.png" pageId="18" pageNumber="473">Fig. 12b</figureCitation>
).
</paragraph>
<paragraph id="0A39B17CAA2A2E0FFDE661E0FA39FDF1" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
Legs (
<figureCitation id="92BDADF9AA2A2E0FFDAB61EFFD00FE49" box="[606,763,471,495]" captionStart="Figure 10" captionStartId="14.[119,188,1993,2014]" captionTargetBox="[95,1502,235,2040]" captionTargetId="figure-3@14.[175,1521,219,2122]" captionTargetPageId="14" captionText="Figure 10 Leptotocepheus murphyi (Mahunka, 1989), adult: a – leg I, right, antiaxial view; b – leg II, right, antiaxial view. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.5173821" httpUri="https://zenodo.org/record/5173821/files/figure.png" pageId="18" pageNumber="473">Figs 10a, 10b</figureCitation>
,
<figureCitation id="92BDADF9AA2A2E0FFCFC61EFFC92FE49" box="[777,873,471,495]" captionStart="Figure 11" captionStartId="15.[119,188,1994,2015]" captionTargetBox="[95,1501,239,2041]" captionTargetId="figure-3@15.[193,1456,219,2006]" captionTargetPageId="15" captionText="Figure 11 Leptotocepheus murphyi (Mahunka, 1989), adult: a – leg III, left, antiaxial view; b – leg IV, left, antiaxial view. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.5173823" httpUri="https://zenodo.org/record/5173823/files/figure.png" pageId="18" pageNumber="473">11a, 11b</figureCitation>
,
<figureCitation id="92BDADF9AA2A2E0FFC8361EFFC2CFE49" box="[886,983,471,495]" captionStart="Figure 12" captionStartId="16.[119,188,1950,1971]" captionTargetBox="[193,1403,219,1926]" captionTargetId="figure-3@16.[193,1403,219,1931]" captionTargetPageId="16" captionText="Figure 12 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – dorsal view; b – lateral view; c – basal part of prodorsum and anterior part of notogaster, anterior view; d – mediobasal part of notogastral setah1, and notogastral foveolae. Scale bar 100 μm (a), scale bar 200 μm (b), scale bar 20 μm (c), scale bar 10 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.5173827" httpUri="https://zenodo.org/record/5173827/files/figure.png" pageId="18" pageNumber="473">12a, 12b</figureCitation>
,
<figureCitation id="92BDADF9AA2A2E0FFC1061EFFBC6FE49" box="[997,1085,471,495]" captionStart="Figure 13" captionStartId="17.[119,188,1456,1477]" captionTargetBox="[118,1527,219,2127]" captionTargetId="figure-3@17.[118,1480,219,2145]" captionTargetPageId="17" captionText="Figure 13 Leptotocepheus murphyi (Mahunka, 1989), adult, SEM photos: a – ventral view; b – anterior view; c – bothridial seta. Scale bar 200 μm (a), scale bar 100 μm (b), scale bar 10 μm (c)." figureDoi="http://doi.org/10.5281/zenodo.5173829" httpUri="https://zenodo.org/record/5173829/files/figure.png" pageId="18" pageNumber="473">13a, 13b</figureCitation>
) – Claw of each leg strong, slightly barbed dorsally. Tarsi with one pair of distinct teeth on dorsal side. Porose areas present on all femora and on trochanters III, IV (well visible in dissected specimens). Formulas of leg setation and solenidia: I (1-4-3-4-16) [1-2-2], II (1-4-3-3-15) [1-1-2], III (2-3-1-2-15) [1-1-0],
</paragraph>
<paragraph id="0A39B17CAA2A2E0FFE1E625AFAAAFD18" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
IV (1-2-2-2-12) [0-1-0]; homology of setae and solenidia indicated in
<tableCitation id="470484C7AA2A2E0FFB0F6259FAB0FDDF" box="[1274,1355,609,633]" captionStart="Table 1" captionStartId="11.[119,175,248,269]" captionText="Table 1 Leg setation and solenidia of adultFissicepheus parastriganovaen. sp. andLeptotocepheus murphyi (Mahunka, 1989)." pageId="18" pageNumber="473">Table 1</tableCitation>
. Solenidion
<subScript id="9602B339AA2A2E0FFA306255FA35FDD8" attach="both" box="[1477,1486,621,638]" fontSize="7" pageId="18" pageNumber="473">1</subScript>
φ on tibiae I setiform, others bacilliform. Famulus of tarsi I short, erect, distally blunt, inserted posterior to solenidia. Setae
<emphasis id="38F26D6EAA2A2E0FFCEE629FFCD3FD18" box="[795,808,679,702]" italics="true" pageId="18" pageNumber="473">u</emphasis>
setiform on tarsi I, II and spiniform on tarsi III, IV.
</paragraph>
</subSubSection>
<subSubSection id="429CE2F7AA2A2E0FFDE662F1FA7BFC80" pageId="18" pageNumber="473" type="materials_examined">
<paragraph id="0A39B17CAA2A2E0FFDE662F1FA7BFC80" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
<emphasis id="38F26D6EAA2A2E0FFDE662F1FD11FD47" bold="true" box="[531,746,713,737]" pageId="18" pageNumber="473">Material examined</emphasis>
— Twenty
<specimenCount id="1C807AF5AA2A2E0FFC9762F2FC05FD47" box="[866,1022,713,737]" pageId="18" pageNumber="473" type="generic">one specimens</specimenCount>
(
<specimenCount id="1C807AF5AA2A2E0FFBF962F1FB7BFD47" box="[1036,1152,713,737]" pageId="18" pageNumber="473" type="female">11 females</specimenCount>
and
<specimenCount id="1C807AF5AA2A2E0FFB4D62F1FAE7FD47" box="[1208,1308,713,737]" pageId="18" pageNumber="473" type="male">10 males</specimenCount>
):
<collectingCountry id="7291F1ECAA2A2E0FFAC562F1FA6BFD47" box="[1328,1424,713,737]" name="Malaysia" pageId="18" pageNumber="473">Malaysia</collectingCountry>
,
<collectingRegion id="C8427F9EAA2A2E0FFA6F62F1FDC6FCA2" country="Malaysia" name="Perak" pageId="18" pageNumber="473">Perak District</collectingRegion>
,
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NE to Gerik, Titiwangsa,
<geoCoordinate id="6FB2D7BBAA2A2E0FFC4762D4FBBFFCA2" box="[946,1092,748,772]" degrees="05" direction="north" minutes="36" orientation="latitude" pageId="18" pageNumber="473" precision="1" seconds="17.4" value="5.604833">05°36’17.4”N</geoCoordinate>
,
<geoCoordinate id="6FB2D7BBAA2A2E0FFBBA62D4FB1EFCA2" box="[1103,1253,748,772]" degrees="101" direction="east" minutes="32" orientation="longitude" pageId="18" pageNumber="473" precision="1" seconds="34.0" value="101.54278">101°32’34.0”E</geoCoordinate>
, alt.
<quantity id="CD7E1C99AA2A2E0FFAEB62D4FA8AFCA5" box="[1310,1393,748,772]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="473" unit="m" value="1100.0">1100 m</quantity>
, forest complex Belum–Temenggor, litter,
<date id="7E3897BCAA2A2E0FFC9B6336FB8FFC80" box="[878,1140,782,806]" pageId="18" pageNumber="473" value="2015-03-30" valueMax="2015-04-13" valueMin="2015-03-30">30 March–13 April 2015</date>
(E. Jendek &amp; O. Šauša).
</paragraph>
</subSubSection>
<subSubSection id="429CE2F7AA2A2E0FFDE66309FC05FA10" pageId="18" pageNumber="473" type="discussion">
<paragraph id="0A39B17CAA2A2E0FFDE66309FDA0FC28" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
<emphasis id="38F26D6EAA2A2E0FFDE66309FD81FCEF" bold="true" box="[531,634,817,841]" pageId="18" pageNumber="473">Remarks</emphasis>
— Morphological traits – Based on the supplementary description and original description (
<bibRefCitation id="6E17CC8DAA2A2E0FFD83636BFCE8FCCD" author="Mahunka S." box="[630,787,851,875]" pageId="18" pageNumber="473" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka 1989</bibRefCitation>
), the following diagnostic morphological traits for
<taxonomicName id="CD86CAFFAA2A2E0FFAC3636CFA57FCCD" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[1334,1452,851,875]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="473" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA2A2E0FFAC3636CFA57FCCD" box="[1334,1452,851,875]" italics="true" pageId="18" pageNumber="473">L. murphyi</emphasis>
</taxonomicName>
are proposed:
</paragraph>
<paragraph id="0A39B17CAA2A2E0FFDE663A0FD0CFB62" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
Body size: 630–946 × 232–398. Body surface microgranulate; prodorsum, notogaster, anogenital region and anterior part of ventral plate foveolate. Rostral and lamellar setae long, setiform, shortly ciliate. Interlamellar and 10 pairs of notogastral setae long, thickened, barbed. Bothridial setae distinctly or slightly dilated mediodistally. Lateral prodorsal and lateral notogastral condyles tubercle-like, medial prodorsal condyles broadly rounded, three medial notogastral condyles (one pair of additional condyles and one median unpair condyle) tubercle-like. Epimeral and genitoaggenital setae setiform, slightly barbed. Adanal and anal setae thickened, barbed. Adanal lyrifissures diagonal. Setae
<emphasis id="38F26D6EAA2A2E0FFB7C64B3FB6DFB04" box="[1161,1174,1163,1186]" italics="true" pageId="18" pageNumber="473">u</emphasis>
setiform on tarsi I, II and spiniform on tarsi III, IV.
</paragraph>
<paragraph id="0A39B17CAA2A2E0FFDE664F7F9FAFAE9" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
Comparison – Our specimens of
<taxonomicName id="CD86CAFFAA2A2E0FFC8464E8FC1DFB41" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[881,998,1231,1255]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="473" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA2A2E0FFC8464E8FC1DFB41" box="[881,998,1231,1255]" italics="true" pageId="18" pageNumber="473">L. murphyi</emphasis>
</taxonomicName>
are morphologically identical to the
<collectingCountry id="7291F1ECAA2A2E0FFA7564F7FA09FB41" box="[1408,1522,1231,1255]" name="Singapore" pageId="18" pageNumber="473">Singapore</collectingCountry>
specimens from the original description (
<bibRefCitation id="6E17CC8DAA2A2E0FFC5A64CAFBB0FAAC" author="Mahunka S." box="[943,1099,1266,1290]" pageId="18" pageNumber="473" pagination="381 - 402" refId="ref5265" refString="Mahunka S. 1989. New and interesting mites from the Geneva Museum LXIV. Oribatids from Singapore (Acari: Oribatida). Rev. suisse Zool., 96 (2): 381 - 402. doi: 10.5962 / bhl. part. 82037" type="journal article" year="1989">Mahunka 1989</bibRefCitation>
). Only one slight difference is present: bothridial setae usually distinctly dilated mediodistally versus bothridial setae near setiform, slightly dilated in median part (only one our specimen has similar structure of bothridial setae).
</paragraph>
<paragraph id="0A39B17CAA2A2E0FFDE66561FC05FA10" blockId="18.[491,1541,229,1462]" pageId="18" pageNumber="473">
Also, we have found that sometimes the median unpaired notogastral condyle is lacking in
<specimenCount id="1C807AF5AA2A2E0FFE1E6544FDA0FA32" box="[491,603,1404,1428]" pageId="18" pageNumber="473" type="male">four males</specimenCount>
in
<collectingCountry id="7291F1ECAA2A2E0FFD8B6544FD20FA32" box="[638,731,1404,1428]" name="Malaysia" pageId="18" pageNumber="473">Malaysia</collectingCountry>
versus always developed. Thus, these morphological nuances must be considered in future identifications of
<taxonomicName id="CD86CAFFAA2A2E0FFC7765A7FC0CFA10" baseAuthorityName="Mahunka" baseAuthorityYear="1989" box="[898,1015,1438,1462]" class="Arachnida" family="Tetracondylidae" genus="Leptotocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="473" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="38F26D6EAA2A2E0FFC7765A7FC0CFA10" box="[898,1015,1438,1462]" italics="true" pageId="18" pageNumber="473">L. murphyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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