treatments-xml/data/6D/61/35/6D613513FFD3CB41FF03FAC5FECFB8FD.xml
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<document id="256C092D4F9D90945089C0383DDAB01F" ID-CLB-Dataset="2548" ID-DOI="10.11646/zootaxa.5091.4.1" ID-GBIF-Dataset="63aff060-8318-4eb1-9d79-f15004b9583a" ID-ISSN="1175-5326" ID-Zenodo-Dep="5863820" ID-ZooBank="FA12D38D-EBF9-4EA1-A413-A203A9FF4EFB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642422905941" checkinUser="plazi" docAuthor="Zhang, Yi-Feng, Meng, Ling-Zeng &amp; Beaver, Roger A." docDate="2022" docId="6D613513FFD3CB41FF03FAC5FECFB8FD" docLanguage="en" docName="zootaxa.5091.4.1.pdf" docOrigin="Zootaxa 5091 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calophagus colombiana Zhang &amp; Meng &amp; Beaver 2022, sp. nov." docType="treatment" docUuid="4D030D66-7082-42A8-B4EE-17AEF9DC9428" docUuidSource="ZooBank" docVersion="7" lastPageNumber="530" masterDocId="91584D6BFFCFCB5CFF94FFF6FFC6BD3C" masterDocTitle="A review of the non-lyctine powder-post beetles of Yunnan (China) with a new genus and new species (Coleoptera: Bostrichidae)" masterLastPageNumber="545" masterPageNumber="501" pageNumber="529" updateTime="1699302562967" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D828FEE7913E6B35233B5E7966656A17" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FE5D9748233A97D08BCB8B8878A3C9FD">
<mods:title id="10E2F6B66B11F1C8086414567F12EA53">A review of the non-lyctine powder-post beetles of Yunnan (China) with a new genus and new species (Coleoptera: Bostrichidae)</mods:title>
</mods:titleInfo>
<mods:name id="FE7F90070FA031201ADA249EDC35AB71" type="personal">
<mods:role id="4B50E93ABCC603DDD230AF5F1E930CAA">
<mods:roleTerm id="82D75D3DF59108C551AD88E07031562B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="870C04F83DFCF41701E4A3EF29C4FCE4">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="8CE45ADC358FD22D824EA14B2609490E" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="65187D9D54772F2BE45C0587CDF58F06">Honghe University, College of Biological &amp; Agricultural Sciences, Mengzi, Yunnan 661199, China. &amp; zhang _ yifeng @ foxmail. com; https: // orcid. org / 0000 - 0003 - 3264 - 9472</mods:affiliation>
<mods:nameIdentifier id="06282FF8DCEDE636331BF6D17AAD399A" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2ADED4883AEF7DC1CCDB5956B67BFC00" type="personal">
<mods:role id="3ED9AE217CAA41F49E677E4F1A34410F">
<mods:roleTerm id="C0D39E2EC628C14AE1C15ED600BC5482">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8B77409A305EAD49C7C89160E98D240">Meng, Ling-Zeng</mods:namePart>
<mods:affiliation id="DDBBA14BE153D0C9BC1A7B7F97CC9F0E">Honghe University, College of Biological &amp; Agricultural Sciences, Mengzi, Yunnan 661199, China.</mods:affiliation>
</mods:name>
<mods:name id="8F03D08730E3FD4A5907270242680D19" type="personal">
<mods:role id="CF946CF11AAA5A905B22B68CE082D638">
<mods:roleTerm id="0C97A2E703ECEAE074E68730608544E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9324C25DD95AEFF32CA071D5DF143C1F">Beaver, Roger A.</mods:namePart>
<mods:affiliation id="412D3593AE72ACC77554243D59E1CE22">161 / 2 Mu 5, Soi Wat Phranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand.</mods:affiliation>
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<mods:typeOfResource id="2B2D6FAE15A59FEC2B284CC03F3DB812">text</mods:typeOfResource>
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<mods:titleInfo id="683E7867AC52C5B35343DE3CE7E23376">
<mods:title id="280DFD3812647EF2AF95BB943515DF3C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9232A8CAA65EF04DF07C32DD03362AD1">
<mods:date id="1952C309E3A3710DE1F5991C931EA3E2">2022</mods:date>
<mods:detail id="A44EF1000DF1080D834C3C03A46FE815" type="pubDate">
<mods:number id="F40748719C2FDF69761BB3916A669566">2022-01-17</mods:number>
</mods:detail>
<mods:detail id="0CFE25202994170B40F09D1973F41B18" type="volume">
<mods:number id="E6E2A6AA1404297685B1FA4E08FE2B6D">5091</mods:number>
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<mods:classification id="68B4C14FEB9E012D5F9700C741BCE011">journal article</mods:classification>
<mods:identifier id="CD30D552DF9D4C97F00EC89E54D1876E" type="CLB-Dataset">2548</mods:identifier>
<mods:identifier id="61E9089B02EACA4133618516C2E9BB48" type="DOI">10.11646/zootaxa.5091.4.1</mods:identifier>
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<mods:identifier id="1F31B9E0AD962BE7384E43973CC96587" type="Zenodo-Dep">5863820</mods:identifier>
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</mods:mods>
<treatment id="6D613513FFD3CB41FF03FAC5FECFB8FD" ID-DOI="http://doi.org/10.5281/zenodo.5872837" ID-GBIF-Taxon="192410534" ID-Zenodo-Dep="5872837" LSID="urn:lsid:zoobank.org:act:4D030D66-7082-42A8-B4EE-17AEF9DC9428" httpUri="http://treatment.plazi.org/id/6D613513FFD3CB41FF03FAC5FECFB8FD" lastPageId="29" lastPageNumber="530" pageId="28" pageNumber="529">
<subSubSection id="ADD2D78EFFD3CB40FF03FAC5FDE4B872" box="[151,546,1331,1358]" pageId="28" pageNumber="529" type="nomenclature">
<paragraph id="E5778405FFD3CB40FF03FAC5FDE4B872" blockId="28.[151,546,1331,1394]" box="[151,546,1331,1358]" pageId="28" pageNumber="529">
<heading id="BE3F3369FFD3CB40FF03FAC5FDE4B872" bold="true" box="[151,546,1331,1358]" fontSize="11" level="1" pageId="28" pageNumber="529" reason="1">
<emphasis id="D7BC5817FFD3CB40FF03FAC5FDE4B872" bold="true" box="[151,546,1331,1358]" pageId="28" pageNumber="529">
<taxonomicName id="22C8FF86FFD3CB40FF03FAC5FE78B872" authority="Zhang &amp; Meng &amp; Beaver, 2022" authorityName="Zhang &amp; Meng &amp; Beaver" authorityYear="2022" box="[151,446,1331,1358]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="D7BC5817FFD3CB40FF03FAC5FE78B872" bold="true" box="[151,446,1331,1358]" italics="true" pageId="28" pageNumber="529">Calophagus colombiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC8FE56CFFD3CB40FE51FAC2FDE4B872" box="[453,546,1332,1358]" pageId="28" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ADD2D78EFFD3CB40FF03FAAEFEF2B84E" box="[151,308,1368,1394]" pageId="28" pageNumber="529" type="description">
<paragraph id="E5778405FFD3CB40FF03FAAEFEF2B84E" blockId="28.[151,546,1331,1394]" box="[151,308,1368,1394]" pageId="28" pageNumber="529">
<figureCitation id="7DF39880FFD3CB40FF03FAAEFF21B84E" box="[151,231,1368,1394]" captionStart="FIGURE 36" captionStartId="34.[151,250,1002,1026]" captionTargetBox="[151,1423,187,978]" captionTargetId="figure-28@34.[151,1435,181,978]" captionTargetPageId="34" captionText="FIGURE 36. (A, B, C, D, E) Lateral (A), dorsal (B), abdomen (C♀, D♂) and longitudinal bands on club antennomeres views of Calophagus colombiana sp.nov.. Total horizontal scale bar of B =1 mm." figureDoi="http://doi.org/10.5281/zenodo.5864012" httpUri="https://zenodo.org/record/5864012/files/figure.png" pageId="28" pageNumber="529">Fig. 36</figureCitation>
(AE)
</paragraph>
</subSubSection>
<subSubSection id="ADD2D78EFFD3CB40FF03FA56FE8ABBAE" pageId="28" pageNumber="529" type="materials_examined">
<paragraph id="E5778405FFD3CB40FF03FA56FE8ABBAE" blockId="28.[151,1436,1440,2042]" pageId="28" pageNumber="529">
<emphasis id="D7BC5817FFD3CB40FF03FA56FE96B886" bold="true" box="[151,336,1440,1466]" pageId="28" pageNumber="529">Type material:</emphasis>
<materialsCitation id="55A08E58FFD3CB40FECBFA56FC82B8E2" ID-GBIF-Occurrence="3446329314" collectingDate="2020-04-19" collectionCode="IZAS" collectorName="LZ Meng." country="China" latitude="22.84731" location="Daweishan National Natural Reserve" longLatPrecision="1" longitude="103.75794" municipality="Female" pageId="28" pageNumber="529" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="3A733AA7FFD3CB40FECBFA56FE0AB886" box="[351,460,1440,1466]" pageId="28" pageNumber="529" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="05131E7FFFD3CB40FE4BFA56FDF0B886" box="[479,566,1440,1466]" pageId="28" pageNumber="529">Female</collectingMunicipality>
:
<collectingCountry id="9DDFC495FFD3CB40FDDDFA56FD60B886" box="[585,678,1440,1466]" name="China" pageId="28" pageNumber="529">CHINA</collectingCountry>
:
<collectingRegion id="270C4AE7FFD3CB40FD2CFA56FCC8B886" box="[696,782,1440,1466]" country="China" name="Yunnan" pageId="28" pageNumber="529">Yunnan</collectingRegion>
<location id="E017D2DEFFD3CB40FC89FA56FB1FB886" LSID="urn:lsid:plazi:treatment:6D613513FFD3CB41FF03FAC5FECFB8FD:E017D2DEFFD3CB40FC89FA56FB1FB886" box="[797,1241,1440,1466]" country="China" latitude="22.84731" longLatPrecision="1" longitude="103.75794" municipality="Female" name="Daweishan National Natural Reserve" pageId="28" pageNumber="529" stateProvince="Yunnan">Daweishan National Natural Reserve</location>
,
<geoCoordinate id="80FCE2C2FFD3CB40FB79FA57FA5EB880" box="[1261,1432,1441,1468]" degrees="22" direction="north" minutes="50" orientation="latitude" pageId="28" pageNumber="529" precision="1" seconds="50.31" value="22.84731">22°5050.31″N</geoCoordinate>
,
<geoCoordinate id="80FCE2C2FFD3CB40FF03FA33FE8BB8DC" box="[151,333,1477,1504]" degrees="103" direction="east" minutes="45" orientation="longitude" pageId="28" pageNumber="529" precision="1" seconds="28.60" value="103.75794">103°4528.60″E</geoCoordinate>
,
<date id="9176A2C5FFD3CB40FECEFA32FE1EB8E2" box="[346,472,1476,1502]" pageId="28" pageNumber="529" value="2020-04-19">
<collectingDate id="81325B2DFFD3CB40FECEFA32FE1EB8E2" box="[346,472,1476,1502]" pageId="28" pageNumber="529" value="2020-04-19">19.IV.2020</collectingDate>
</date>
, 1893m, leg.
<collectorName id="483DE1D3FFD3CB40FDE1FA32FD2EB8E2" box="[629,744,1476,1502]" pageId="28" pageNumber="529">LZ Meng.</collectorName>
(
<collectionCode id="83D91CC0FFD3CB40FD6CFA32FCFAB8E2" box="[760,828,1476,1502]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="28" pageNumber="529">IZAS</collectionCode>
)
</materialsCitation>
.
<typeStatus id="3A733AA7FFD3CB40FCDBFA32FC73B8E2" box="[847,949,1476,1502]" pageId="28" pageNumber="529" type="allotype">Allotype</typeStatus>
: Male: as
<typeStatus id="3A733AA7FFD3CB40FBBFFA32FB57B8E2" box="[1067,1169,1476,1502]" pageId="28" pageNumber="529" type="holotype">holotype</typeStatus>
.
<typeStatus id="3A733AA7FFD3CB40FB09FA32FAC8B8E2" box="[1181,1294,1476,1502]" pageId="28" pageNumber="529" type="paratype">Paratypes</typeStatus>
: as
<typeStatus id="3A733AA7FFD3CB40FAAEFA32FA5AB8E2" box="[1338,1436,1476,1502]" pageId="28" pageNumber="529" type="holotype">holotype</typeStatus>
(
<specimenCount id="F3CE4F8CFFD3CB40FF36FA1EFF1CBB3E" box="[162,218,1512,1538]" pageId="28" pageNumber="529" type="female">4♀♀</specimenCount>
,
<specimenCount id="F3CE4F8CFFD3CB40FF7FFA1EFEE0BB3E" box="[235,294,1512,1538]" pageId="28" pageNumber="529" type="male">4♂♂</specimenCount>
); as previous except:
<geoCoordinate id="80FCE2C2FFD3CB40FDBDFA1FFD12BB38" box="[553,724,1513,1540]" degrees="22" direction="north" minutes="50" orientation="latitude" pageId="28" pageNumber="529" precision="1" seconds="52.90" value="22.848028">22°5052.90″N</geoCoordinate>
,
<geoCoordinate id="80FCE2C2FFD3CB40FD77FA1FFC5FBB38" box="[739,921,1513,1540]" degrees="103" direction="east" minutes="45" orientation="longitude" pageId="28" pageNumber="529" precision="1" seconds="28.95" value="103.75804">103°4528.95″E</geoCoordinate>
,
<date id="9176A2C5FFD3CB40FC3DFA1EFBD8BB3E" box="[937,1054,1512,1538]" pageId="28" pageNumber="529" value="2020-05-29">29.V.2020</date>
, 1872m (
<specimenCount id="F3CE4F8CFFD3CB40FB06FA1EFB75BB3E" box="[1170,1203,1512,1538]" pageId="28" pageNumber="529" type="female">1♀</specimenCount>
,
<specimenCount id="F3CE4F8CFFD3CB40FB51FA1EFAC7BB3E" box="[1221,1281,1512,1538]" pageId="28" pageNumber="529" type="male">3♂♂</specimenCount>
); as previous except:
<date id="9176A2C5FFD3CB40FF67F9FAFEA9BB1A" box="[243,367,1548,1574]" pageId="28" pageNumber="529" value="2020-04-19">19.IV.2020</date>
(
<specimenCount id="F3CE4F8CFFD3CB40FE17F9FAFE6EBB1A" box="[387,424,1548,1574]" pageId="28" pageNumber="529" type="male">1♂</specimenCount>
); as previous except:
<geoCoordinate id="80FCE2C2FFD3CB40FD3EF9FBFC93BB14" box="[682,853,1549,1576]" degrees="22" direction="north" minutes="50" orientation="latitude" pageId="28" pageNumber="529" precision="1" seconds="52.14" value="22.847816">22°5052.14″N</geoCoordinate>
,
<geoCoordinate id="80FCE2C2FFD3CB40FCF0F9FBFBDFBB14" box="[868,1049,1549,1576]" degrees="103" direction="east" minutes="45" orientation="longitude" pageId="28" pageNumber="529" precision="1" seconds="28.59" value="103.75794">103°4528.59″E</geoCoordinate>
,
<date id="9176A2C5FFD3CB40FBBCF9FAFB5BBB1A" box="[1064,1181,1548,1574]" pageId="28" pageNumber="529" value="2019-05-28">28.V.2019</date>
, 1881m (
<specimenCount id="F3CE4F8CFFD3CB40FA9BF9FAFAF5BB1A" box="[1295,1331,1548,1574]" pageId="28" pageNumber="529" type="male">1♂</specimenCount>
); as previous except:
<geoCoordinate id="80FCE2C2FFD3CB40FEA8F9C7FE21BB70" box="[316,487,1585,1612]" degrees="22" direction="north" minutes="50" orientation="latitude" pageId="28" pageNumber="529" precision="1" seconds="55.03" value="22.84862">22°5055.03″N</geoCoordinate>
,
<geoCoordinate id="80FCE2C2FFD3CB40FE62F9C7FD6ABB70" box="[502,684,1585,1612]" degrees="103" direction="east" minutes="45" orientation="longitude" pageId="28" pageNumber="529" precision="1" seconds="29.29" value="103.75813">103°4529.29″E</geoCoordinate>
,
<date id="9176A2C5FFD3CB40FD28F9C6FCFCBB76" box="[700,826,1584,1610]" pageId="28" pageNumber="529" value="2020-04-19">19.IV.2020</date>
, 1843m (
<specimenCount id="F3CE4F8CFFD3CB40FC39F9C6FC2FBB76" box="[941,1001,1584,1610]" pageId="28" pageNumber="529" type="male">3♂♂</specimenCount>
); as previous except:
<geoCoordinate id="80FCE2C2FFD3CB40FB79F9C7FA5EBB70" box="[1261,1432,1585,1612]" degrees="22" direction="north" minutes="54" orientation="latitude" pageId="28" pageNumber="529" precision="1" seconds="30.79" value="22.908552">22°5430.79″N</geoCoordinate>
,
<geoCoordinate id="80FCE2C2FFD3CB40FF03F9A3FE8BBB4C" box="[151,333,1621,1648]" degrees="103" direction="east" minutes="41" orientation="longitude" pageId="28" pageNumber="529" precision="1" seconds="50.17" value="103.69727">103°4150.17″E</geoCoordinate>
,
<date id="9176A2C5FFD3CB40FECDF9A2FE08BB52" box="[345,462,1620,1646]" pageId="28" pageNumber="529" value="2019-05-29">29.V.2019</date>
, 2130m (
<specimenCount id="F3CE4F8CFFD3CB40FDAEF9A2FD9CBB52" box="[570,602,1620,1646]" pageId="28" pageNumber="529" type="female">1♀</specimenCount>
); as previous except:
<geoCoordinate id="80FCE2C2FFD3CB40FCC6F9A3FC3ABB4C" box="[850,1020,1621,1648]" degrees="22" direction="north" minutes="54" orientation="latitude" pageId="28" pageNumber="529" precision="1" seconds="30.41" value="22.908447">22°5430.41″N</geoCoordinate>
,
<geoCoordinate id="80FCE2C2FFD3CB40FB9CF9A3FB78BB4C" box="[1032,1214,1621,1648]" degrees="103" direction="east" minutes="41" orientation="longitude" pageId="28" pageNumber="529" precision="1" seconds="50.32" value="103.69731">103°4150.32″E</geoCoordinate>
,
<date id="9176A2C5FFD3CB40FB5EF9A2FA84BB52" box="[1226,1346,1620,1646]" pageId="28" pageNumber="529" value="2018-10-20">20.X.2018</date>
, 2136m (
<specimenCount id="F3CE4F8CFFD3CB40FF35F98EFF02BBAE" box="[161,196,1656,1682]" pageId="28" pageNumber="529" type="male">1♂</specimenCount>
) (all
<collectionCode id="83D91CC0FFD3CB40FE94F98EFE86BBAE" box="[256,320,1656,1682]" country="Colombia" name="Universidad de Córdoba" pageId="28" pageNumber="529" type="Herbarium">HUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="ADD2D78EFFD3CB41FF53F96AFD0EBED1" lastPageId="29" lastPageNumber="530" pageId="28" pageNumber="529" type="description">
<paragraph id="E5778405FFD3CB40FF53F96AFD6FBA1E" blockId="28.[151,1436,1440,2042]" pageId="28" pageNumber="529">
<emphasis id="D7BC5817FFD3CB40FF53F96AFE9FBB8A" bold="true" box="[199,345,1692,1718]" pageId="28" pageNumber="529">Description.</emphasis>
Length 7.08.0 mm, 2.4 × longer than wide (n = 4), body cylindrical, elongated, and slightly widened posteriorly; pronotum yellowish-brown, elytra sometimes bicoloured, becoming darker from yellowish-brown anterior to dark-brown at apical declivity, sometimes elytra wholly darker; ventral parts yellowish-brown anteriorly, abdomen blackish-brown; anterior tibiae black.
</paragraph>
<paragraph id="E5778405FFD3CB41FF53F8DAFD0ABDC5" blockId="28.[151,1436,1440,2042]" lastBlockId="29.[151,1437,151,1473]" lastPageId="29" lastPageNumber="530" pageId="28" pageNumber="529">
<emphasis id="D7BC5817FFD3CB40FF53F8DAFEC1BA7A" bold="true" box="[199,263,1836,1862]" pageId="28" pageNumber="529">Head</emphasis>
. Labrum concave at middle and covered with dense downwardly directed setae, especially at the anterior margin. Epistoma narrow, its surface densely granulate, apical margin arcuately emarginate, with moderately dense, soft, golden yellow hairlike setae directed medially. Fronto-clypeal suture sulciform, quite distinct. Frons narrow, wider than epistoma, its surface densely granulate, convex at middle, setae near epistoma curved towards middle of frons. Vertex conspicuously wrinkled. Antennae with 10 antennomeres, funiculus with 5, the funicular antennomeres gradually becoming more transverse towards club of 3 antennomeres; first club antennomere somewhat oval, the second and third elongate rectangular, the third longest, both sides of the club antennomeres with two longitudinal bands of short and dense golden yellow hairs, less distinct on the apical antennomere. Eyes slightly flattened, without setae behind ocular margins (
<figureCitation id="7DF39880FFD2CB41FDA8FF29FD79BDC5" box="[572,703,223,249]" captionStart="FIGURE 39" captionStartId="37.[151,250,1604,1628]" captionTargetBox="[151,1435,752,1581]" captionTargetId="figure-269@37.[151,1435,741,1581]" captionTargetPageId="37" captionText="FIGURE 39. (A, B, C, D, E, F) Front and anterior 45° angel views on head of Calophagus colombiana sp.nov. (A, D), Xylodrypta guochuanii sp.nov. (B, E) and Xylodrypta zhenghei sp.nov. (C, F)." figureDoi="http://doi.org/10.5281/zenodo.5864032" httpUri="https://zenodo.org/record/5864032/files/figure.png" pageId="29" pageNumber="530">Fig.39A, D</figureCitation>
).
</paragraph>
<paragraph id="E5778405FFD2CB41FF53FEF5FF19BCC9" blockId="29.[151,1437,151,1473]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FEF5FE81BC21" bold="true" box="[199,327,259,285]" pageId="29" pageNumber="530">Pronotum.</emphasis>
Transverse, 1.25 × wider than long (n = 4), strongly narrowed in front and widest behind middle, finely and more or less roughly punctured on area above anterior margin between uncinate teeth, glabrous, shining, smooth and impunctate surface on posterior half of pronotum and laterally. Lateral carina visible only posteriorly as an indistinct fold. Anterior angles each with a short, upturned tooth, behind these a row of larger asperities forming the edge of the rasp; asperities on anterior slope small, densely placed, becoming smaller towards summit. Posterior angles angulate, anterolateral part of pronotum with sparse punctures with dense, long, white, recumbent pubescence.
</paragraph>
<paragraph id="E5778405FFD2CB41FF53FE08FBCEBF25" blockId="29.[151,1437,151,1473]" box="[199,1032,510,537]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FE08FE40BF24" bold="true" box="[199,390,510,536]" pageId="29" pageNumber="530">Scutellar shield.</emphasis>
Small, tongue-shaped, slightly convex, finely punctured.
</paragraph>
<paragraph id="E5778405FFD2CB41FF53FDD5FB78BFCD" blockId="29.[151,1437,151,1473]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FDD5FEDCBF01" bold="true" box="[199,282,547,573]" pageId="29" pageNumber="530">Elytra.</emphasis>
Length to width ratio 1.7 × (n = 4); Glabrous and shining anteriorly, punctures small, shallow near base, increasing in density, size and depth posteriorly, sometimes coalescent near declivity, and interspaces very narrow. Margin of declivity with three pairs of small, slightly protruding, bluntly tipped tubercles, the first pair adjacent to suture, the second and third widely separated, the third larger, placed a little below the midpoint of the declivity; declivital surface densely punctured, the punctures circular, the base smooth, the diameter slightly less than the interspaces; suture distinctly raised in lower two-thirds of declivity, slightly less so at the apex.
</paragraph>
<paragraph id="E5778405FFD2CB41FF53FD0CFD5CBEBD" blockId="29.[151,1437,151,1473]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FD0CFEFABE28" bold="true" box="[199,316,762,788]" pageId="29" pageNumber="530">Abdomen</emphasis>
. Ventrites with dense, fine pubescence. Male: last ventrite with pleural pieces, ventries 1-4 unmodified. Female: penultimate ventrite with a median, distinctly convex process directed posteriorly and slightly ventrally, terminating in a row of four separate long sharply pointed spines, of almost equal length, the median pair more strongly ventrally curved (
<figureCitation id="7DF39880FFD2CB41FD94FC91FD4BBEBD" box="[512,653,871,897]" captionStart="FIGURE 36" captionStartId="34.[151,250,1002,1026]" captionTargetBox="[151,1423,187,978]" captionTargetId="figure-28@34.[151,1435,181,978]" captionTargetPageId="34" captionText="FIGURE 36. (A, B, C, D, E) Lateral (A), dorsal (B), abdomen (C♀, D♂) and longitudinal bands on club antennomeres views of Calophagus colombiana sp.nov.. Total horizontal scale bar of B =1 mm." figureDoi="http://doi.org/10.5281/zenodo.5864012" httpUri="https://zenodo.org/record/5864012/files/figure.png" pageId="29" pageNumber="530">Figs 36A, C</figureCitation>
).
</paragraph>
<paragraph id="E5778405FFD2CB41FF53FC7DFD0EBED1" blockId="29.[151,1437,151,1473]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FC7DFEC0BE99" bold="true" box="[199,262,907,933]" pageId="29" pageNumber="530">Legs.</emphasis>
Anterior tibia normal, widest apically, with a row of small teeth externally, covered with sparse golden yellow setae. Second and third segments of tarsi usually distinctly wider than following segments, and bearing golden-yellowish, long and thick brushes of hairs.
</paragraph>
</subSubSection>
<subSubSection id="ADD2D78EFFD2CB41FF53FC00FC56B869" pageId="29" pageNumber="530" type="diagnosis">
<paragraph id="E5778405FFD2CB41FF53FC00FC56B869" blockId="29.[151,1437,151,1473]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FC00FE84B92C" bold="true" box="[199,322,1014,1040]" pageId="29" pageNumber="530">Diagnosis.</emphasis>
The species is placed provisionally in
<taxonomicName id="22C8FF86FFD2CB41FC9FFC01FBDBB92D" authority="Lesne. The" authorityName="Lesne. The" box="[779,1053,1015,1041]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="D7BC5817FFD2CB41FC9FFC01FC54B92D" box="[779,914,1015,1041]" italics="true" pageId="29" pageNumber="530">Calophagus</emphasis>
Lesne. The
</taxonomicName>
genus at present contains only a single species,
<taxonomicName id="22C8FF86FFD2CB41FED0FBEDFD31B909" authority="Lesne, 1902" authorityName="Lesne" authorityYear="1902" box="[324,759,1051,1077]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis id="D7BC5817FFD2CB41FED0FBEDFE1AB909" box="[324,476,1051,1077]" italics="true" pageId="29" pageNumber="530">C. pekinensis</emphasis>
Lesne, 1902 (Liu 2021)
</taxonomicName>
. This is the only
<collectingRegion id="270C4AE7FFD2CB41FC55FBEDFBDBB909" box="[961,1053,1051,1077]" country="Oman" name="Shamal ash Sharqiyah" pageId="29" pageNumber="530">Oriental</collectingRegion>
genus of
<taxonomicName id="22C8FF86FFD2CB41FB05FBEDFAE6B909" authorityName="Lesne" authorityYear="1921" box="[1169,1312,1051,1077]" class="Magnoliopsida" family="Combretaceae" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="531" phylum="Tracheophyta" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
which has longitudinal bands of hairs on the club antennomeres, as found in the present species. It shares the thick brush of hairs on the frons, the three rather small tubercles on the margin of the elytra declivity, the presence of pleural pieces in the male, and the general habitus of
<taxonomicName id="22C8FF86FFD2CB41FDC1FB71FD2AB99D" authorityName="Lesne" authorityYear="1902" box="[597,748,1159,1185]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis id="D7BC5817FFD2CB41FDC1FB71FD2AB99D" box="[597,748,1159,1185]" italics="true" pageId="29" pageNumber="530">C. pekinensis</emphasis>
</taxonomicName>
. However, in contrast to
<taxonomicName id="22C8FF86FFD2CB41FB9AFB71FB72B99D" authorityName="Zhang &amp; Meng &amp; Beaver" authorityYear="2022" box="[1038,1204,1159,1185]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="colombiana">
<emphasis id="D7BC5817FFD2CB41FB9AFB71FB72B99D" box="[1038,1204,1159,1185]" italics="true" pageId="29" pageNumber="530">C. colombiana</emphasis>
</taxonomicName>
,
<taxonomicName id="22C8FF86FFD2CB41FB50FB71FA9DB99D" authorityName="Lesne" authorityYear="1902" box="[1220,1371,1159,1185]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis id="D7BC5817FFD2CB41FB50FB71FA9DB99D" box="[1220,1371,1159,1185]" italics="true" pageId="29" pageNumber="530">C. pekinensis</emphasis>
</taxonomicName>
completely lacks a lateral carina on the pronotum, and the female ventrites are unmodified (Liu 2021).
<taxonomicName id="22C8FF86FFD2CB41FB62FB5DFA5AB9F9" authorityName="Zhang &amp; Meng &amp; Beaver" authorityYear="2022" box="[1270,1436,1195,1221]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="colombiana">
<emphasis id="D7BC5817FFD2CB41FB62FB5DFA5AB9F9" box="[1270,1436,1195,1221]" italics="true" pageId="29" pageNumber="530">C. colombiana</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="22C8FF86FFD2CB41FECDFB39FE29B9D5" authorityName="Lesne" authorityYear="1902" box="[345,495,1231,1257]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis id="D7BC5817FFD2CB41FECDFB39FE29B9D5" box="[345,495,1231,1257]" italics="true" pageId="29" pageNumber="530">C. pekinensis</emphasis>
</taxonomicName>
in the coarse punctation of the posterior part of the elytra, which in
<taxonomicName id="22C8FF86FFD2CB41FB79FB39FA45B9D5" authorityName="Lesne" authorityYear="1902" box="[1261,1411,1231,1257]" class="Insecta" family="Bostrichidae" genus="Calophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis id="D7BC5817FFD2CB41FB79FB39FA45B9D5" box="[1261,1411,1231,1257]" italics="true" pageId="29" pageNumber="530">C. pekinensis</emphasis>
</taxonomicName>
is smooth and very finely punctured. Most genera of
<taxonomicName id="22C8FF86FFD2CB41FD58FB05FC9DB831" authorityName="Lesne" authorityYear="1921" box="[716,859,1267,1293]" class="Magnoliopsida" family="Combretaceae" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="531" phylum="Tracheophyta" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
are in need of revision, although some progress has been made (e.g.
<bibRefCitation id="8159F9F4FFD2CB41FED9FAE1FDE0B80D" author="Liu, L. - Y. &amp; Beaver, R. A." box="[333,550,1303,1329]" pageId="29" pageNumber="530" pagination="380" refId="ref32470" refString="Liu, L. - Y. &amp; Beaver, R. A. (2017) A review of the powderpost beetle genus, Xylopertha Guerin-Meneville, 1845, with a new species and new synonymy (Coleoptera: Bostrichidae: Bostrichinae: Xyloperthini). European Journal of Taxonomy, 380. [published online] https: // doi. org / 10.5852 / ejt. 2017.380" type="journal article" year="2017">Liu &amp; Beaver 2017</bibRefCitation>
; Liu 2021;
<bibRefCitation id="8159F9F4FFD2CB41FD30FAE1FC8DB80D" author="Liu, L. - Y. &amp; Beaver, R. A. &amp; Sanguansub, S." box="[676,843,1303,1329]" pageId="29" pageNumber="530" pagination="234 - 240" refId="ref32756" refString="Liu, L. - Y., Beaver, R. A. &amp; Sanguansub, S. (2021) Revision of the genus Xylodrypta Lesne 1901 (Coleoptera: Bostrichidae: Bostrichinae: Xyloperthini) with a new species and a key to species. Zootaxa, 5005 (2), 234 - 240. https: // doi. org / 10.11646 / zootaxa. 5005.2.9" type="journal article" year="2021">
Liu
<emphasis id="D7BC5817FFD2CB41FD45FAE1FCC4B80D" box="[721,770,1303,1329]" italics="true" pageId="29" pageNumber="530">et al</emphasis>
. 2021
</bibRefCitation>
), and further studies, preferably comparing DNA, are needed to determine whether the present species is correctly placed.
</paragraph>
</subSubSection>
<subSubSection id="ADD2D78EFFD2CB41FF53FAA9FECFB8FD" pageId="29" pageNumber="530" type="etymology">
<paragraph id="E5778405FFD2CB41FF53FAA9FECFB8FD" blockId="29.[151,1437,151,1473]" pageId="29" pageNumber="530">
<emphasis id="D7BC5817FFD2CB41FF53FAA9FE88B845" bold="true" box="[199,334,1375,1401]" pageId="29" pageNumber="530">Etymology.</emphasis>
The species is named in reference to the famous voyager, Cristóbal Colón (Spanish, 1451-1506), as both Chinese authors of this manuscript have dreamed, during their youth, of being a captain and sailing around the world.
</paragraph>
</subSubSection>
</treatment>
</document>