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<document id="8972E1A273E4D668B14232DB800842E6" ID-CLB-Dataset="20873" ID-DOI="10.11646/zootaxa.3786.1.5" ID-GBIF-Dataset="91db6810-ed6b-4f49-8d74-7b52138d3b61" ID-ISSN="1175-5326" ID-Zenodo-Dep="4913383" ID-ZooBank="786AEFE5-374D-49DB-A084-AB0BEECD7FDF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623170264580" checkinUser="felipe" docAuthor="Maruyama, Munetoshi, Bartolozzi, Luca, Inui, Yoko, Tanaka, Hiroshi O., Hyodo, Fujio, Shimizu-Kaya, Usun, Takematsu, Yoko, Hishi, Takuo &amp; Itioka, Takao" docDate="2014" docId="6B4925292E38F14CFF477117FD51FD07" docLanguage="en" docName="zt03786p078.pdf" docOrigin="Zootaxa 3786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pycnotarsobrentus inuiae Maruyama &amp; Bartolozzi &amp; Inui &amp; Tanaka &amp; Hyodo &amp; Shimizu-Kaya &amp; Takematsu &amp; Hishi &amp; Itioka 2014, sp. nov." docType="treatment" docVersion="5" lastPageNumber="77" masterDocId="97705D512E39F148FFD07638FFEEFFCA" masterDocTitle="A new genus and species of myrmecophilous brentid beetle (Coleoptera: Brentidae) inhabiting the myrmecophytic epiphytes in the Bornean rainforest canopy" masterLastPageNumber="78" masterPageNumber="73" pageNumber="74" updateTime="1699005028685" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7F38926834F918E89C05FD9701D316AE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0F7D4EEC817F2E86AFA150AEDC6ED79D">
<mods:title id="4DB5CA6501C0131AE39E6A11150EA1DF">A new genus and species of myrmecophilous brentid beetle (Coleoptera: Brentidae) inhabiting the myrmecophytic epiphytes in the Bornean rainforest canopy</mods:title>
</mods:titleInfo>
<mods:name id="96E0EEA5AB1533D4CBF1B592EBD914A9" type="personal">
<mods:role id="2B37A0428DD2AC72849BB1F4D07EFFB0">
<mods:roleTerm id="05E230CE2E6FA3B186B208A3CD448C9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="823298EAAD9CEB9424E68BFA0BB38030">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="05394A2E1C094A404906AA3E8362330A">The Kyushu University Museum, Hakozaki, Fukuoka, 812 - 8581 Japan. E-mail: dendrolasius @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="31C1A7255809D8980F1FC5A0B8A42E44" type="email">dendrolasius@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="23EBF5E7BC60B3D30621CBFCAE03EC4E" type="personal">
<mods:role id="D103F010B8DFD7B331345B2CD0CA1ACD">
<mods:roleTerm id="EF5AE916269A571E6032B90DB61CDE58">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B920E23DCAE9513418DBE8A3F345055A">Bartolozzi, Luca</mods:namePart>
<mods:affiliation id="7493C9AE0575F13215C02FF83953CB7D">Sezione di Zoologia &quot; La Specola &quot;, Museo di Storia Naturale, Università di Firenze, via Romana 17, 50125 Firenze, Italy.</mods:affiliation>
</mods:name>
<mods:name id="EB10907516FA7FB553077496211BCB57" type="personal">
<mods:role id="DFC1B964C7980C30BC74EEC2C22F0049">
<mods:roleTerm id="260A811D74C17D9ED358D5DC448A46C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EDFC5CAB831E9D00E63D27919CD9D56">Inui, Yoko</mods:namePart>
<mods:affiliation id="C3B0633FEA72EAE04870352F706E8564">Department of Arts and Sciences, Faculty of Education, Osaka Kyoiku University 4 - 698 - 1 Asahigaoka, Kashiwara, Osaka, 582 - 8582 Japan</mods:affiliation>
</mods:name>
<mods:name id="2C99CE8CBCE34A9C4EC02B1CC033462A" type="personal">
<mods:role id="2EEB5A3C0840C3FE824B6BA504BE8192">
<mods:roleTerm id="2862B2CEB615CAFC8B0B24F058327E14">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A79D11B5CB343524F7F3F192511C56D">Tanaka, Hiroshi O.</mods:namePart>
<mods:affiliation id="20C1D6B4BE84CEFEBDAE683313E23C13">Graduate School of Human and Environmental Studies, Kyoto University, Japan</mods:affiliation>
</mods:name>
<mods:name id="5C32067EDECAE59C767ABD712E4BA1A4" type="personal">
<mods:role id="54CB5203C73835E3171050366EA77EB7">
<mods:roleTerm id="5203C2416AF1EAD4AFAD37C61848E0C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="71DA58F64CC54E77F21E5ADBC1D9E916">Hyodo, Fujio</mods:namePart>
<mods:affiliation id="7596E5B1AFD4EC38C5AB9E74F0AF1FFF">Research Core for Interdisciplinary Sciences, Okayama University, 3 - 1 - 1, Tsushimanaka, Okayama, 700 - 8530 Japan</mods:affiliation>
</mods:name>
<mods:name id="D26EB02422E1C5233947B17580F0B1B2" type="personal">
<mods:role id="63B6C77527FB60AD40453F11DBCB3683">
<mods:roleTerm id="834E3C9E935084C640E9EC0B2A4F6C74">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C638C10FD4128543F8B253F487D544C">Shimizu-Kaya, Usun</mods:namePart>
<mods:affiliation id="03CF01214D21C35B5C33700639A478A0">Graduate School of Human and Environmental Studies, Kyoto University, Japan</mods:affiliation>
</mods:name>
<mods:name id="4096EC0BF31C78726BD09F6A4442C181" type="personal">
<mods:role id="F65B1973E143B353B1830B38A8D01BCF">
<mods:roleTerm id="D2C418C5B4B9C08D1F98A8651C4E674B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="362884E4C96C92C01328AC54CC7092F3">Takematsu, Yoko</mods:namePart>
<mods:affiliation id="4F08B0E523EF61DB307D36F1B8BA8B40">Department of Biological and Environmental Sciences, Faculty of Agriculture, Yamaguchi University, 1677 - 1 Yoshida, Yamaguchi 753 - 8515 Japan</mods:affiliation>
</mods:name>
<mods:name id="071ED0F136501142E1AC59BB98F4A257" type="personal">
<mods:role id="B00CF56E860251E9FC0CB41E5EC36640">
<mods:roleTerm id="54F9F50E223645AE3C7BC7BE9D4A6DA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="359FB079AB3D01909D8D199DC7237FE3">Hishi, Takuo</mods:namePart>
<mods:affiliation id="369BF09164D3AC8906172F33401EA104">Kyushu University Forest, 394 Tsubakuro, Sasaguri-cho, Fukuoka, 811 - 2415 Japan</mods:affiliation>
</mods:name>
<mods:name id="02027E986E5E212871C778E7B57BF767" type="personal">
<mods:role id="2A8EFBBFC589E83780A8457B0CAD33FE">
<mods:roleTerm id="E2E66BD128D61B004DB8EC597DCB231E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1C75D0848EC3BFE881376E41C314FC1">Itioka, Takao</mods:namePart>
<mods:affiliation id="9B26154AF8DF5D1B0D393324250D2F68">Graduate School of Human and Environmental Studies, Kyoto University, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="AD8A335436A4FABBAE7BC477E8D5211B">text</mods:typeOfResource>
<mods:relatedItem id="4228C08EF668CB315E9F792B09D3A628" type="host">
<mods:titleInfo id="6594C68A949C626CC037E61D174D9FA1">
<mods:title id="8E01197FB3C9F1F2F04B0BC2420407C6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DB6F2F28148E9D737440923A03F692CA">
<mods:date id="1D3E87E7A75ED4916530FC521AACDEAC">2014</mods:date>
<mods:detail id="EFFB660DB0DBE2A24E0D0B99208B5E12" type="pubDate">
<mods:number id="156D4DAD5AE56D8EB54AF1DD66D8F652">2014-04-08</mods:number>
</mods:detail>
<mods:detail id="A02D0D5B75B8ED09F5E756C96F775D27" type="volume">
<mods:number id="AB54DEB07DB83BCADF10AA6B8A929EBF">3786</mods:number>
</mods:detail>
<mods:detail id="1E03838833B544888CB5B7DCA9B0D90E" type="issue">
<mods:number id="69DD234536DC74D60ED0ED5B6EE7259F">1</mods:number>
</mods:detail>
<mods:extent id="4AA8118E18911026C0315985DC2EDF55" unit="page">
<mods:start id="4CB6965953E2B592A10AEF91CC05EDCD">73</mods:start>
<mods:end id="46CF45396B646B2EA49B3E9E89EE844A">78</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="03A8703631974A05A4B16A93766FCE70">journal article</mods:classification>
<mods:identifier id="0E41C8A9E922760898D9DF119F96592D" type="CLB-Dataset">20873</mods:identifier>
<mods:identifier id="F39B4EEF7B93657FC272E77F9C41A28E" type="DOI">10.11646/zootaxa.3786.1.5</mods:identifier>
<mods:identifier id="53A347E37E689BE0BA141126C59C1F6F" type="GBIF-Dataset">91db6810-ed6b-4f49-8d74-7b52138d3b61</mods:identifier>
<mods:identifier id="CA560D6D1F91E3FFADE0FEED1F8B322F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1165359C5BA9C60DABBD2032DE0E01EF" type="Zenodo-Dep">4913383</mods:identifier>
<mods:identifier id="B9ECBA2111A36DD985978EB37ABDDDF8" type="ZooBank">786AEFE5-374D-49DB-A084-AB0BEECD7FDF</mods:identifier>
</mods:mods>
<treatment id="6B4925292E38F14CFF477117FD51FD07" ID-DOI="http://doi.org/10.5281/zenodo.4913387" ID-GBIF-Taxon="192936709" ID-Zenodo-Dep="4913387" LSID="urn:lsid:plazi:treatment:6B4925292E38F14CFF477117FD51FD07" httpUri="http://treatment.plazi.org/id/6B4925292E38F14CFF477117FD51FD07" lastPageId="4" lastPageNumber="77" pageId="1" pageNumber="74">
<subSubSection id="ABFAC7B42E38F149FF477117FC9AF880" box="[151,884,1839,1866]" pageId="1" pageNumber="74" type="nomenclature">
<paragraph id="E35F943F2E38F149FF477117FC9AF880" blockId="1.[151,884,1839,1866]" box="[151,884,1839,1866]" pageId="1" pageNumber="74">
<heading id="B81723532E38F149FF477117FC9AF880" bold="true" box="[151,884,1839,1866]" fontSize="11" level="1" pageId="1" pageNumber="74" reason="1">
<emphasis id="D194482D2E38F149FF477117FC9AF880" bold="true" box="[151,884,1839,1866]" pageId="1" pageNumber="74">
<taxonomicName id="24E0EFBC2E38F149FF477117FCE7F880" authority="Maruyama &amp; Bartolozzi" authorityName="Maruyama &amp; Bartolozzi &amp; Inui &amp; Tanaka &amp; Hyodo &amp; Shimizu-Kaya &amp; Takematsu &amp; Hishi &amp; Itioka" authorityYear="2014" box="[151,777,1839,1866]" class="Insecta" family="Brentidae" genus="Pycnotarsobrentus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="74" phylum="Arthropoda" rank="species" species="inuiae" status="sp. nov.">
<emphasis id="D194482D2E38F149FF477117FE22F883" bold="true" box="[151,460,1839,1865]" italics="true" pageId="1" pageNumber="74">Pycnotarsobrentus inuiae</emphasis>
Maruyama &amp; Bartolozzi
</taxonomicName>
,
<taxonomicNameLabel id="CAA7F5562E38F149FCC67108FC9AF880" box="[790,884,1840,1866]" pageId="1" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABFAC7B42E38F149FF477149FD8EF83C" pageId="1" pageNumber="74" type="materials_examined">
<paragraph id="E35F943F2E38F149FF477149FD8EF83C" blockId="1.[151,1436,1905,2038]" pageId="1" pageNumber="74">
<emphasis id="D194482D2E38F149FF477149FEA8F840" bold="true" box="[151,326,1905,1930]" inLineHeading="true" pageId="1" pageNumber="74" reason="1">Type material.</emphasis>
<materialsCitation id="53889E622E38F149FE9F7149FF53F864" ID-GBIF-Occurrence="3459914303" collectingDate="2011-09-22" country="Malaysia" location="Lambir Hills National Park" municipality="Itioka" pageId="1" pageNumber="74" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="3C5B2A9D2E38F149FE9F7149FE54F840" box="[335,442,1905,1930]" pageId="1" pageNumber="74" type="holotype">Holotype</typeStatus>
,
<specimenCount id="F5E65FB62E38F149FE16714AFE35F843" box="[454,475,1906,1929]" pageId="1" pageNumber="74" type="male">♂</specimenCount>
(DFS),
<location id="E63FC2E42E38F149FDEA7149FC9DF840" LSID="urn:lsid:plazi:treatment:6B4925292E38F14CFF477117FD51FD07:E63FC2E42E38F149FDEA7149FC9DF840" box="[570,883,1905,1930]" country="Malaysia" municipality="Itioka" name="Lambir Hills National Park" pageId="1" pageNumber="74">Lambir Hills National Park</location>
, Sarawak, E.
<collectingCountry id="9BF7D4AF2E38F149FBC17149FB95F840" box="[1041,1147,1905,1930]" name="Malaysia" pageId="1" pageNumber="74">Malaysia</collectingCountry>
,
<date id="975EB2FF2E38F149FB587149FAE3F840" box="[1160,1293,1905,1930]" pageId="1" pageNumber="74" value="2011-09-22">
<collectingDate id="871A4B172E38F149FB587149FAE3F840" box="[1160,1293,1905,1930]" pageId="1" pageNumber="74" value="2011-09-22">22 IX 2011</collectingDate>
</date>
,
<collectingMunicipality id="033B0E452E38F149FAC97149FAB4F840" box="[1305,1370,1905,1930]" pageId="1" pageNumber="74">Itioka</collectingMunicipality>
et al. leg
</materialsCitation>
.
<materialsCitation id="53889E622E38F149FF1971AEFCE5F864" ID-GBIF-Occurrence="3459914301" box="[201,779,1941,1966]" collectingDate="2011-09-28" country="Malaysia" location="Lambir Hills National Park" municipality="Itioka" pageId="1" pageNumber="74" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="3C5B2A9D2E38F149FF1971AEFED7F864" box="[201,313,1942,1966]" pageId="1" pageNumber="74" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="F5E65FB62E38F149FE9671ADFE84F864" box="[326,362,1941,1966]" pageId="1" pageNumber="74" type="male">3♂</specimenCount>
(KUM), same data, but
<date id="975EB2FF2E38F149FD5571ADFCE5F864" box="[645,779,1941,1966]" pageId="1" pageNumber="74" value="2011-09-28">
<collectingDate id="871A4B172E38F149FD5571ADFCE5F864" box="[645,779,1941,1966]" pageId="1" pageNumber="74" value="2011-09-28">28 IX 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="53889E622E38F149FCC971ADFB3CF864" ID-GBIF-Occurrence="3459914302" box="[793,1234,1941,1966]" collectionCode="MZUF" country="Malaysia" pageId="1" pageNumber="74" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="F5E65FB62E38F149FCC971ADFCD2F864" box="[793,828,1941,1966]" pageId="1" pageNumber="74" type="male">1♂</specimenCount>
(
<collectionCode id="85F10CFA2E38F149FC9D71ADFC73F864" box="[845,925,1941,1966]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="1" pageNumber="74">MZUF</collectionCode>
, collection number 17085)
</materialsCitation>
,
<materialsCitation id="53889E622E38F149FB3071ADFD40F818" ID-GBIF-Occurrence="3459914304" collectingDate="2005-09-17" collectionCode="T" country="Malaysia" pageId="1" pageNumber="74" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F5E65FB62E38F149FB3071ADFAEDF864" box="[1248,1283,1941,1966]" pageId="1" pageNumber="74" type="female">1♀</specimenCount>
(DFS), same data but
<date id="975EB2FF2E38F149FED07181FE67F818" box="[256,393,1977,2002]" pageId="1" pageNumber="74" value="2005-09-17">
<collectingDate id="871A4B172E38F149FED07181FE67F818" box="[256,393,1977,2002]" pageId="1" pageNumber="74" value="2005-09-17">17 IX 2005</collectingDate>
</date>
(ex
<collectionCode id="85F10CFA2E38F149FE147182FE39F81B" box="[452,471,1978,2001]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="74">T</collectionCode>
. Itioka collection)
</materialsCitation>
. All the specimens were collected from nests of
<taxonomicName id="24E0EFBC2E38F149FB257182FF12F83F" authorityName="F. Smith" authorityYear="1857" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="74" phylum="Arthropoda" rank="species" species="difformis">
<emphasis id="D194482D2E38F149FB257182FF12F83F" italics="true" pageId="1" pageNumber="74">Crematogaster difformis</emphasis>
</taxonomicName>
in domatia of the canopy ferns.
</paragraph>
</subSubSection>
<caption id="B79FC4B72E3BF14AFF4771B6FB73F828" ID-DOI="http://doi.org/10.5281/zenodo.4913385" ID-Zenodo-Dep="4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="2" pageNumber="75" startId="2.[151,263,1934,1957]" targetBox="[180,1397,203,1906]" targetPageId="2">
<paragraph id="E35F943F2E3BF14AFF4771B6FB73F828" blockId="2.[151,1436,1934,2018]" pageId="2" pageNumber="75">
<emphasis id="D194482D2E3BF14AFF4771B6FED4F86E" bold="true" box="[151,314,1934,1957]" pageId="2" pageNumber="75">FIGURES 1–7.</emphasis>
<taxonomicName id="24E0EFBC2E3BF14AFE9171B7FDABF86E" authority="Maruyama &amp; Bartolozzi &amp; Inui &amp; Tanaka &amp; Hyodo &amp; Shimizu-Kaya &amp; Takematsu &amp; Hishi &amp; Itioka, 2014" authorityName="Maruyama &amp; Bartolozzi &amp; Inui &amp; Tanaka &amp; Hyodo &amp; Shimizu-Kaya &amp; Takematsu &amp; Hishi &amp; Itioka" authorityYear="2014" box="[321,581,1935,1956]" class="Insecta" family="Brentidae" genus="Pycnotarsobrentus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="species" species="inuiae" status="gen. nov. and sp. nov.">
<emphasis id="D194482D2E3BF14AFE9171B7FDABF86E" box="[321,581,1935,1956]" italics="true" pageId="2" pageNumber="75">Pycnotarsobrentus inuiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAA7F5562E3BF14AFD9D71B7FCDEF86E" box="[589,816,1934,1956]" pageId="2" pageNumber="75" rank="species">
<emphasis id="D194482D2E3BF14AFD9D71B7FD42F86E" bold="true" box="[589,684,1935,1956]" pageId="2" pageNumber="75">gen. nov.</emphasis>
and
<emphasis id="D194482D2E3BF14AFD0F71B7FCDEF86E" bold="true" box="[735,816,1935,1956]" pageId="2" pageNumber="75">sp. nov.</emphasis>
</taxonomicNameLabel>
1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view.
</paragraph>
</caption>
<caption id="B79FC4B72E3AF14BFF477355FC1FFA69" ID-DOI="http://doi.org/10.5281/zenodo.4913391" ID-Zenodo-Dep="4913391" httpUri="https://zenodo.org/record/4913391/files/figure.png" pageId="3" pageNumber="76" startId="3.[151,263,1389,1412]" targetBox="[151,1436,207,1347]" targetPageId="3">
<paragraph id="E35F943F2E3AF14BFF477355FC1FFA69" blockId="3.[151,1436,1389,1443]" pageId="3" pageNumber="76">
<emphasis id="D194482D2E3AF14BFF477355FEA0FA49" bold="true" box="[151,334,1389,1412]" pageId="3" pageNumber="76">FIGURES 8–10.</emphasis>
Habitat of
<taxonomicName id="24E0EFBC2E3AF14BFE077356FD0CFA4E" authority="Maruyama &amp; Bartolozzi &amp; Inui &amp; Tanaka &amp; Hyodo &amp; Shimizu-Kaya &amp; Takematsu &amp; Hishi &amp; Itioka, 2014" authorityName="Maruyama &amp; Bartolozzi &amp; Inui &amp; Tanaka &amp; Hyodo &amp; Shimizu-Kaya &amp; Takematsu &amp; Hishi &amp; Itioka" authorityYear="2014" box="[471,738,1390,1412]" class="Insecta" family="Brentidae" genus="Pycnotarsobrentus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="inuiae" status="gen. nov. and sp. nov.">
<emphasis id="D194482D2E3AF14BFE077356FD0CFA4E" box="[471,738,1390,1412]" italics="true" pageId="3" pageNumber="76">Pycnotarsobrentus inuiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAA7F5562E3AF14BFD217357FC1EFA4E" box="[753,1008,1390,1412]" pageId="3" pageNumber="76" rank="species">
<emphasis id="D194482D2E3AF14BFD217357FCB9FA4E" bold="true" box="[753,855,1391,1412]" pageId="3" pageNumber="76">gen. nov.</emphasis>
and
<emphasis id="D194482D2E3AF14BFC487357FC1EFA4E" bold="true" box="[920,1008,1391,1412]" pageId="3" pageNumber="76">sp. nov.</emphasis>
</taxonomicNameLabel>
8—
<taxonomicName id="24E0EFBC2E3AF14BFBF47356FA8BFA4E" authority="Copel." authorityName="Copel." box="[1060,1381,1390,1412]" class="Polypodiopsida" family="Polypodiaceae" genus="Platycerium" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="76" phylum="Tracheophyta" rank="species" species="crustacea">
<emphasis id="D194482D2E3AF14BFBF47356FAFFFA4E" box="[1060,1297,1390,1412]" italics="true" pageId="3" pageNumber="76">Platycerium crustacea</emphasis>
Copel.
</taxonomicName>
, 9—
<taxonomicName id="24E0EFBC2E3AF14BFF4773B6FE3EFA69" authority="H. Christ." authorityName="H. Christ." box="[151,464,1421,1443]" class="Polypodiopsida" family="Polypodiaceae" genus="Lecanopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="76" phylum="Tracheophyta" rank="species" species="ridleyi">
<emphasis id="D194482D2E3AF14BFF4773B6FE8DFA68" box="[151,355,1421,1443]" italics="true" pageId="3" pageNumber="76">Lecanopteris ridleyi</emphasis>
H. Christ.
</taxonomicName>
, 10, 11—living male in a domatia of
<taxonomicName id="24E0EFBC2E3AF14BFC9973B6FC1FFA69" box="[841,1009,1422,1443]" class="Polypodiopsida" family="Polypodiaceae" genus="Lecanopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="76" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="D194482D2E3AF14BFC9973B6FC23FA69" box="[841,973,1422,1443]" italics="true" pageId="3" pageNumber="76">Lecanopteris</emphasis>
sp.
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="ABFAC7B42E3AF14CFF1773F7FD38FE42" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76" type="description">
<paragraph id="E35F943F2E3AF14BFF1773F7FBFBFA22" blockId="3.[151,1436,1487,2016]" box="[199,1045,1487,1512]" pageId="3" pageNumber="76">
<emphasis id="D194482D2E3AF14BFF1773F7FE4DFA22" bold="true" box="[199,419,1487,1512]" pageId="3" pageNumber="76">
<emphasis id="D194482D2E3AF14BFF1773F7FEB7FA22" box="[199,345,1487,1512]" inLineHeading="true" pageId="3" pageNumber="76" reason="1">Description.</emphasis>
Male.
</emphasis>
Body (
<figureCitation id="7BDB88BA2E3AF14BFE2673F7FD8EFA22" box="[502,608,1487,1512]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="3" pageNumber="76">Figs. 1, 2</figureCitation>
) dark brown, with legs slightly lighter.
</paragraph>
<paragraph id="E35F943F2E3AF14BFF1773CBFF3BF92E" blockId="3.[151,1436,1487,2016]" pageId="3" pageNumber="76">
Head (
<figureCitation id="7BDB88BA2E3AF14BFEC773CBFEB1F9C6" box="[279,351,1523,1548]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="3" pageNumber="76">Fig. 3</figureCitation>
) short, eyes totally occupying its lateral sides until basal constriction; vertex connected to mesorostral plate on same plane; surface of vertex with sparse yellow minute setae; metarostrum about as long as head, laterally concave; lateral apophysis preceded by minute tooth; mesorostrum large, flat, laterally rounded; prorostrum short, almost parallel sided; lateral margins of mesorostrum and prorostrum moderately punctuate; underside of meta- and mesorostrum moderately covered with setiferous punctures; antennal segments cylindrical, rather compressed, with rugose surface; segments II-X of antennae all wider than long; segment XI twice as long as wide.
</paragraph>
<paragraph id="E35F943F2E3AF14BFF1770C8FB91F8E1" blockId="3.[151,1436,1487,2016]" pageId="3" pageNumber="76">
Pronotum (
<figureCitation id="7BDB88BA2E3AF14BFE9570D7FE6BF8C2" box="[325,389,1775,1800]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="3" pageNumber="76">Fig. 1</figureCitation>
) 1.1× as long as wide, with similar width at anterior and posterior margins, widest at middle; finely punctured on disc, strongly rugose at sides, with some scattered erect golden setae.
</paragraph>
<paragraph id="E35F943F2E3AF14BFF17710FFB4AF8BE" blockId="3.[151,1436,1487,2016]" pageId="3" pageNumber="76">
Elytra (
<figureCitation id="7BDB88BA2E3AF14BFECD710FFE8EF89A" box="[285,352,1847,1872]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="3" pageNumber="76">Fig. 1</figureCitation>
) 2.4× as long as pronotum; all striae and interstriae present; humeral calli slightly prominent; interstriae 3 and 5 slightly elevated; surface with sparse suberect small setae, denser at sides.
</paragraph>
<paragraph id="E35F943F2E3AF14CFF1771B8FF3CFF32" blockId="3.[151,1436,1487,2016]" lastBlockId="4.[151,1436,151,717]" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76">
Legs (
<figureCitation id="7BDB88BA2E3AF14BFEC07147FE6EF852" box="[272,384,1919,1944]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="3" pageNumber="76">Figs. 1, 2</figureCitation>
) with femora weakly dilated from base to apex, laterally compressed basally, then weakly swollen; tibiae (
<figureCitation id="7BDB88BA2E3AF14BFE9D719BFE7FF876" box="[333,401,1955,1980]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="3" pageNumber="76">Fig. 4</figureCitation>
) with inner margin almost straight, serrate; outer margin gently curved; surface smooth but along outer margin with granulate setiferous punctures; tarsi short, segment I slightly longer than II, segment III about twice as long as II, deeply hollowed at apex, last tarsal segment very short, its basal part deeply inside hollow of segment III, surface of segments I-III rugose; underside of segment III with two longitudinal hairy pads.
</paragraph>
<paragraph id="E35F943F2E3DF14CFF17773CFCA9FEAF" blockId="4.[151,1436,151,717]" pageId="4" pageNumber="77">
<emphasis id="D194482D2E3DF14CFF17773CFECBFED7" bold="true" box="[199,293,260,285]" inLineHeading="true" pageId="4" pageNumber="77" reason="1">Female.</emphasis>
Body (
<figureCitation id="7BDB88BA2E3DF14CFEAC773CFE04FED7" box="[380,490,260,285]" captionStart="FIGURES 1–7" captionStartId="2.[151,263,1934,1957]" captionTargetBox="[180,1397,203,1906]" captionTargetId="figure-66@2.[168,1404,193,1912]" captionTargetPageId="2" captionText="FIGURES 1–7. Pycnotarsobrentus inuiae gen. nov. and sp. nov. 1—Male habitus, dorsal view (paratype; color became paler during preserving in ethanol), 2—same individual, ventral view, 3—ditto, head, dorsal view, 4—right fore tibia and tarsus, 5— female habitus, dorsal view (paratype), 6—same individual, ventral view, 7—ditto, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4913385" httpUri="https://zenodo.org/record/4913385/files/figure.png" pageId="4" pageNumber="77">Figs. 5, 6</figureCitation>
) more matte, densely micro-reticulate overall. Prorostrum cylindrical, densely and coarsely punctured, punctation denser but smaller than on mesorostrum. Pronotum more coarsely punctured, except along anterior margin. Elytra more densely punctured.
</paragraph>
<paragraph id="E35F943F2E3DF14CFF177757FD38FE42" blockId="4.[151,1436,151,717]" box="[199,726,367,392]" pageId="4" pageNumber="77">Body length: 8.5–6.2 mm (including rostrum).</paragraph>
</subSubSection>
<subSubSection id="ABFAC7B42E3DF14CFF1777ACFEE6FE3F" pageId="4" pageNumber="77" type="diagnosis">
<paragraph id="E35F943F2E3DF14CFF1777ACFEE6FE3F" blockId="4.[151,1436,151,717]" pageId="4" pageNumber="77">
<emphasis id="D194482D2E3DF14CFF1777ACFE20FE67" bold="true" box="[199,462,404,429]" inLineHeading="true" pageId="4" pageNumber="77" reason="1">Differential diagnosis.</emphasis>
No species of
<taxonomicName id="24E0EFBC2E3DF14CFDA677ACFCE9FE67" authorityName="Semenov-Tian-Schanskij" authorityYear="1892" box="[630,775,404,429]" class="Insecta" family="Brentidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="tribe" tribe="Eremoxenini">Eremoxenini</taxonomicName>
with similar morphological modifications are known from the Oriental region and the new taxon is easily distinguishable from all other Asian
<taxonomicName id="24E0EFBC2E3DF14CFB8F778FFB1EFE1A" authorityName="Semenov-Tian-Schanskij" authorityYear="1892" box="[1119,1264,439,464]" class="Insecta" family="Brentidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="tribe" tribe="Eremoxenini">Eremoxenini</taxonomicName>
by the generic diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="ABFAC7B42E3DF14CFF1777C7FD51FD07" pageId="4" pageNumber="77" type="etymology">
<paragraph id="E35F943F2E3DF14CFF1777C7FE11FDF7" blockId="4.[151,1436,151,717]" pageId="4" pageNumber="77">
<emphasis id="D194482D2E3DF14CFF1777C7FEA0FDD2" bold="true" box="[199,334,511,536]" inLineHeading="true" pageId="4" pageNumber="77" reason="1">Etymology.</emphasis>
Dedicated to Dr. Yoko Inui, one of the collectors, for her invaluable contribution to the knowledge of tree canopy ant communities.
</paragraph>
<paragraph id="E35F943F2E3DF14CFF17747FFD51FD07" blockId="4.[151,1436,151,717]" pageId="4" pageNumber="77">
<emphasis id="D194482D2E3DF14CFF17747FFEA4FD95" box="[199,330,583,608]" inLineHeading="true" pageId="4" pageNumber="77" reason="1">
<emphasis id="D194482D2E3DF14CFF17747FFEADFDAA" bold="true" box="[199,323,583,608]" pageId="4" pageNumber="77">Bionomics</emphasis>
.
</emphasis>
The beetles were found exclusively inside the nests of
<taxonomicName id="24E0EFBC2E3DF14CFC677470FBD2FDAA" authorityName="F. Smith" authorityYear="1857" box="[951,1084,584,608]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="difformis">
<emphasis id="D194482D2E3DF14CFC677470FBD2FDAA" box="[951,1084,584,608]" italics="true" pageId="4" pageNumber="77">C. difformis</emphasis>
</taxonomicName>
in the domatia of the epiphytic ferns (
<figureCitation id="7BDB88BA2E3DF14CFF0E7454FEA7FD4F" box="[222,329,620,645]" captionStart="FIGURES 8–10" captionStartId="3.[151,263,1389,1412]" captionTargetBox="[151,1436,207,1347]" captionTargetId="figure-268@3.[151,1437,207,1347]" captionTargetPageId="3" captionText="FIGURES 8–10. Habitat of Pycnotarsobrentus inuiae gen. nov. and sp. nov. 8—Platycerium crustacea Copel., 9— Lecanopteris ridleyi H. Christ., 10, 11—living male in a domatia of Lecanopteris sp." figureDoi="http://doi.org/10.5281/zenodo.4913391" httpUri="https://zenodo.org/record/4913391/files/figure.png" pageId="4" pageNumber="77">Figs. 8, 9</figureCitation>
). They were observed to walk slowly when the nests were excavated (
<figureCitation id="7BDB88BA2E3DF14CFB8B7454FB40FD4F" box="[1115,1198,620,645]" captionStart="FIGURES 8–10" captionStartId="3.[151,263,1389,1412]" captionTargetBox="[151,1436,207,1347]" captionTargetId="figure-268@3.[151,1437,207,1347]" captionTargetPageId="3" captionText="FIGURES 8–10. Habitat of Pycnotarsobrentus inuiae gen. nov. and sp. nov. 8—Platycerium crustacea Copel., 9— Lecanopteris ridleyi H. Christ., 10, 11—living male in a domatia of Lecanopteris sp." figureDoi="http://doi.org/10.5281/zenodo.4913391" httpUri="https://zenodo.org/record/4913391/files/figure.png" pageId="4" pageNumber="77">Fig. 10</figureCitation>
, 11), and the ants did not pay any attention to them. Ecological studies on the communities of the myrmecophiles in the nests of
<taxonomicName id="24E0EFBC2E3DF14CFA5274A8FF12FD06" authorityName="F. Smith" authorityYear="1857" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="difformis">
<emphasis id="D194482D2E3DF14CFA5274A8FF12FD06" italics="true" pageId="4" pageNumber="77">C. difformis</emphasis>
</taxonomicName>
are in progress by the Japanese authors.
</paragraph>
</subSubSection>
</treatment>
</document>
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