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<document id="2BFB686EB9D409DB6798B132E82603CF" ID-DOI="10.26107/RBZ-2021-0005" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351678" ID-ZooBank="D63943BE-00B0-409E-BCE4-96942D33114E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630206537254" checkinUser="diego" docAuthor="Tran, A. D. &amp; Zettel, H." docDate="2021" docId="610CF11BE00BFFD0FC3223FAFC01F996" docLanguage="en" docName="RBZ-2021-0005.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Ranatra stali Montandon 1905" docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="9D358963E00FFFD7FFE32768FFF2FFE3" masterDocTitle="Taxonomic review of the Ranatra gracilis group sensu Lansbury, 1972 (Nepomorpha: Nepidae), with descriptions of four new species" masterLastPageNumber="70" masterPageNumber="45" pageNumber="49" updateTime="1699185027225" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="24F42A0854F0DAA86CD9D3597C4FCD72">Taxonomic review of the Ranatra gracilis group sensu Lansbury, 1972 (Nepomorpha: Nepidae), with descriptions of four new species</mods:title>
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<mods:namePart id="30752FE50F8750EA5765A78EB0C67751">Tran, A. D.</mods:namePart>
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<mods:namePart id="5754112A876A589245BF43E2197DD7D2">Zettel, H.</mods:namePart>
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<mods:title id="E302DB91823A140AAF661F785D1021B7">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="5B8AF25FE6CBAC15D2ABF256773F4769">
<mods:date id="7C0DBD64113D3C155EC313FD1265E413">2021</mods:date>
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<mods:number id="9C9A3CAC5E8F22E3E9741084AF84A577">2021-02-25</mods:number>
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<mods:number id="179B626C27530358BB8333B4DA8FC25D">69</mods:number>
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<treatment id="610CF11BE00BFFD0FC3223FAFC01F996" ID-DOI="http://doi.org/10.5281/zenodo.5449953" ID-Zenodo-Dep="5449953" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996" httpUri="http://treatment.plazi.org/id/610CF11BE00BFFD0FC3223FAFC01F996" lastPageId="7" lastPageNumber="52" pageId="4" pageNumber="49">
<subSubSection id="A1BF1386E00BFFD3FC3223FAFAC4FB49" box="[977,1334,1169,1194]" pageId="4" pageNumber="49" type="nomenclature">
<paragraph id="E91A400DE00BFFD3FC3223FAFAC4FB49" blockId="4.[977,1334,1169,1226]" box="[977,1334,1169,1194]" pageId="4" pageNumber="49">
<heading id="B252F761E00BFFD3FC3223FAFAC4FB49" bold="true" box="[977,1334,1169,1194]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="49" reason="2">
<taxonomicName id="2EA53B8EE00BFFD3FC3223FAFAC4FB49" ID-CoL="792W8" authority="Montandon, 1905" authorityName="Montandon" authorityYear="1905" box="[977,1334,1169,1194]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE00BFFD3FC3223FAFAC4FB49" bold="true" box="[977,1334,1169,1194]" pageId="4" pageNumber="49">
<emphasis id="DBD19C1FE00BFFD3FC3223FAFB90FB4A" bold="true" box="[977,1122,1169,1193]" italics="true" pageId="4" pageNumber="49">Ranatra stali</emphasis>
<bibRefCitation id="8D343DFCE00BFFD3FB8F23FAFAC4FB49" author="Montandon AL" box="[1132,1334,1170,1194]" pageId="4" pageNumber="49" pagination="389 - 398" refId="ref20192" refString="Montandon AL (1905) Especes nouvelles ou peu connues du genre Ranatra. Buletinul Societat ̦ ii de S ̦ ciint ̦ e din Bucures ̦ ci, Romania, 14: 389 - 398." type="journal article" year="1905">Montandon, 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E00BFFD3FBDF23DAFB3BFB29" box="[1084,1225,1202,1226]" pageId="4" pageNumber="49" type="description">
<paragraph id="E91A400DE00BFFD3FBDF23DAFB3BFB29" blockId="4.[977,1334,1169,1226]" box="[1084,1225,1202,1226]" pageId="4" pageNumber="49">
(
<figureCitation id="719E5C88E00BFFD3FBA723DAFB57FB29" box="[1092,1189,1202,1226]" captionStart="Fig" captionStartId="5.[113,148,2000,2022]" captionTargetBox="[113,1451,253,1984]" captionTargetId="figure-10@5.[113,1451,253,1984]" captionTargetPageId="5" captionText="Fig. 1. Habitus of Ranatra spp., all males. A, R. stali Montandon, 1905; B, R. bisaya, new species; C, R. bendanilloi, new species; D, R. brevicauda, new species; E, R. pangantihoni, new species (all of same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351680" httpUri="https://zenodo.org/record/5351680/files/figure.png" pageId="4" pageNumber="49">Figs. 1A</figureCitation>
,
<figureCitation id="719E5C88E00BFFD3FB5023DAFB33FB29" box="[1203,1217,1202,1226]" captionStart="Fig" captionStartId="6.[136,171,1985,2007]" captionTargetBox="[141,1469,259,1965]" captionTargetId="figure-7@6.[136,1476,236,1970]" captionTargetPageId="6" captionText="Fig. 2. Ranatra stali Montandon, 1905, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, operculum; H, paramere; lateral view; I, apical part of paramere, inner view. (B, C same scale; D, E same scale; F, G same scale, H, I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351682" httpUri="https://zenodo.org/record/5351682/files/figure.png" pageId="4" pageNumber="49">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E00BFFD3FCA02399FA50FAA3" pageId="4" pageNumber="49" type="reference_group">
<paragraph id="E91A400DE00BFFD3FCA02399FAB9FAE5" blockId="4.[835,1473,1264,1344]" box="[835,1355,1264,1286]" pageId="4" pageNumber="49">
<treatmentCitationGroup id="C9B56723E00BFFD3FCA02399FAB9FAE5" box="[835,1355,1264,1286]" pageId="4" pageNumber="49">
<treatmentCitation id="6804661CE00BFFD3FCA02399FB78FAE5" author="Stal C" box="[835,1162,1264,1286]" page="707" pageId="4" pageNumber="49" year="1870">
<taxonomicName id="2EA53B8EE00BFFD3FCA02399FB78FAE5" ID-CoL="6X37T" authority="Stal, 1870: 707" authorityName="Stal" authorityPageNumber="707" authorityYear="1870" box="[835,1162,1264,1286]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="parmata">
<emphasis id="DBD19C1FE00BFFD3FCA02399FC19FAE5" box="[835,1003,1265,1286]" italics="true" pageId="4" pageNumber="49">Ranatra parmata</emphasis>
<bibRefCitation id="8D343DFCE00BFFD3FC162398FB78FAE5" author="Stal C" box="[1013,1162,1264,1286]" pageId="4" pageNumber="49" pagination="607 - 776" refId="ref20830" refString="Stal C (1870) Hemiptera insularum Philippinarum. Ofversigt af Kungliga Vetenskaps-Akademiens Forhandlingar, 27 (7): 607 - 776." type="journal article" year="1870">Stål, 1870: 707</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E91A400DE00BFFD3FCA02266FA50FAA3" blockId="4.[835,1473,1264,1344]" pageId="4" pageNumber="49">
<treatmentCitationGroup id="C9B56723E00BFFD3FCA02266FA50FAA3" pageId="4" pageNumber="49">
<treatmentCitation id="6804661CE00BFFD3FCA02266FB39FAC1" author="Montandon AL" box="[835,1227,1293,1315]" page="391" pageId="4" pageNumber="49" year="1905">
<taxonomicName id="2EA53B8EE00BFFD3FCA02266FB39FAC1" ID-CoL="792W8" authority="Montandon, 1905: 390 - 391" authorityName="Montandon" authorityPageNumber="390 - 391" authorityYear="1905" box="[835,1227,1293,1315]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE00BFFD3FCA02266FC32FAC1" box="[835,960,1293,1315]" italics="true" pageId="4" pageNumber="49">Ranatra stali</emphasis>
<bibRefCitation id="8D343DFCE00BFFD3FC202265FB39FAC1" author="Montandon AL" box="[963,1227,1293,1315]" pageId="4" pageNumber="49" pagination="389 - 398" refId="ref20192" refString="Montandon AL (1905) Especes nouvelles ou peu connues du genre Ranatra. Buletinul Societat ̦ ii de S ̦ ciint ̦ e din Bucures ̦ ci, Romania, 14: 389 - 398." type="journal article" year="1905">Montandon, 1905: 390391</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="361EFEAFE00BFFD3FB342265FAF0FAC1" box="[1239,1282,1293,1314]" pageId="4" pageNumber="49">type</typeStatus>
locality: Mindanao,
<collectingCountry id="91B2009DE00BFFD3FC8A2242FC28FAA3" box="[873,986,1322,1344]" name="Philippines" pageId="4" pageNumber="49">Philippines</collectingCountry>
). —
<treatmentCitation id="6804661CE00BFFD3FBE82242FAF7FAA3" author="Lansbury I" box="[1035,1285,1322,1344]" page="296" pageId="4" pageNumber="49" year="1972">
<bibRefCitation id="8D343DFCE00BFFD3FBE82242FAF7FAA3" author="Lansbury I" box="[1035,1285,1322,1344]" pageId="4" pageNumber="49" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansbury, 1972: 296298</bibRefCitation>
</treatmentCitation>
(redescription).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E00BFFD3FCA0220DFBB3F87D" pageId="4" pageNumber="49" type="materials_examined">
<paragraph id="E91A400DE00BFFD3FCA0220DFB3AF9DE" blockId="4.[835,1474,1381,1597]" pageId="4" pageNumber="49">
<emphasis id="DBD19C1FE00BFFD3FCA0220DFB84FA9E" bold="true" box="[835,1142,1381,1405]" pageId="4" pageNumber="49">Type material examined.</emphasis>
<materialsCitation id="59CD4A50E00BFFD3FB61220DFB3AF9DE" collectionCode="NRMS" county="Haglund" location="Type" municipality="Typus" pageId="4" pageNumber="49" specimenCount="1" stateProvince="Stockholm" typeStatus="lectotype">
<emphasis id="DBD19C1FE00BFFD3FB61220DFB0BFA9E" bold="true" box="[1154,1273,1381,1405]" pageId="4" pageNumber="49">
<typeStatus id="361EFEAFE00BFFD3FB61220DFB0BFA9E" box="[1154,1273,1381,1405]" pageId="4" pageNumber="49" type="lectotype">Lectotype</typeStatus>
</emphasis>
(female,
<collectionCode id="8FB4D8C8E00BFFD3FA8E220DFA4FFA9E" box="[1389,1469,1381,1405]" pageId="4" pageNumber="49">NRMS</collectionCode>
, designated by
<bibRefCitation id="8D343DFCE00BFFD3FC0B22EDFB6EFA7E" author="Lansbury I" box="[1000,1180,1413,1437]" pageId="4" pageNumber="49" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansbury, 1972</bibRefCitation>
): “Mindanao.”, “collectio\
<collectingCounty id="007B3881E00BFFD3FCA722CDFC5AFA5E" box="[836,936,1445,1469]" pageId="4" pageNumber="49">Haglund</collectingCounty>
”, “Type.”, “
<collectingMunicipality id="097EDA77E00BFFD3FBD622CDFB8FFA5E" box="[1077,1149,1445,1469]" pageId="4" pageNumber="49">Typus</collectingMunicipality>
” [red label], “
<taxonomicName id="2EA53B8EE00BFFD3FAC722CDFA8EFA5E" authorityName="Fabricius" authorityYear="1790" box="[1316,1404,1445,1469]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">Ranatra</taxonomicName>
\
<taxonomicName id="2EA53B8EE00BFFD3FA6C22CDFC4CFA3E" authority="Montandon" authorityName="Montandon" authorityYear="1905" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="stali">Ståli Montandon</taxonomicName>
\
<location id="EC7A16D6E00BFFD3FC2E22AEFBF6FA3D" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996:EC7A16D6E00BFFD3FC2E22AEFBF6FA3D" box="[973,1028,1478,1502]" county="Haglund" municipality="Typus" name="Type" pageId="4" pageNumber="49" stateProvince="Stockholm">Type</location>
1903.” [handwritten, last line in red ink], “480\ 62” [red label, last line handwritten], “51\ 67” [red label, last line handwritten], “
<location id="EC7A16D6E00BFFD3FB62216DFAD5F9FE" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996:EC7A16D6E00BFFD3FB62216DFAD5F9FE" box="[1153,1319,1541,1565]" county="Haglund" municipality="Typus" name="Naturhistoriska" pageId="4" pageNumber="49" stateProvince="Stockholm">Naturhistoriska</location>
\
<location id="EC7A16D6E00BFFD3FAD4216DFA48F9FE" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996:EC7A16D6E00BFFD3FAD4216DFA48F9FE" box="[1335,1466,1541,1565]" county="Haglund" municipality="Typus" name="Rijksmuseet" pageId="4" pageNumber="49" stateProvince="Stockholm">Rijksmuseet</location>
\
<collectingRegion id="2B618EEFE00BFFD3FCA0214DFC44F9DE" box="[835,950,1573,1597]" country="Sweden" name="Stockholm" pageId="4" pageNumber="49">Stockholm</collectingRegion>
\
<location id="EC7A16D6E00BFFD3FC24214EFC0CF9DE" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996:EC7A16D6E00BFFD3FC24214EFC0CF9DE" box="[967,1022,1574,1597]" county="Haglund" municipality="Typus" name="Loan" pageId="4" pageNumber="49" stateProvince="Stockholm">Loan</location>
no 251/01”, “14”.
</materialsCitation>
</paragraph>
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<emphasis id="DBD19C1FE00BFFD3FCA0210DFB5BF99E" bold="true" box="[835,1193,1637,1661]" pageId="4" pageNumber="49">
<collectingRegion id="2B618EEFE00BFFD3FCA0210DFC4FF99E" box="[835,957,1637,1661]" pageId="4" pageNumber="49">Additional</collectingRegion>
material examined.
</emphasis>
<materialsCitation id="59CD4A50E00BFFD3FB51210DFB1BF97E" collectionCode="NHMW, UPLB, USC, ZCW, ZRC" country="Philippines" location="Mindanao" pageId="4" pageNumber="49" specimenCount="1" stateProvince="Mindanao">
<collectingCountry id="91B2009DE00BFFD3FB51210DFAA2F99E" box="[1202,1360,1637,1661]" name="Philippines" pageId="4" pageNumber="49">PHILIPPINES</collectingCountry>
(
<collectionCode id="8FB4D8C8E00BFFD3FA81210DFA4CF99E" box="[1378,1470,1637,1661]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="49" type="Museum">NHMW</collectionCode>
,
<collectionCode id="8FB4D8C8E00BFFD3FCA021EEFC78F97E" box="[835,906,1670,1693]" country="Philippines" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="4" pageNumber="49">UPLB</collectionCode>
,
<collectionCode id="8FB4D8C8E00BFFD3FC7521EDFC3EF97E" box="[918,972,1669,1693]" pageId="4" pageNumber="49">USC</collectionCode>
,
<collectionCode id="8FB4D8C8E00BFFD3FC3A21EDFBE5F97E" box="[985,1047,1669,1693]" pageId="4" pageNumber="49">ZCW</collectionCode>
,
<collectionCode id="8FB4D8C8E00BFFD3FBC721EDFBAFF97E" box="[1060,1117,1669,1693]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="49">ZRC</collectionCode>
):
<collectingRegion id="2B618EEFE00BFFD3FB9321EDFB1BF97E" box="[1136,1257,1669,1693]" pageId="4" pageNumber="49">
<emphasis id="DBD19C1FE00BFFD3FB9321EDFB1BF97E" bold="true" box="[1136,1257,1669,1693]" pageId="4" pageNumber="49">Mindanao</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="DBD19C1FE00BFFD3FB0A21EDFB1CF97E" bold="true" box="[1257,1262,1669,1693]" pageId="4" pageNumber="49">:</emphasis>
<materialsCitation id="59CD4A50E00BFFD3FB1521EDFA51F95E" collectorName="Malaybalay &amp; Kaamulan Site" country="Philippines" elevation="650" location="Bukidnon Province" pageId="4" pageNumber="49" specimenCount="4" specimenCount-male="4" stateProvince="Bukidnon">
<specimenCount id="FFA38B84E00BFFD3FB1521EDFABFF97E" box="[1270,1357,1669,1693]" pageId="4" pageNumber="49" type="male">4 males</specimenCount>
,
<collectingRegion id="2B618EEFE00BFFD3FABA21EDFC5EF95E" country="Philippines" name="Bukidnon" pageId="4" pageNumber="49">Bukidnon Province</collectingRegion>
,
<collectorName id="445025DBE00BFFD3FC5F21CDFBB7F95E" box="[956,1093,1701,1725]" pageId="4" pageNumber="49">Malaybalay</collectorName>
,
<collectorName id="445025DBE00BFFD3FBB621CDFAF6F95E" box="[1109,1284,1701,1725]" pageId="4" pageNumber="49">Kaamulan Site</collectorName>
,
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<quantity id="2E5DEDE8E00BFFD3FAF621CDFAADF95E" box="[1301,1375,1701,1725]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="49" unit="m" value="650.0">650 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
;
<materialsCitation id="59CD4A50E00BFFD3FA5621CDFA30F91E" country="Philippines" county="San Francisco" elevation="54" latitude="8.456667" location="Tagkunayai creek" longLatPrecision="21" longitude="125.97" municipality="Bayogan" pageId="4" pageNumber="49" specimenCount="2" specimenCount-female="2" stateProvince="Agusan del Sur">
<specimenCount id="FFA38B84E00BFFD3FA5621CDFC68F93E" pageId="4" pageNumber="49" type="female">2 females</specimenCount>
,
<collectingRegion id="2B618EEFE00BFFD3FC4721ADFB42F93E" box="[932,1200,1733,1757]" country="Philippines" name="Agusan del Sur" pageId="4" pageNumber="49">Agusan del Sur Province</collectingRegion>
,
<collectingCounty id="007B3881E00BFFD3FB5921ADFAA1F93E" box="[1210,1363,1733,1757]" pageId="4" pageNumber="49">San Francisco</collectingCounty>
,
<collectingMunicipality id="097EDA77E00BFFD3FABE21AEFA4CF93D" box="[1373,1470,1734,1758]" pageId="4" pageNumber="49">Bayogan</collectingMunicipality>
,
<location id="EC7A16D6E00BFFD3FCA0218DFBE2F91E" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996:EC7A16D6E00BFFD3FCA0218DFBE2F91E" box="[835,1040,1765,1789]" country="Philippines" county="San Francisco" latitude="8.456667" longLatPrecision="21" longitude="125.97" municipality="Bayogan" name="Tagkunayai creek" pageId="4" pageNumber="49" stateProvince="Agusan del Sur">Tagkunayai creek</location>
,
<geoCoordinate id="8C9126CAE00BFFD3FBFC218CFB6AF91E" box="[1055,1176,1764,1789]" degrees="8" direction="north" minutes="27" orientation="latitude" pageId="4" pageNumber="49" precision="15" seconds="24" value="8.456667">8°2724″N</geoCoordinate>
,
<geoCoordinate id="8C9126CAE00BFFD3FB44218CFACBF91E" box="[1191,1337,1764,1789]" degrees="125" direction="east" minutes="58" orientation="longitude" pageId="4" pageNumber="49" precision="15" seconds="12" value="125.97">125°5812″E</geoCoordinate>
,
<elevation id="6288A73EE00BFFD3FAAB218DFA30F91E" box="[1352,1474,1765,1789]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="49" unit="m" value="54.0">
<quantity id="2E5DEDE8E00BFFD3FAAB218DFA71F91E" box="[1352,1411,1765,1789]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="49" unit="m" value="54.0">54 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
<materialsCitation id="59CD4A50E00BFFD3FCA7206DFB5CF8DE" collectorName="Veriato &amp; El Amigo" country="Philippines" location="Veriato Falls" pageId="4" pageNumber="49" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Samar">
<emphasis id="DBD19C1FE00BFFD3FCA7206DFC69F8FE" bold="true" box="[836,923,1797,1821]" pageId="4" pageNumber="49">
<collectingRegion id="2B618EEFE00BFFD3FCA7206DFC65F8FE" box="[836,919,1797,1821]" country="Philippines" name="Samar" pageId="4" pageNumber="49">Samar</collectingRegion>
:
</emphasis>
<specimenCount id="FFA38B84E00BFFD3FC45206DFBF1F8FE" box="[934,1027,1797,1821]" pageId="4" pageNumber="49" type="male">4 males</specimenCount>
,
<specimenCount id="FFA38B84E00BFFD3FBF0206DFB77F8FE" box="[1043,1157,1797,1821]" pageId="4" pageNumber="49" type="female">2 females</specimenCount>
,
<collectingRegion id="2B618EEFE00BFFD3FB76206DFA4FF8FE" box="[1173,1469,1797,1821]" country="Philippines" name="Northern Samar" pageId="4" pageNumber="49">Northern Samar Province</collectingRegion>
,
<collectorName id="445025DBE00BFFD3FCA7204DFC65F8DE" box="[836,919,1829,1853]" pageId="4" pageNumber="49">Veriato</collectorName>
,
<collectorName id="445025DBE00BFFD3FC47204DFBE3F8DE" box="[932,1041,1829,1853]" pageId="4" pageNumber="49">El Amigo</collectorName>
,
<location id="EC7A16D6E00BFFD3FBFD204DFB5CF8DE" LSID="urn:lsid:plazi:treatment:610CF11BE00BFFD0FC3223FAFC01F996:EC7A16D6E00BFFD3FBFD204DFB5CF8DE" box="[1054,1198,1829,1853]" country="Philippines" name="Veriato Falls" pageId="4" pageNumber="49" stateProvince="Samar">Veriato Falls</location>
</materialsCitation>
;
<materialsCitation id="59CD4A50E00BFFD3FB58204DFB1DF8BE" collectorName="San Joaquin" country="Philippines" location="Northern Samar Province" pageId="4" pageNumber="49" specimenCount="1" specimenCount-male="1" stateProvince="Northern Samar">
<specimenCount id="FFA38B84E00BFFD3FB58204DFAFAF8DE" box="[1211,1288,1829,1853]" pageId="4" pageNumber="49" type="male">1 male</specimenCount>
,
<collectingRegion id="2B618EEFE00BFFD3FAF6204DFC57F8BE" country="Philippines" name="Northern Samar" pageId="4" pageNumber="49">Northern Samar Province</collectingRegion>
,
<collectorName id="445025DBE00BFFD3FC52202DFBCAF8BE" box="[945,1080,1861,1885]" pageId="4" pageNumber="49">San Joaquin</collectorName>
, stream near sea
</materialsCitation>
;
<materialsCitation id="59CD4A50E00BFFD3FB1F202DFAA1F89E" collectorName="Western &amp; E Basey &amp; Sohoton NP" country="Philippines" location="Samar Province" pageId="4" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Samar">
<specimenCount id="FFA38B84E00BFFD3FB1F202DFAAEF8BE" box="[1276,1372,1861,1885]" pageId="4" pageNumber="49" type="female">1 female</specimenCount>
,
<collectorName id="445025DBE00BFFD3FA8A202EFA30F8BE" box="[1385,1474,1862,1885]" pageId="4" pageNumber="49">Western</collectorName>
<collectingRegion id="2B618EEFE00BFFD3FCA0200DFC0BF89E" box="[835,1017,1893,1917]" country="Philippines" name="Samar" pageId="4" pageNumber="49">Samar Province</collectingRegion>
,
<collectorName id="445025DBE00BFFD3FBEB200EFB98F89D" box="[1032,1130,1894,1918]" pageId="4" pageNumber="49">E Basey</collectorName>
,
<collectorName id="445025DBE00BFFD3FB9A200DFAF4F89E" box="[1145,1286,1893,1917]" pageId="4" pageNumber="49">Sohoton NP</collectorName>
, creek
</materialsCitation>
;
<materialsCitation id="59CD4A50E00BFFD3FA80200DFBCCF87D" collectorName="Western &amp; Basey" country="Philippines" location="Samar" pageId="4" pageNumber="49" specimenCount="2" specimenCount-male="2" stateProvince="Samar">
<specimenCount id="FFA38B84E00BFFD3FA80200DFA4CF89E" box="[1379,1470,1893,1917]" pageId="4" pageNumber="49" type="male">2 males</specimenCount>
,
<collectorName id="445025DBE00BFFD3FCA720EEFC6FF87E" box="[836,925,1926,1949]" pageId="4" pageNumber="49">Western</collectorName>
<collectingRegion id="2B618EEFE00BFFD3FC4620EDFC1EF87E" box="[933,1004,1925,1949]" country="Philippines" name="Samar" pageId="4" pageNumber="49">Samar</collectingRegion>
,
<collectorName id="445025DBE00BFFD3FC1A20EEFBCCF87D" box="[1017,1086,1926,1950]" pageId="4" pageNumber="49">Basey</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E00BFFD0FCA020ADFEC1FA76" lastPageId="7" lastPageNumber="52" pageId="4" pageNumber="49" type="description">
<paragraph id="E91A400DE00BFFD3FCA020ADFBB8F81E" blockId="4.[835,1474,1989,2045]" pageId="4" pageNumber="49">
<emphasis id="DBD19C1FE00BFFD3FCA020ADFB96F83E" bold="true" box="[835,1124,1989,2013]" pageId="4" pageNumber="49">
Description of
<typeStatus id="361EFEAFE00BFFD3FC1620ADFBADF83E" box="[1013,1119,1989,2013]" pageId="4" pageNumber="49" type="lectotype">lectotype</typeStatus>
.
</emphasis>
Colour uniformly light brown, probably faded by light.
</paragraph>
<caption id="BDDA1085E00AFFD2FF9220B8FC9CF7E0" ID-DOI="http://doi.org/10.5281/zenodo.5351680" ID-Zenodo-Dep="5351680" httpUri="https://zenodo.org/record/5351680/files/figure.png" pageId="5" pageNumber="50" startId="5.[113,148,2000,2022]" targetBox="[113,1451,253,1984]" targetPageId="5">
<paragraph id="E91A400DE00AFFD2FF9220B8FC9CF7E0" blockId="5.[113,1452,2000,2051]" pageId="5" pageNumber="50">
Fig. 1. Habitus of
<taxonomicName id="2EA53B8EE00AFFD2FEC620B9FE57F806" box="[293,421,2000,2022]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="DBD19C1FE00AFFD2FEC620B9FE86F805" box="[293,372,2001,2022]" italics="true" pageId="5" pageNumber="50">Ranatra</emphasis>
spp.
</taxonomicName>
, all males. A,
<taxonomicName id="2EA53B8EE00AFFD2FDD020B9FCC2F805" authority="Montandon, 1905" authorityName="Montandon" authorityYear="1905" box="[563,816,2000,2022]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE00AFFD2FDD020B9FD8AF806" box="[563,632,2000,2022]" italics="true" pageId="5" pageNumber="50">R. stali</emphasis>
<bibRefCitation id="8D343DFCE00AFFD2FD6220B8FCC2F805" author="Montandon AL" box="[641,816,2000,2022]" pageId="5" pageNumber="50" pagination="389 - 398" refId="ref20192" refString="Montandon AL (1905) Especes nouvelles ou peu connues du genre Ranatra. Buletinul Societat ̦ ii de S ̦ ciint ̦ e din Bucures ̦ ci, Romania, 14: 389 - 398." type="journal article" year="1905">Montandon, 1905</bibRefCitation>
</taxonomicName>
; B,
<taxonomicName id="2EA53B8EE00AFFD2FCB420B9FC40F806" authority="Tran &amp; Zettel, 2021" authorityName="Tran &amp; Zettel" authorityYear="2021" box="[855,946,2000,2022]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="bisaya" status="sp. nov.">
<emphasis id="DBD19C1FE00AFFD2FCB420B9FC40F806" box="[855,946,2000,2022]" italics="true" pageId="5" pageNumber="50">R. bisaya</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C0E22164E00AFFD2FC5C20B8FBC4F805" box="[959,1078,2000,2022]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
; C,
<taxonomicName id="2EA53B8EE00AFFD2FBBC20B9FB1AF806" authority="Tran &amp; Zettel, 2021" authorityName="Tran &amp; Zettel" authorityYear="2021" box="[1119,1256,2000,2022]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="bendanilloi" status="sp. nov.">
<emphasis id="DBD19C1FE00AFFD2FBBC20B9FB1AF806" box="[1119,1256,2000,2022]" italics="true" pageId="5" pageNumber="50">R. bendanilloi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C0E22164E00AFFD2FB1620B8FA9FF805" box="[1269,1389,2000,2022]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
; D,
<taxonomicName id="2EA53B8EE00AFFD2FA7420B9FF2FF7E1" authority="Tran &amp; Zettel, 2021" authorityName="Tran &amp; Zettel" authorityYear="2021" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="brevicauda" status="sp. nov.">
<emphasis id="DBD19C1FE00AFFD2FA7420B9FF2FF7E1" italics="true" pageId="5" pageNumber="50">R. brevicauda</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C0E22164E00AFFD2FF082085FE91F7E0" box="[235,355,2029,2051]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
; E,
<taxonomicName id="2EA53B8EE00AFFD2FE682085FDDAF7E1" authority="Tran &amp; Zettel, 2021" authorityName="Tran &amp; Zettel" authorityYear="2021" box="[395,552,2029,2050]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="pangantihoni" status="sp. nov.">
<emphasis id="DBD19C1FE00AFFD2FE682085FDDAF7E1" box="[395,552,2029,2050]" italics="true" pageId="5" pageNumber="50">R. pangantihoni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C0E22164E00AFFD2FDD62085FD58F7E0" box="[565,682,2029,2051]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
(all of same scale).
</paragraph>
</caption>
<caption id="BDDA1085E009FFD1FF6B20A9FD9AF7F2" ID-DOI="http://doi.org/10.5281/zenodo.5351682" ID-Zenodo-Dep="5351682" httpUri="https://zenodo.org/record/5351682/files/figure.png" pageId="6" pageNumber="51" startId="6.[136,171,1985,2007]" targetBox="[141,1469,259,1965]" targetPageId="6">
<paragraph id="E91A400DE009FFD1FF6B20A9FD9AF7F2" blockId="6.[136,1474,1985,2065]" pageId="6" pageNumber="51">
Fig. 2.
<taxonomicName id="2EA53B8EE009FFD1FF3220AAFDFBF834" authority="Montandon, 1905" authorityName="Montandon" authorityYear="1905" box="[209,521,1985,2007]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE009FFD1FF3220AAFEA3F835" box="[209,337,1985,2007]" italics="true" pageId="6" pageNumber="51">Ranatra stali</emphasis>
<bibRefCitation id="8D343DFCE009FFD1FEB920A9FDFBF834" author="Montandon AL" box="[346,521,1985,2007]" pageId="6" pageNumber="51" pagination="389 - 398" refId="ref20192" refString="Montandon AL (1905) Especes nouvelles ou peu connues du genre Ranatra. Buletinul Societat ̦ ii de S ̦ ciint ̦ e din Bucures ̦ ci, Romania, 14: 389 - 398." type="journal article" year="1905">Montandon, 1905</bibRefCitation>
</taxonomicName>
, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, operculum; H, paramere; lateral view; I, apical part of paramere, inner view. (B, C same scale; D, E same scale; F, G same scale, H, I same scale).
</paragraph>
</caption>
<paragraph id="E91A400DE008FFD0FF9227D6FEA5FDD6" blockId="7.[113,753,189,565]" pageId="7" pageNumber="52">Measurements. Body length 39.6; respiratory siphon&gt; 9.0 (apex broken); width of head 3.27; width of eye 1.05; interocular width 1.17; rostral segment 3: 0.72, segment 4: 0.52; anterior width of pronotum 2.75; posterior width of pronotum 3.40; length of pronotum 9.90; length of anterior lobe of pronotum 7.6; length of posterior lobe 3.5. Length of distal part of fore femur 4.0; proximal part of fore femur 6.1. Maximum width of fore femur at basal part 0.94, at median tooth 0.92. Lengths of leg segments: fore leg: coxa 6.7, femur 10.1, tibia 3.9, tarsus 0.8; middle leg: femur 16.8, tibia 15.6, tarsus broken off; hind leg: femur 16.7, tibia 19.6, tarsus 2.4.</paragraph>
<paragraph id="E91A400DE008FFD0FF922535FDF3FB76" blockId="7.[113,754,605,1173]" pageId="7" pageNumber="52">
Structural characteristics. Clypeus as high as lorum, clypeoloral sulcus shallow; vertex with prominent, high tubercle. Eye slightly narrower than interocular space. Anterior collar of pronotum raised to paired tubercles. Anterior lobe of pronotum 2.2× as long as posterior lobe, sublateral swellings caudally on pronotum distinct. Mesoscutellum 1.9× as long as broad, dorsal surface smooth, with paired grooves on caudal half. Prosternum with paired broad, shallow longitudinal depressions separated by low and blunt median carina; median carina rather indistinct on posterior half. Mesosternum invisible in
<typeStatus id="361EFEAFE008FFD0FD6324F5FD1EFC56" box="[640,748,925,949]" pageId="7" pageNumber="52" type="lectotype">lectotype</typeStatus>
. Metasternum anteromedially depressed, posterior twothirds with low median carina, posterior margin angularly emarginated, with sublateral grooves. Smallest distance between hind coxae ca. 0.8× distance between middle coxae. Hemelytra posteriorly reaching to anterior third of abdominal tergum VI. Hind femur not reaching operculum. Operculum projecting beyond apex of abdomen.
</paragraph>
<paragraph id="E91A400DE008FFD0FF9223D5FEC1FA76" blockId="7.[113,753,1213,1429]" pageId="7" pageNumber="52">
<emphasis id="DBD19C1FE008FFD0FF9223D5FDD3FB36" bold="true" box="[113,545,1213,1237]" pageId="7" pageNumber="52">Measurement of additional specimens.</emphasis>
Males: body length 3538; length of siphon 12.915.9; width of head 3.003.41; interocular width 1.041.27; width of eye 0.971.07; pronotal length 9.12, anterior pronotal length 7.3; posterior pronotal length 3.0; anterior width of pronotum 2.49; posterior width of pronotum 2.91. Females: body length 3943; length of siphon 14.115.2.
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E008FFD0FF9222D5FB44FF16" pageId="7" pageNumber="52" type="diagnosis">
<paragraph id="E91A400DE008FFD0FF9222D5FB44FF16" blockId="7.[113,753,1469,2069]" lastBlockId="7.[813,1452,189,245]" pageId="7" pageNumber="52">
<emphasis id="DBD19C1FE008FFD0FF9222D5FF1FFA36" bold="true" box="[113,237,1469,1493]" pageId="7" pageNumber="52">Diagnosis.</emphasis>
Body length: males 3538, females 3943; ratio of siphon length: body length ca. 0.350.42; lorum higher than clypeus and with a nodule dorsally; ratio of eye width: interocular width ca. 0.841.00; pronotal length ca. 1.341.52× fore coxa length; hemelytra reaching about anterior third of abdominal tergum VI; posterior margin of metasternum angularly emarginated, with sublateral grooves (
<figureCitation id="719E5C88E008FFD0FF9A21F5FF39F956" box="[121,203,1693,1717]" captionStart="Fig" captionStartId="6.[136,171,1985,2007]" captionTargetBox="[141,1469,259,1965]" captionTargetId="figure-7@6.[136,1476,236,1970]" captionTargetPageId="6" captionText="Fig. 2. Ranatra stali Montandon, 1905, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, operculum; H, paramere; lateral view; I, apical part of paramere, inner view. (B, C same scale; D, E same scale; F, G same scale, H, I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351682" httpUri="https://zenodo.org/record/5351682/files/figure.png" pageId="7" pageNumber="52">Fig. 2F</figureCitation>
); flexor side of fore femur with a tooth and a carina at ca. 0.4 distal part, pre-apical part with a pair of small teeth on either side of flexor side (
<figureCitation id="719E5C88E008FFD0FE4B21B5FDEFF916" box="[424,541,1757,1781]" captionStart="Fig" captionStartId="6.[136,171,1985,2007]" captionTargetBox="[141,1469,259,1965]" captionTargetId="figure-7@6.[136,1476,236,1970]" captionTargetPageId="6" captionText="Fig. 2. Ranatra stali Montandon, 1905, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, operculum; H, paramere; lateral view; I, apical part of paramere, inner view. (B, C same scale; D, E same scale; F, G same scale, H, I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351682" httpUri="https://zenodo.org/record/5351682/files/figure.png" pageId="7" pageNumber="52">Fig. 2D, E</figureCitation>
); hind femur, when folded back parallel to body at most reaching posterior margin of abdominal sternum VI (in males) or just surpassing midlength of sternum VI (in females). Male paramere: thickest at basal third, constricted at ca. distal third, distal third bent downwards; ventral margin gradually tapering from middle part towards the constricted part, then followed by a small, obtuse, trapezoidal process bearing tuft of setae; from inner view, pre-apical process appearing sub-rectangular due to a small round projection shorter than tuft of setae; apical hook thick, curved with rounded tip; gap between hook and pre-apical hook narrow (
<figureCitation id="719E5C88E008FFD0FBD927B5FB5BFF16" box="[1082,1193,221,245]" captionStart="Fig" captionStartId="6.[136,171,1985,2007]" captionTargetBox="[141,1469,259,1965]" captionTargetId="figure-7@6.[136,1476,236,1970]" captionTargetPageId="6" captionText="Fig. 2. Ranatra stali Montandon, 1905, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, operculum; H, paramere; lateral view; I, apical part of paramere, inner view. (B, C same scale; D, E same scale; F, G same scale, H, I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351682" httpUri="https://zenodo.org/record/5351682/files/figure.png" pageId="7" pageNumber="52">Fig. 2H, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E008FFD0FCCE2675FBA6FA16" pageId="7" pageNumber="52" type="discussion">
<paragraph id="E91A400DE008FFD0FCCE2675FA5EFE16" blockId="7.[812,1452,285,501]" pageId="7" pageNumber="52">
<emphasis id="DBD19C1FE008FFD0FCCE2675FC53FED6" bold="true" box="[813,929,285,309]" pageId="7" pageNumber="52">Remarks.</emphasis>
<bibRefCitation id="8D343DFCE008FFD0FC4E2675FB74FED6" author="Montandon AL" box="[941,1158,285,309]" pageId="7" pageNumber="52" pagination="389 - 398" refId="ref20192" refString="Montandon AL (1905) Especes nouvelles ou peu connues du genre Ranatra. Buletinul Societat ̦ ii de S ̦ ciint ̦ e din Bucures ̦ ci, Romania, 14: 389 - 398." type="journal article" year="1905">Montandon (1905)</bibRefCitation>
noted that the “
<taxonomicName id="2EA53B8EE008FFD0FAAC2676FC75FEB6" authorityName="Stal" authorityYear="1870" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="parmata">
<emphasis id="DBD19C1FE008FFD0FAAC2676FC75FEB6" italics="true" pageId="7" pageNumber="52">Ranatra parmata</emphasis>
</taxonomicName>
” from the
<collectingCountry id="91B2009DE008FFD0FC1C2655FB85FEB6" box="[1023,1143,317,341]" name="Philippines" pageId="7" pageNumber="52">Philippines</collectingCountry>
reported by
<bibRefCitation id="8D343DFCE008FFD0FAE12655FA88FEB6" author="Stal C" box="[1282,1402,317,341]" pageId="7" pageNumber="52" pagination="607 - 776" refId="ref20830" refString="Stal C (1870) Hemiptera insularum Philippinarum. Ofversigt af Kungliga Vetenskaps-Akademiens Forhandlingar, 27 (7): 607 - 776." type="journal article" year="1870">Stål (1870)</bibRefCitation>
was actually a new species and described it as
<taxonomicName id="2EA53B8EE008FFD0FB112636FA8CFE96" authorityName="Montandon" authorityYear="1905" box="[1266,1406,349,373]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FB112636FA8CFE96" box="[1266,1406,349,373]" italics="true" pageId="7" pageNumber="52">Ranatra stali</emphasis>
</taxonomicName>
. He also included
<specimenCount id="FFA38B84E008FFD0FC582616FBA6FE76" box="[955,1108,381,405]" pageId="7" pageNumber="52" type="generic">two specimens</specimenCount>
from Borneo and from
<collectingCountry id="91B2009DE008FFD0FAA52615FA5EFE76" box="[1350,1452,381,405]" name="Indonesia" pageId="7" pageNumber="52">Moluccas</collectingCountry>
as
<taxonomicName id="2EA53B8EE008FFD0FCAE26F6FC12FE56" authorityName="Montandon" authorityYear="1905" box="[845,992,413,437]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FCAE26F6FC12FE56" box="[845,992,413,437]" italics="true" pageId="7" pageNumber="52">Ranatra stali</emphasis>
</taxonomicName>
, but subsequently, he separated those as another taxon,
<taxonomicName id="2EA53B8EE008FFD0FC3326D6FB69FE36" authorityName="Montandon" authorityYear="1910" box="[976,1179,445,469]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="spinifrons">
<emphasis id="DBD19C1FE008FFD0FC3326D6FB69FE36" box="[976,1179,445,469]" italics="true" pageId="7" pageNumber="52">Ranatra spinifrons</emphasis>
</taxonomicName>
(see
<bibRefCitation id="8D343DFCE008FFD0FB3B26D5FA55FE36" author="Montandon AL" box="[1240,1447,445,469]" pageId="7" pageNumber="52" pagination="163 - 171" refId="ref20294" refString="Montandon AL (1910 b) Notes et descriptions d'especes nouvelles. Annales Musei Nationalis Hungarici, 8: 163 - 171." type="journal article" year="1910">Montandon, 1910b</bibRefCitation>
,
<bibRefCitation id="8D343DFCE008FFD0FCCE26B5FC97FE16" author="Montandon AL" box="[813,869,477,501]" pageId="7" pageNumber="52" pagination="118 - 125" refId="ref20317" refString="Montandon AL (1914) Nouvelles observations sur quelques formes peu connus de la fam. Nepidae, et descriptions d'especes nouvelles. Buletinul Societat ̦ ii Romane de S ̦ ciint ̦ e, 23: 118 - 125." type="journal article" year="1914">1914</bibRefCitation>
, also see
<bibRefCitation id="8D343DFCE008FFD0FC3326B6FB59FE16" author="Tran AD &amp; Poggi R" box="[976,1195,477,501]" pageId="7" pageNumber="52" pagination="236 - 246" refId="ref20899" refString="Tran AD &amp; Poggi R (2019) The genus Ranatra Fabricius (Heteroptera: Nepidae) in Borneo, with a redescription of Ranatra spinifrons Montandon and the description of a new species. Zootaxa, 4555 (2): 236 - 246." type="journal article" year="2019">Tran &amp; Poggi, 2019</bibRefCitation>
for further discussion).
</paragraph>
<paragraph id="E91A400DE008FFD0FCCE2575FB48FBB6" blockId="7.[812,1452,541,1109]" pageId="7" pageNumber="52">
The identity of
<taxonomicName id="2EA53B8EE008FFD0FC312576FBACFDD6" authorityName="Montandon" authorityYear="1905" box="[978,1118,541,565]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FC312576FBACFDD6" box="[978,1118,541,565]" italics="true" pageId="7" pageNumber="52">Ranatra stali</emphasis>
</taxonomicName>
was fixed by
<bibRefCitation id="8D343DFCE008FFD0FB112575FA55FDD6" author="Lansbury I" box="[1266,1447,541,565]" pageId="7" pageNumber="52" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansbury (1972)</bibRefCitation>
, who designated a female
<typeStatus id="361EFEAFE008FFD0FBB22556FB54FDB5" box="[1105,1190,574,598]" pageId="7" pageNumber="52" type="syntype">syntype</typeStatus>
from Mindanao as the
<typeStatus id="361EFEAFE008FFD0FCCE2535FC7DFD96" box="[813,911,605,629]" pageId="7" pageNumber="52" type="lectotype">lectotype</typeStatus>
and described it (the interpretation of
<taxonomicName id="2EA53B8EE008FFD0FAA32536FA7DFD96" authorityName="Montandon" authorityYear="1905" box="[1344,1423,605,629]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FAA32536FA7DFD96" box="[1344,1423,605,629]" italics="true" pageId="7" pageNumber="52">R. stali</emphasis>
</taxonomicName>
is based on this specimen; see description above and
<figureCitation id="719E5C88E008FFD0FAAA2515FA54FD76" box="[1353,1446,637,661]" captionStart="Fig" captionStartId="5.[113,148,2000,2022]" captionTargetBox="[113,1451,253,1984]" captionTargetId="figure-10@5.[113,1451,253,1984]" captionTargetPageId="5" captionText="Fig. 1. Habitus of Ranatra spp., all males. A, R. stali Montandon, 1905; B, R. bisaya, new species; C, R. bendanilloi, new species; D, R. brevicauda, new species; E, R. pangantihoni, new species (all of same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351680" httpUri="https://zenodo.org/record/5351680/files/figure.png" pageId="7" pageNumber="52">Figs. 1A</figureCitation>
,
<figureCitation id="719E5C88E008FFD0FCCE25F5FCCFFD56" box="[813,829,669,693]" captionStart="Fig" captionStartId="6.[136,171,1985,2007]" captionTargetBox="[141,1469,259,1965]" captionTargetId="figure-7@6.[136,1476,236,1970]" captionTargetPageId="6" captionText="Fig. 2. Ranatra stali Montandon, 1905, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, operculum; H, paramere; lateral view; I, apical part of paramere, inner view. (B, C same scale; D, E same scale; F, G same scale, H, I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351682" httpUri="https://zenodo.org/record/5351682/files/figure.png" pageId="7" pageNumber="52">2</figureCitation>
).
<bibRefCitation id="8D343DFCE008FFD0FCAD25F5FC0EFD56" author="Lansbury I" box="[846,1020,669,693]" pageId="7" pageNumber="52" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansbury (1972)</bibRefCitation>
also included and treated specimens from
<collectingRegion id="2B618EEFE008FFD0FCCF25D5FC58FD36" box="[812,938,701,725]" country="Philippines" name="Zamboanga" pageId="7" pageNumber="52">Zamboanga</collectingRegion>
(western Mindanao),
<collectingRegion id="2B618EEFE008FFD0FB7125D6FB23FD35" box="[1170,1233,702,726]" country="Philippines" name="Southern Leyte" pageId="7" pageNumber="52">Leyte</collectingRegion>
, and northern Luzon (
<collectingRegion id="2B618EEFE008FFD0FCD725B5FBF6FD16" box="[820,1028,733,757]" country="Philippines" name="Mountain Province" pageId="7" pageNumber="52">Mountain Province</collectingRegion>
) as conspecific to
<taxonomicName id="2EA53B8EE008FFD0FB2625B6FAFBFD16" authorityName="Montandon" authorityYear="1905" box="[1221,1289,733,757]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FB2625B6FAFBFD16" box="[1221,1289,733,757]" italics="true" pageId="7" pageNumber="52">R stali</emphasis>
</taxonomicName>
. Subsequently, J.
<bibRefCitation id="8D343DFCE008FFD0FCAB2595FB8AFCF6" author="Polhemus JT &amp; Reisen WK" box="[840,1144,765,789]" pageId="7" pageNumber="52" pagination="259 - 294" refId="ref20677" refString="Polhemus JT &amp; Reisen WK (1976) Aquatic Hemiptera of the Philippines. Kalikasan, Philippine Journal of Biology, 5 (3): 259 - 294." type="journal article" year="1976">Polhemus &amp; Reisen (1976)</bibRefCitation>
also reported
<taxonomicName id="2EA53B8EE008FFD0FAFE2596FA9FFCF6" authorityName="Montandon" authorityYear="1905" box="[1309,1389,765,789]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FAFE2596FA9FFCF6" box="[1309,1389,765,789]" italics="true" pageId="7" pageNumber="52">R. stali</emphasis>
</taxonomicName>
from Luzon, but based on only nymphs. After comparative study of numerous specimens from various major islands of the
<collectingCountry id="91B2009DE008FFD0FCCE2435FC5AFC96" box="[813,936,861,885]" name="Philippines" pageId="7" pageNumber="52">Philippines</collectingCountry>
, we recognise five distinct and related species. One is
<taxonomicName id="2EA53B8EE008FFD0FC9D2416FBFCFC76" authorityName="Montandon" authorityYear="1905" box="[894,1038,893,917]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FC9D2416FBFCFC76" box="[894,1038,893,917]" italics="true" pageId="7" pageNumber="52">Ranatra stali</emphasis>
</taxonomicName>
, with its distribution restricted to the biogeographic subregion of Greater Mindanao (records from Mindanao and
<collectingRegion id="2B618EEFE008FFD0FC3124D5FBE9FC36" box="[978,1051,957,981]" country="Philippines" name="Samar" pageId="7" pageNumber="52">Samar</collectingRegion>
). The other four are new to science. Based on extensive samples from major islands of the
<collectingCountry id="91B2009DE008FFD0FCCE2495FC58FBF6" box="[813,938,1021,1045]" name="Philippines" pageId="7" pageNumber="52">Philippines</collectingCountry>
, we conclude that the specimens from Luzon studied by
<bibRefCitation id="8D343DFCE008FFD0FC412375FBA7FBD6" author="Lansbury I" box="[930,1109,1053,1077]" pageId="7" pageNumber="52" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansbury (1972)</bibRefCitation>
and
<bibRefCitation id="8D343DFCE008FFD0FB6A2375FA5EFBD6" author="Polhemus JT &amp; Reisen WK" box="[1161,1452,1053,1077]" pageId="7" pageNumber="52" pagination="259 - 294" refId="ref20677" refString="Polhemus JT &amp; Reisen WK (1976) Aquatic Hemiptera of the Philippines. Kalikasan, Philippine Journal of Biology, 5 (3): 259 - 294." type="journal article" year="1976">Polhemus &amp; Reisen (1976)</bibRefCitation>
were not
<taxonomicName id="2EA53B8EE008FFD0FC772356FC13FBB6" authorityName="Montandon" authorityYear="1905" box="[916,993,1085,1109]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FC772356FC13FBB6" box="[916,993,1085,1109]" italics="true" pageId="7" pageNumber="52">R. stali</emphasis>
</taxonomicName>
, but a new species.
</paragraph>
<paragraph id="E91A400DE008FFD0FCCE2315FA62FA76" blockId="7.[812,1452,1149,1429]" pageId="7" pageNumber="52">
<bibRefCitation id="8D343DFCE008FFD0FCCE2315FC10FB76" author="Lansbury I" box="[813,994,1149,1173]" pageId="7" pageNumber="52" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansbury (1972)</bibRefCitation>
noted that
<taxonomicName id="2EA53B8EE008FFD0FB832316FB5CFB76" authorityName="Montandon" authorityYear="1905" box="[1120,1198,1149,1173]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FB832316FB5CFB76" box="[1120,1198,1149,1173]" italics="true" pageId="7" pageNumber="52">R. stali</emphasis>
</taxonomicName>
was rarely represented in collections. The main reason seems to be that in earlier times more attention was paid to collecting in stagnant water bodies than in streams. However, all species of the
<taxonomicName id="2EA53B8EE008FFD0FABD23B6FA59FB16" authorityName="Montandon" authorityYear="1905" box="[1374,1451,1245,1269]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="stali">
<emphasis id="DBD19C1FE008FFD0FABD23B6FA59FB16" box="[1374,1451,1245,1269]" italics="true" pageId="7" pageNumber="52">R. stali</emphasis>
</taxonomicName>
complex live at the edges of lentic sections of running waters. Rarely, specimens are also found in pools of intermittent streams, too. Shaded habitats are generally preferred. Often specimens sit hidden at undercuts of banks or between the roots of trees hanging into the water (Zettel, pers. obs.).
</paragraph>
<paragraph id="E91A400DE008FFD0FCCE22D6FBA6FA16" blockId="7.[813,1453,1469,1525]" pageId="7" pageNumber="52">
For further comparative notes, see Remarks under
<taxonomicName id="2EA53B8EE008FFD0FA7722D6FC49FA16" authority="Tran &amp; Zettel, 2021" authorityName="Tran &amp; Zettel" authorityYear="2021" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="pangantihoni" status="sp. nov.">
<emphasis id="DBD19C1FE008FFD0FA7722D6FC49FA16" italics="true" pageId="7" pageNumber="52">R. pangantihoni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C0E22164E008FFD0FC2922B6FBBDFA16" box="[970,1103,1501,1525]" pageId="7" pageNumber="52" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="A1BF1386E008FFD0FCCE2175FC01F996" pageId="7" pageNumber="52" type="distribution">
<paragraph id="E91A400DE008FFD0FCCE2175FC01F996" blockId="7.[813,1453,1565,1653]" pageId="7" pageNumber="52">
<emphasis id="DBD19C1FE008FFD0FCCE2175FC39F9D6" bold="true" box="[813,971,1565,1589]" pageId="7" pageNumber="52">Distribution.</emphasis>
<collectingCountry id="91B2009DE008FFD0FC342175FBADF9D6" box="[983,1119,1565,1589]" name="Philippines" pageId="7" pageNumber="52">Philippines</collectingCountry>
: Greater Mindanao Region: Mindanao and
<collectingRegion id="2B618EEFE008FFD0FC2E2155FBE7F9B6" box="[973,1045,1597,1621]" country="Philippines" name="Samar" pageId="7" pageNumber="52">Samar</collectingRegion>
.
<bibRefCitation id="8D343DFCE008FFD0FBFC2155FB17F9B6" author="Lansbury I" box="[1055,1253,1597,1621]" pageId="7" pageNumber="52" pagination="287 - 341" refId="ref19903" refString="Lansbury I (1972) A review of the Oriental species of Ranatra Fabricius (Hemiptera-Heteroptera: Nepidae). Transactions of the Royal Entomological Society of London, 124: 287 - 341." type="journal article" year="1972">Lansburys (1972)</bibRefCitation>
record from
<collectingRegion id="2B618EEFE008FFD0FA932156FA5FF9B5" box="[1392,1453,1598,1622]" country="Philippines" name="Southern Leyte" pageId="7" pageNumber="52">Leyte</collectingRegion>
needs verification.
</paragraph>
</subSubSection>
</treatment>
</document>