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Genus
<taxonomicName id="0EAF4D3EFFBC7966FF41FD256624FD0C" ID-CoL="7P563" authority="Walker, 1896" authorityName="Walker" authorityYear="1896" box="[238,538,655,682]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FF41FD256551FD0F" bold="true" box="[238,367,655,682]" italics="true" pageId="4" pageNumber="505">Halicoides</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FED9FD256624FD0C" author="Walker, A. O." box="[374,538,655,681]" pageId="4" pageNumber="505" pagination="343 - 346" refId="ref15195" refString="Walker, A. O. (1896) On two new species of Amphipoda Gammarina. Annals and Magazine of Natural History, Series 6, 17, 343 - 346, pl. 16. https: // doi. org / 10.1080 / 00222939608680377" type="journal article" year="1896">Walker, 1896</bibRefCitation>
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<taxonomicName id="0EAF4D3EFFBC7966FF38FD6465EFFD43" ID-CoL="7P563" authority="Walker, 1896: 344" authorityName="Walker" authorityPageNumber="344" authorityYear="1896" box="[151,465,718,742]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FF38FD64653BFD43" box="[151,261,718,742]" italics="true" pageId="4" pageNumber="505">Halicoides</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FEA0FD6465EFFD43" author="Walker, A. O." box="[271,465,718,742]" pageId="4" pageNumber="505" pagination="343 - 346" refId="ref15195" refString="Walker, A. O. (1896) On two new species of Amphipoda Gammarina. Annals and Magazine of Natural History, Series 6, 17, 343 - 346, pl. 16. https: // doi. org / 10.1080 / 00222939608680377" type="journal article" year="1896">Walker, 1896: 344</bibRefCitation>
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.—
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<bibRefCitation id="AD3E4B4CFFBC7966FE41FD6466FCFD43" author="Stebbing, T. R. R." box="[494,706,718,742]" pageId="4" pageNumber="505" pagination="1 - 806" refId="ref15083" refString="Stebbing, T. R. R. (1906) Amphipoda I. Gammaridea. Das Tierreich, 21, 1 - 806, figs. 1 - 127." type="journal article" year="1906">Stebbing, 1906: 221</bibRefCitation>
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.—
<treatmentCitation id="480E10ACFFBC7966FD4FFD646794FD43" author="Karaman, G. S." box="[736,938,718,742]" page="19" pageId="4" pageNumber="505" year="1974">
<bibRefCitation id="AD3E4B4CFFBC7966FD4FFD646794FD43" author="Karaman, G. S." box="[736,938,718,742]" pageId="4" pageNumber="505" pagination="1 - 46" refId="ref14245" refString="Karaman, G. S. (1974) Revision of the family Pardaliscidae with diagnosis of genera, distribution of species and bibliography. 43. Contribution to the knowledge of the Amphipoda. Acta Adriatica, 15, 1 - 46." type="journal article" year="1974">Karaman, 1974: 19</bibRefCitation>
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.—
<treatmentCitation id="480E10ACFFBC7966FC66FD6460A1FD43" author="Thurston, M. H." box="[969,1183,718,742]" page="148" pageId="4" pageNumber="505" year="1976">
<bibRefCitation id="AD3E4B4CFFBC7966FC66FD6460A1FD43" author="Thurston, M. H." box="[969,1183,718,742]" pageId="4" pageNumber="505" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston, 1976: 148</bibRefCitation>
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.—
<treatmentCitation id="480E10ACFFBC7966FB12FD64650EFCA0" author="Barnard, J. L. &amp; Karaman, G. S." page="577" pageId="4" pageNumber="505" year="1991">
<bibRefCitation id="AD3E4B4CFFBC7966FB12FD64650EFCA0" author="Barnard, J. L. &amp; Karaman, G. S." pageId="4" pageNumber="505" pagination="1 - 866" refId="ref13224" refString="Barnard, J. L. &amp; Karaman, G. S. (1991) The families and genera of marine gammaridean amphipods (except marine Gammaroidea). Parts 1 and 2. Records of the Australian Museum, Supplement 13, 1 - 866. https: // doi. org / 10.3853 / j. 0812 - 7387.13.1991.367" type="journal article" year="1991">Barnard &amp; Karaman, 1991: 577</bibRefCitation>
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.
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<taxonomicName id="0EAF4D3EFFBC7966FF38FCA66639FC81" ID-CoL="BFG9F" authority="J. L. Bamard, 1961: 78" authorityName="J. L. Bamard" authorityPageNumber="78" authorityYear="1961" box="[151,519,779,804]" class="Malacostraca" family="Pardaliscidae" genus="Pardisynopia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FF38FCA66521FC81" box="[151,287,780,804]" italics="true" pageId="4" pageNumber="505">Pardisynopia</emphasis>
J.L. Bamard, 1961: 78
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(
<taxonomicName id="0EAF4D3EFFBC7966FDB9FCA667F4FC86" ID-CoL="4DJHX" authority="J. L. Barnard, 1961" authorityName="J. L. Barnard" authorityYear="1961" box="[534,970,779,804]" class="Malacostraca" family="Pardaliscidae" genus="Pardisynopia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="tambiella">
<emphasis id="FBDBEAAFFFBC7966FDB9FCA6673AFC81" box="[534,772,780,804]" italics="true" pageId="4" pageNumber="505">Pardisynopia tambiella</emphasis>
J.L.
<bibRefCitation id="AD3E4B4CFFBC7966FC98FCA667F4FC86" author="Barnard, J. L." box="[823,970,779,804]" pageId="4" pageNumber="505" pagination="23 - 128" refId="ref13124" refString="Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 - 6000 meters. Galathea Report, 5, 23 - 128." type="journal article" year="1961">Barnard, 1961</bibRefCitation>
</taxonomicName>
, original designation).
</paragraph>
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<emphasis id="FBDBEAAFFFBC7966FF38FCF8650CFCCE" bold="true" box="[151,306,849,876]" pageId="4" pageNumber="505">
<typeStatus id="1614881FFFBC7966FF38FCF864ECFCC9" box="[151,210,850,876]" pageId="4" pageNumber="505">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="0EAF4D3EFFBC7966FE96FCF866FCFCC9" ID-CoL="3JCV5" authority="Walker, 1896" authorityName="Walker" authorityYear="1896" box="[313,706,850,876]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBC7966FE96FCF86619FCC9" box="[313,551,850,876]" italics="true" pageId="4" pageNumber="505">Halicoides anomalus</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FD82FCF866FCFCC9" author="Walker, A. O." box="[557,706,850,876]" pageId="4" pageNumber="505" pagination="343 - 346" refId="ref15195" refString="Walker, A. O. (1896) On two new species of Amphipoda Gammarina. Annals and Magazine of Natural History, Series 6, 17, 343 - 346, pl. 16. https: // doi. org / 10.1080 / 00222939608680377" type="journal article" year="1896">Walker, 1896</bibRefCitation>
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<subSubSection id="81B56536FFBC7966FF68FCDF656DFA09" pageId="4" pageNumber="505" type="diagnosis">
<paragraph id="C91036BDFFBC7966FF68FCDF6149FB5D" blockId="4.[151,1437,849,2028]" pageId="4" pageNumber="505">
<emphasis id="FBDBEAAFFFBC7966FF68FCDF657CFC2A" bold="true" box="[199,322,885,911]" pageId="4" pageNumber="505">Diagnosis.</emphasis>
Rostrum well developed, reaching half-length of article 1 of peduncle of antenna 1. Eyes absent. Antennae 12 with long flagellum. Antenna 1, peduncle short, article 1 a little longer than articles 2 and 3;
<emphasis id="FBDBEAAFFFBC7966FF38FC1465C5FC5E" bold="true" pageId="4" pageNumber="505">accessory flagellum present. In males, article 1 of primary flagellum with callynophore; article 1 of primary and accessory flagellum fused</emphasis>
. Mouthparts forming quadrate bundle. Mandibles, incisors weakly toothed on both sides, palp fully developed,
<emphasis id="FBDBEAAFFFBC7966FE75FBAF602FFB85" bold="true" box="[474,1041,1029,1056]" pageId="4" pageNumber="505">article 3 size variable, but shorter the article 2</emphasis>
. Lower lip, inner lobes coalesced. Maxilla l, palp not expanded apically. Maxilla 2 well developed, slender plates, about 3x longer than wide, inner margin of inner plate setose. Coxae 17 short, wider than long. Gnathopods 12 simple, slender, carpus not lobate, dactylus claw-shaped, without inner teeth.
<emphasis id="FBDBEAAFFFBC7966FD2AFBD861ABFB29" bold="true" box="[645,1429,1138,1164]" pageId="4" pageNumber="505">Pereopods 34, merus and carpus moderately to strongly robust</emphasis>
. Pereopods 57 elongated, propodus at least 10x longer than wide.
<emphasis id="FBDBEAAFFFBC7966FCD0FB3F61ABFB15" bold="true" box="[895,1429,1173,1200]" pageId="4" pageNumber="505">Urosomites 12 with teeth vestigial or absent</emphasis>
;
<emphasis id="FBDBEAAFFFBC7966FF38FB1366DBFB71" bold="true" box="[151,741,1209,1236]" pageId="4" pageNumber="505">urosomite 2 with long seta on posterodorsal angle</emphasis>
. Uropods 12 well developed, setose, ramus apically rounded with robust seta; uropod 3 lanceolate, setose. Telson at least 2x longer than wide, deeply cleft (more than 70%).
</paragraph>
<paragraph id="C91036BDFFBC7966FF68FAA8656DFA09" blockId="4.[151,1437,849,2028]" pageId="4" pageNumber="505">
<emphasis id="FBDBEAAFFFBC7966FF68FAA86587FABE" bold="true" box="[199,441,1281,1308]" pageId="4" pageNumber="505">Included 14 species.</emphasis>
<taxonomicName id="0EAF4D3EFFBC7966FE6DFAA96763FAB8" authority="(K. H. Barnard, 1925)" baseAuthorityName="K. H. Barnard" baseAuthorityYear="1925" box="[450,861,1282,1309]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anacantha">
<emphasis id="FBDBEAAFFFBC7966FE6DFAA96662FAB9" box="[450,604,1282,1308]" italics="true" pageId="4" pageNumber="505">H. anacantha</emphasis>
(K.H.
<bibRefCitation id="AD3E4B4CFFBC7966FD01FAA8676BFAB8" author="Barnard, K. H." box="[686,853,1282,1309]" pageId="4" pageNumber="505" pagination="319 - 380" refId="ref13286" refString="Barnard, K. H. (1925) Contributions to the Crustacean Fauna of South Africa n ° 8. Further Additions to the List of Amphipoda. Annals of the South African Museum, 20 (5), 319 - 380." type="journal article" year="1925">Barnard, 1925</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FCC4FAA960A1FAB9" authority="Walker, 1896" authorityName="Walker" authorityYear="1896" box="[875,1183,1282,1308]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBC7966FCC4FAA967C2FAB9" box="[875,1020,1282,1308]" italics="true" pageId="4" pageNumber="505">H. anomalus</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FBAAFAA860A1FAB9" author="Walker, A. O." box="[1029,1183,1282,1308]" pageId="4" pageNumber="505" pagination="343 - 346" refId="ref15195" refString="Walker, A. O. (1896) On two new species of Amphipoda Gammarina. Annals and Magazine of Natural History, Series 6, 17, 343 - 346, pl. 16. https: // doi. org / 10.1080 / 00222939608680377" type="journal article" year="1896">Walker, 1896</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FB02FAA9657FFAE5" authority="Johansen &amp; Vader, 2018" authorityName="Johansen &amp; Vader" authorityYear="2018" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FBDBEAAFFFBC7966FB02FAA96113FAB9" box="[1197,1325,1282,1308]" italics="true" pageId="4" pageNumber="505">H. borealis</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FA99FAA8657FFAE5" author="Johansen, P. O. &amp; Vader, W." pageId="4" pageNumber="505" pagination="1133 - 1145" refId="ref14186" refString="Johansen, P. O. &amp; Vader, W. (2018) Halicoides borealis, a new species of Pardaliscidae (Crustacea: Amphipoda) from the northern North Sea. Journal of Natural History, 52 (17 - 18), 1133 - 1145. https: // doi. org / 10.1080 / 00222933.2018.1453097" type="journal article" year="2018">Johansen &amp; Vader, 2018</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FEE0FA8D65D6FAE5" authorityName="Do Nascimento &amp; Serejo" authorityYear="2024" box="[335,488,1319,1344]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="campensis" status="sp. nov.">
<emphasis id="FBDBEAAFFFBC7966FEE0FA8D65D6FAE5" box="[335,488,1319,1344]" italics="true" pageId="4" pageNumber="505">H. campensis</emphasis>
</taxonomicName>
<emphasis id="FBDBEAAFFFBC7966FE5EFA8C6673FAE5" bold="true" box="[497,589,1318,1344]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="E0E857D4FFBC7966FE5EFA8C6673FAE5" box="[497,589,1318,1344]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="0EAF4D3EFFBC7966FDF1FA8D6791FAE5" authority="Thurston, 1976" authorityName="Thurston" authorityYear="1976" box="[606,943,1318,1344]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="discoveryi">
<emphasis id="FBDBEAAFFFBC7966FDF1FA8D66CBFAE5" box="[606,757,1318,1344]" italics="true" pageId="4" pageNumber="505">H. discoveryi</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FD51FA8C6791FAE5" author="Thurston, M. H." box="[766,943,1318,1344]" pageId="4" pageNumber="505" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston, 1976</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FC12FA8D6005FAE5" authorityName="Do Nascimento &amp; Serejo" authorityYear="2024" box="[957,1083,1319,1344]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="iemanja" status="sp. nov.">
<emphasis id="FBDBEAAFFFBC7966FC12FA8D6005FAE5" box="[957,1083,1319,1344]" italics="true" pageId="4" pageNumber="505">H. iemanja</emphasis>
</taxonomicName>
<emphasis id="FBDBEAAFFFBC7966FBEBFA8C609EFAE5" bold="true" box="[1092,1184,1318,1344]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="E0E857D4FFBC7966FBEBFA8C609EFAE5" box="[1092,1184,1318,1344]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="0EAF4D3EFFBC7966FB1EFA8D65B2FAC1" authority="Birstein &amp; M. Vinogradov, 1964" authorityName="Birstein &amp; M. Vinogradov" authorityYear="1964" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FBDBEAAFFFBC7966FB1EFA8D6125FAE5" box="[1201,1307,1318,1344]" italics="true" pageId="4" pageNumber="505">H. indica</emphasis>
Birstein &amp; M. Vinogradov, 1964
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FE36FAE16695FAC1" authority="Ren, 2012" authorityName="Ren" authorityYear="2012" box="[409,683,1354,1380]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="latilobata">
<emphasis id="FBDBEAAFFFBC7966FE36FAE16614FAC1" box="[409,554,1354,1380]" italics="true" pageId="4" pageNumber="505">H. latilobata</emphasis>
<bibRefCitation id="AD3E4B4CFFBC7966FD9CFAE06695FAC1" author="Ren, X." box="[563,683,1354,1380]" pageId="4" pageNumber="505" refId="ref14763" refString="Ren, X. (2012) Crustacea, Amphipoda, Gammaridea (II). Fauna Sinica, Invertebrata. vol. 43. Science Press, Beijing, 651 pp." type="book" year="2012">Ren, 2012</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FD17FAE167C4FAC1" authority="(J. L. Barnard, 1971)" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1971" box="[696,1018,1354,1380]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="lolo">
<emphasis id="FBDBEAAFFFBC7966FD17FAE16736FAC1" box="[696,776,1354,1380]" italics="true" pageId="4" pageNumber="505">H. lolo</emphasis>
(J.L.
<bibRefCitation id="AD3E4B4CFFBC7966FCE3FAE067CFFAC1" author="Barnard, J. L." box="[844,1009,1354,1380]" pageId="4" pageNumber="505" pagination="1 - 86" refId="ref13183" refString="Barnard, J. L. (1971) Gammaridean Amphipoda from a deep-sea transect off Oregon. Smithsonian Contributions to Zoology, 61, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.61" type="journal article" year="1971">Barnard, 1971</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FBA8FAE164ECFA2D" authority="Birstein &amp; M. Vinogradov, 1960" authorityName="Birstein &amp; M. Vinogradov" authorityYear="1960" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="FBDBEAAFFFBC7966FBA8FAE1605DFAC1" box="[1031,1123,1355,1380]" italics="true" pageId="4" pageNumber="505">H. nana</emphasis>
Birstein &amp; M. Vinogradov, 1960
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FF72FAC56670FA2D" authority="(J. L. Barnard, 1962)" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1962" box="[221,590,1390,1416]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="synopiae">
<emphasis id="FBDBEAAFFFBC7966FF72FAC5655DFA2D" box="[221,355,1391,1416]" italics="true" pageId="4" pageNumber="505">H. synopiae</emphasis>
(J.L.
<bibRefCitation id="AD3E4B4CFFBC7966FE0CFAC46678FA2D" author="Barnard, J. L." box="[419,582,1390,1416]" pageId="4" pageNumber="505" pagination="1 - 78" refId="ref13152" refString="Barnard, J. L. (1962) South Atlantic abyssal amphipods collected by R. V. Vema. Vema Research Series, 1, 1 - 78." type="journal article" year="1962">Barnard, 1962</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FDF5FAC567ECFA2D" authority="(J. L. Barnard, 1961)" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1961" box="[602,978,1390,1416]" class="Malacostraca" family="Pardaliscidae" genus="Pardisynopia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="tambiella">
<emphasis id="FBDBEAAFFFBC7966FDF5FAC566DBFA2D" box="[602,741,1390,1416]" italics="true" pageId="4" pageNumber="505">H. tambiella</emphasis>
(J.L.
<bibRefCitation id="AD3E4B4CFFBC7966FC8AFAC467F7FA2D" author="Barnard, J. L." box="[805,969,1390,1416]" pageId="4" pageNumber="505" pagination="23 - 128" refId="ref13124" refString="Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 - 6000 meters. Galathea Report, 5, 23 - 128." type="journal article" year="1961">Barnard, 1961</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FC73FAC56122FA2D" authority="(Stephensen, 1931)" baseAuthorityName="Stephensen" baseAuthorityYear="1931" box="[988,1308,1390,1416]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="FBDBEAAFFFBC7966FC73FAC56005FA2D" box="[988,1083,1391,1416]" italics="true" pageId="4" pageNumber="505">H. tertia</emphasis>
(
<bibRefCitation id="AD3E4B4CFFBC7966FBE7FAC4612DFA2D" author="Stephensen, K." box="[1096,1299,1390,1416]" pageId="4" pageNumber="505" pagination="179 - 290" refId="ref15114" refString="Stephensen, K. (1931) Crustacea Malacostraca. VII. (Amphipoda III). Danish Ingolf Expedition, 3 (11), 179 - 290." type="journal article" year="1931">Stephensen, 1931</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="0EAF4D3EFFBC7966FA88FAC56571FA09" authority="(Ledoyer, 1973)" baseAuthorityName="Ledoyer" baseAuthorityYear="1973" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="FBDBEAAFFFBC7966FA88FAC561A2FA2D" box="[1319,1436,1390,1416]" italics="true" pageId="4" pageNumber="505">H. walkeri</emphasis>
(
<bibRefCitation id="AD3E4B4CFFBC7966FF30FA386579FA09" author="Ledoyer, M." box="[159,327,1426,1452]" pageId="4" pageNumber="505" pagination="881 - 898" refId="ref14362" refString="Ledoyer, M. (1973) Amphipodes gammariens nouveaux ou peu connus de la region de Marseille. Tethys, 4 (4), 881 - 898." type="journal article" year="1973">Ledoyer, 1973</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="81B56536FFBC7967FF68FA1C6532FE74" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="discussion">
<paragraph id="C91036BDFFBC7966FF68FA1C6596F955" blockId="4.[151,1437,849,2028]" pageId="4" pageNumber="505">
<emphasis id="FBDBEAAFFFBC7966FF68FA1C6503FA75" bold="true" box="[199,317,1462,1488]" pageId="4" pageNumber="505">Remarks:</emphasis>
The scale-like structure in the second article of the peduncle of antenna 1, and the absence of accessory flagellum in the
<taxonomicName id="0EAF4D3EFFBC7966FE66FA716664FA51" authorityName="Walker" authorityYear="1896" box="[457,602,1498,1524]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBC7966FE66FA716664FA51" box="[457,602,1498,1524]" italics="true" pageId="4" pageNumber="505">H. anomalus</emphasis>
</taxonomicName>
<typeStatus id="1614881FFFBC7966FDCDFA7066ACFA51" box="[610,658,1498,1524]" pageId="4" pageNumber="505">type</typeStatus>
material, caused confusions and conflict in the status of
<taxonomicName id="0EAF4D3EFFBC7966FAB3FA7061ABFA51" authorityName="Walker" authorityYear="1896" box="[1308,1429,1498,1524]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FAB3FA7061ABFA51" box="[1308,1429,1498,1524]" italics="true" pageId="4" pageNumber="505">Halicoides</emphasis>
</taxonomicName>
. The structure was understood by
<bibRefCitation id="AD3E4B4CFFBC7966FDA2FA546691F9BD" author="Walker, A. O." box="[525,687,1534,1560]" pageId="4" pageNumber="505" pagination="343 - 346" refId="ref15195" refString="Walker, A. O. (1896) On two new species of Amphipoda Gammarina. Annals and Magazine of Natural History, Series 6, 17, 343 - 346, pl. 16. https: // doi. org / 10.1080 / 00222939608680377" type="journal article" year="1896">Walker (1896)</bibRefCitation>
as a scale-like accessory flagellum dislocated to the second article of the peduncle, rather than on the third, which would be a unique character among all
<taxonomicName id="0EAF4D3EFFBC7966FBD8F98860C1F999" authorityName="Latreille" authorityYear="1816" box="[1143,1279,1570,1596]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="order">Amphipoda</taxonomicName>
. The question remained until
<taxonomicName id="0EAF4D3EFFBC7966FEEEF9EC6623F9C5" authorityName="Johansen &amp; Vader" authorityYear="2018" box="[321,541,1606,1632]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FBDBEAAFFFBC7966FEEEF9EC6623F9C5" box="[321,541,1606,1632]" italics="true" pageId="4" pageNumber="505">Halicoides borealis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="AD3E4B4CFFBC7966FD43F9EC6033F9C5" author="Johansen, P. O. &amp; Vader, W." box="[748,1037,1606,1632]" pageId="4" pageNumber="505" pagination="1133 - 1145" refId="ref14186" refString="Johansen, P. O. &amp; Vader, W. (2018) Halicoides borealis, a new species of Pardaliscidae (Crustacea: Amphipoda) from the northern North Sea. Journal of Natural History, 52 (17 - 18), 1133 - 1145. https: // doi. org / 10.1080 / 00222933.2018.1453097" type="journal article" year="2018">Johansen &amp; Vader (2018)</bibRefCitation>
. In this species, an oval structure at the end of article 2 was found in male antenna 1, in addition to a well-developed accessory flagellum.
<bibRefCitation id="AD3E4B4CFFBC7966FAB7F9C06515F90D" author="Johansen, P. O. &amp; Vader, W." pageId="4" pageNumber="505" pagination="1133 - 1145" refId="ref14186" refString="Johansen, P. O. &amp; Vader, W. (2018) Halicoides borealis, a new species of Pardaliscidae (Crustacea: Amphipoda) from the northern North Sea. Journal of Natural History, 52 (17 - 18), 1133 - 1145. https: // doi. org / 10.1080 / 00222933.2018.1453097" type="journal article" year="2018">Johansen &amp; Vader (2018)</bibRefCitation>
had access to the unpublished holotype-drawings of
<taxonomicName id="0EAF4D3EFFBC7966FC29F9256029F90D" authorityName="Walker" authorityYear="1896" box="[902,1047,1678,1704]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBC7966FC29F9256029F90D" box="[902,1047,1678,1704]" italics="true" pageId="4" pageNumber="505">H. anomalus</emphasis>
</taxonomicName>
, re-examined by
<bibRefCitation id="AD3E4B4CFFBC7966FB72F92461A9F90D" author="Thurston, M. H." box="[1245,1431,1678,1704]" pageId="4" pageNumber="505" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston (1976)</bibRefCitation>
, and considered that the structures of
<taxonomicName id="0EAF4D3EFFBC7966FD9EF91966FEF969" authorityName="Walker" authorityYear="1896" box="[561,704,1714,1740]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBC7966FD9EF91966FEF969" box="[561,704,1714,1740]" italics="true" pageId="4" pageNumber="505">H. anomalus</emphasis>
</taxonomicName>
and
<taxonomicName id="0EAF4D3EFFBC7966FD59F919674DF969" authorityName="Johansen &amp; Vader" authorityYear="2018" box="[758,883,1714,1740]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FBDBEAAFFFBC7966FD59F919674DF969" box="[758,883,1714,1740]" italics="true" pageId="4" pageNumber="505">H. borealis</emphasis>
</taxonomicName>
are the same, which may have a sensory function to male sexual behavior.
</paragraph>
<paragraph id="C91036BDFFBC7966FF68F9506118F849" blockId="4.[151,1437,849,2028]" pageId="4" pageNumber="505">
Taking these into account, we assume that
<taxonomicName id="0EAF4D3EFFBC7966FD07F9516706F8B1" authorityName="Walker" authorityYear="1896" box="[680,824,1786,1812]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBC7966FD07F9516706F8B1" box="[680,824,1786,1812]" italics="true" pageId="4" pageNumber="505">H. anomalus</emphasis>
</taxonomicName>
possess accessory flagellum together with the sensory oval structure, but this flagellum was missing accidentaly in the
<typeStatus id="1614881FFFBC7966FCD3F8B46792F89D" box="[892,940,1822,1848]" pageId="4" pageNumber="505">type</typeStatus>
material, as suggested by
<bibRefCitation id="AD3E4B4CFFBC7966FB72F8B461A6F89D" author="Thurston, M. H." box="[1245,1432,1822,1848]" pageId="4" pageNumber="505" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston (1976)</bibRefCitation>
. Thus, since every other
<taxonomicName id="0EAF4D3EFFBC7966FE0EF8E8664FF8F9" box="[417,625,1858,1884]" pageId="4" pageNumber="505">
<emphasis id="FBDBEAAFFFBC7966FE0EF8E86624F8F9" box="[417,538,1858,1884]" italics="true" pageId="4" pageNumber="505">Halicoides</emphasis>
species
</taxonomicName>
has a well-developed accessory flagellum in antenna 1, we consider it as a diagnostic character of
<taxonomicName id="0EAF4D3EFFBC7966FE00F8CC6616F825" authorityName="Walker" authorityYear="1896" box="[431,552,1894,1920]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FE00F8CC6616F825" box="[431,552,1894,1920]" italics="true" pageId="4" pageNumber="505">Halicoides</emphasis>
</taxonomicName>
. Further on,
<taxonomicName id="0EAF4D3EFFBC7966FD17F8CC670FF825" authorityName="Walker" authorityYear="1896" box="[696,817,1894,1920]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FD17F8CC670FF825" box="[696,817,1894,1920]" italics="true" pageId="4" pageNumber="505">Halicoides</emphasis>
</taxonomicName>
can be defined by a combination of characters, initially defined by
<bibRefCitation id="AD3E4B4CFFBC7966FEB5F82065EBF801" author="Thurston, M. H." box="[282,469,1930,1956]" pageId="4" pageNumber="505" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston (1976)</bibRefCitation>
, that will be compared with the closer genus
<taxonomicName id="0EAF4D3EFFBC7966FC48F820600CF801" authorityName="Boeck" authorityYear="1871" box="[999,1074,1930,1956]" class="Malacostraca" family="Pardaliscidae" genus="Halice" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBC7966FC48F820600CF801" box="[999,1074,1930,1956]" italics="true" pageId="4" pageNumber="505">Halice</emphasis>
</taxonomicName>
(in parentheses): 1) absence or vestigial teeth on urosomites 1 and 2 (versus long teeth); pereopods 3 and 4 moderately to strongly robust (versus thin and not expanded); presence of a single seta on the posterior margin of urosomite 2 (versus absent).
</paragraph>
<paragraph id="C91036BDFFBD7967FF68FF3D6532FE74" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="506">
The presence of a seta on urosomite 2 has not been originally described for some species, but a re-examination of
<typeStatus id="1614881FFFBD7967FF19FF1164D8FF70" box="[182,230,187,213]" pageId="5" pageNumber="506">type</typeStatus>
material of
<taxonomicName id="0EAF4D3EFFBD7967FEC1FF116638FF70" baseAuthorityName="K. H. Barnard" baseAuthorityYear="1925" box="[366,518,187,213]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="anacantha">
<emphasis id="FBDBEAAFFFBD7967FEC1FF116638FF70" box="[366,518,187,213]" italics="true" pageId="5" pageNumber="506">H. anacantha</emphasis>
</taxonomicName>
,
<taxonomicName id="0EAF4D3EFFBD7967FDBAFF11669AFF70" authorityName="Walker" authorityYear="1896" box="[533,676,187,213]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="FBDBEAAFFFBD7967FDBAFF11669AFF70" box="[533,676,187,213]" italics="true" pageId="5" pageNumber="506">H. anomalus</emphasis>
</taxonomicName>
,
<taxonomicName id="0EAF4D3EFFBD7967FD1DFF116706FF71" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1962" box="[690,824,187,212]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="synopiae">
<emphasis id="FBDBEAAFFFBD7967FD1DFF116706FF71" box="[690,824,187,212]" italics="true" pageId="5" pageNumber="506">H. synopiae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EAF4D3EFFBD7967FCC1FF1167C5FF70" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1961" box="[878,1019,187,213]" class="Malacostraca" family="Pardaliscidae" genus="Pardisynopia" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="tambiella">
<emphasis id="FBDBEAAFFFBD7967FCC1FF1167C5FF70" box="[878,1019,187,213]" italics="true" pageId="5" pageNumber="506">H. tambiella</emphasis>
</taxonomicName>
by
<bibRefCitation id="AD3E4B4CFFBD7967FB8BFF1160E5FF70" author="Thurston, M. H." box="[1060,1243,187,213]" pageId="5" pageNumber="506" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston (1976)</bibRefCitation>
has shown that a seta is present on urosomite 2.
<bibRefCitation id="AD3E4B4CFFBD7967FE45FF75669EFF5C" author="Thurston, M. H." box="[490,672,223,249]" pageId="5" pageNumber="506" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston (1976)</bibRefCitation>
suggested that this seta may be presented in all species of
<taxonomicName id="0EAF4D3EFFBD7967FA8CFF7561A2FF5C" authorityName="Walker" authorityYear="1896" box="[1315,1436,223,249]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBD7967FA8CFF7561A2FF5C" box="[1315,1436,223,249]" italics="true" pageId="5" pageNumber="506">Halicoides</emphasis>
</taxonomicName>
and thus constitutes a character of generic value. The presence of this characteristic seta was also noticed by the first author in
<taxonomicName id="0EAF4D3EFFBD7967FE94FE8D65B5FEE4" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1971" box="[315,395,295,321]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="lolo">
<emphasis id="FBDBEAAFFFBD7967FE94FE8D65B5FEE4" box="[315,395,295,321]" italics="true" pageId="5" pageNumber="506">H. lolo</emphasis>
</taxonomicName>
and
<taxonomicName id="0EAF4D3EFFBD7967FE6CFE8D6675FEE5" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1962" box="[451,587,295,320]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="synopiae">
<emphasis id="FBDBEAAFFFBD7967FE6CFE8D6675FEE5" box="[451,587,295,320]" italics="true" pageId="5" pageNumber="506">H. synopiae</emphasis>
</taxonomicName>
when re-examining material of both species at the Smithsonian Institution (2019). On the other hand,
<taxonomicName id="0EAF4D3EFFBD7967FE7BFEE16601FEC0" authorityName="Birstein &amp; M. Vinogradov" authorityYear="1964" box="[468,575,331,357]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FBDBEAAFFFBD7967FE7BFEE16601FEC0" box="[468,575,331,357]" italics="true" pageId="5" pageNumber="506">H. indica</emphasis>
</taxonomicName>
,
<taxonomicName id="0EAF4D3EFFBD7967FDE0FEE166DFFEC0" authorityName="Ren" authorityYear="2012" box="[591,737,331,357]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="latilobata">
<emphasis id="FBDBEAAFFFBD7967FDE0FEE166DFFEC0" box="[591,737,331,357]" italics="true" pageId="5" pageNumber="506">H. latilobata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EAF4D3EFFBD7967FCB2FEE16744FEC1" authorityName="Birstein &amp; M. Vinogradov" authorityYear="1960" box="[797,890,331,356]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="FBDBEAAFFFBD7967FCB2FEE16744FEC1" box="[797,890,331,356]" italics="true" pageId="5" pageNumber="506">H. nana</emphasis>
</taxonomicName>
have no mention of the presence of the seta in their descriptions, and it was not possible to verify material of these species. Nevertheless, we agree with
<bibRefCitation id="AD3E4B4CFFBD7967FA98FEC564DCFE08" author="Thurston, M. H." pageId="5" pageNumber="506" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston (1976)</bibRefCitation>
that it is probable that all
<taxonomicName id="0EAF4D3EFFBD7967FDBAFE3966D7FE08" box="[533,745,403,429]" pageId="5" pageNumber="506">
<emphasis id="FBDBEAAFFFBD7967FDBAFE3966B0FE08" box="[533,654,403,429]" italics="true" pageId="5" pageNumber="506">Halicoides</emphasis>
species
</taxonomicName>
have the seta, being an important diagnostic character to be examined.
</paragraph>
</subSubSection>
<subSubSection id="81B56536FFBD7967FF68FE7065FAFC24" pageId="5" pageNumber="506" type="distribution">
<paragraph id="C91036BDFFBD7967FF68FE70662FFD54" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="506">
<emphasis id="FBDBEAAFFFBD7967FF68FE7065C8FE51" bold="true" box="[199,502,474,501]" pageId="5" pageNumber="506">Distribution and habitat.</emphasis>
Marine, cosmopolitan.
<taxonomicName id="0EAF4D3EFFBD7967FCA8FE7167A7FE50" authorityName="Ren" authorityYear="2012" box="[775,921,475,501]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="latilobata">
<emphasis id="FBDBEAAFFFBD7967FCA8FE7167A7FE50" box="[775,921,475,501]" italics="true" pageId="5" pageNumber="506">H. latilobata</emphasis>
</taxonomicName>
from East
<collectingCountry id="B1B8762DFFBD7967FBB4FE716060FE50" box="[1051,1118,475,501]" name="China" pageId="5" pageNumber="506">China</collectingCountry>
Sea (
<bibRefCitation id="AD3E4B4CFFBD7967FB0EFE716128FE50" author="Ren, X." box="[1185,1302,475,501]" pageId="5" pageNumber="506" refId="ref14763" refString="Ren, X. (2012) Crustacea, Amphipoda, Gammaridea (II). Fauna Sinica, Invertebrata. vol. 43. Science Press, Beijing, 651 pp." type="book" year="2012">Ren 2012</bibRefCitation>
).
<taxonomicName id="0EAF4D3EFFBD7967FA84FE7161ABFE50" authorityName="Birstein &amp; M. Vinogradov" authorityYear="1964" box="[1323,1429,475,501]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FBDBEAAFFFBD7967FA84FE7161ABFE50" box="[1323,1429,475,501]" italics="true" pageId="5" pageNumber="506">H. indica</emphasis>
</taxonomicName>
,
<taxonomicName id="0EAF4D3EFFBD7967FF38FE55651BFDBC" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1961" box="[151,293,511,537]" class="Malacostraca" family="Pardaliscidae" genus="Pardisynopia" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="tambiella">
<emphasis id="FBDBEAAFFFBD7967FF38FE55651BFDBC" box="[151,293,511,537]" italics="true" pageId="5" pageNumber="506">H. tambiella</emphasis>
</taxonomicName>
,
<taxonomicName id="0EAF4D3EFFBD7967FE9AFE556582FDBD" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1962" box="[309,444,511,536]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="synopiae">
<emphasis id="FBDBEAAFFFBD7967FE9AFE556582FDBD" box="[309,444,511,536]" italics="true" pageId="5" pageNumber="506">H. synopiae</emphasis>
</taxonomicName>
,
<taxonomicName id="0EAF4D3EFFBD7967FE63FE556616FDBD" authorityName="Birstein &amp; M. Vinogradov" authorityYear="1960" box="[460,552,511,536]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="FBDBEAAFFFBD7967FE63FE556616FDBD" box="[460,552,511,536]" italics="true" pageId="5" pageNumber="506">H. nana</emphasis>
</taxonomicName>
and
<taxonomicName id="0EAF4D3EFFBD7967FDCEFE55668FFDBC" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1971" box="[609,689,511,537]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="lolo">
<emphasis id="FBDBEAAFFFBD7967FDCEFE55668FFDBC" box="[609,689,511,537]" italics="true" pageId="5" pageNumber="506">H. lolo</emphasis>
</taxonomicName>
from Indian Ocean, Tasmanian Sea, Southern California, Eastern Pacific, respectively (
<bibRefCitation id="AD3E4B4CFFBD7967FE25FD8966EAFD98" author="Birstein, J. A. &amp; Vinogradov, M. E." box="[394,724,547,573]" pageId="5" pageNumber="506" pagination="152 - 195" refId="ref13381" refString="Birstein, J. A. &amp; Vinogradov, M. E. (1964). Pelagischeskie gammaridy severnoi chasti Indiiskogo Okeana [The pelagic amphipods gammarids of the northern part of the Indian Ocean]. Trudy Instituta Okeanologii, Akademia Nauk SSSR, 65, 152 - 195." type="journal article" year="1964">Birstein &amp; Vinogradov, 1964</bibRefCitation>
; J.L.
<bibRefCitation id="AD3E4B4CFFBD7967FCBDFD896791FD98" author="Barnard, J. L." box="[786,943,547,573]" pageId="5" pageNumber="506" pagination="23 - 128" refId="ref13124" refString="Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 - 6000 meters. Galathea Report, 5, 23 - 128." type="journal article" year="1961">Barnard 1961</bibRefCitation>
; J.L.
<bibRefCitation id="AD3E4B4CFFBD7967FC42FD8960B4FD98" author="Barnard, J. L." box="[1005,1162,547,573]" pageId="5" pageNumber="506" pagination="1 - 78" refId="ref13152" refString="Barnard, J. L. (1962) South Atlantic abyssal amphipods collected by R. V. Vema. Vema Research Series, 1, 1 - 78." type="journal article" year="1962">Barnard 1962</bibRefCitation>
;
<bibRefCitation id="AD3E4B4CFFBD7967FB39FD8964ECFDC4" author="Birstein, J. A. &amp; Vinogradov, M. E." pageId="5" pageNumber="506" pagination="165 - 241" refId="ref13331" refString="Birstein, J. A. &amp; Vinogradov, M. E. (1960) Pelagischeskie gammaridy tropischeskoi chasti Ticogo Okeana. [Pelagic Gammaridea of the tropical part of the Pacific Ocean]. Trudy Instituta Okeanologii, Akademia Nauk SSSR, 34, 165 - 241." type="journal article" year="1960">Birstein &amp; Vinogradov, 1960</bibRefCitation>
;
<bibRefCitation id="AD3E4B4CFFBD7967FF73FDED6541FDC4" author="Barnard, J. L." box="[220,383,583,609]" pageId="5" pageNumber="506" pagination="1 - 86" refId="ref13183" refString="Barnard, J. L. (1971) Gammaridean Amphipoda from a deep-sea transect off Oregon. Smithsonian Contributions to Zoology, 61, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.61" type="journal article" year="1971">Barnard, 1971</bibRefCitation>
).
<taxonomicName id="0EAF4D3EFFBD7967FE3EFDED6616FDC4" baseAuthorityName="K. H. Barnard" baseAuthorityYear="1925" box="[401,552,583,609]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="anacantha">
<emphasis id="FBDBEAAFFFBD7967FE3EFDED6616FDC4" box="[401,552,583,609]" italics="true" pageId="5" pageNumber="506">H. anacantha</emphasis>
</taxonomicName>
from
<collectingCountry id="B1B8762DFFBD7967FDC6FDED66C8FDC4" box="[617,758,583,609]" name="South Africa" pageId="5" pageNumber="506">South Africa</collectingCountry>
(K.H.
<bibRefCitation id="AD3E4B4CFFBD7967FCEEFDED67E7FDC4" author="Barnard, K. H." box="[833,985,582,609]" pageId="5" pageNumber="506" pagination="319 - 380" refId="ref13286" refString="Barnard, K. H. (1925) Contributions to the Crustacean Fauna of South Africa n ° 8. Further Additions to the List of Amphipoda. Annals of the South African Museum, 20 (5), 319 - 380." type="journal article" year="1925">Barnard 1925</bibRefCitation>
) and
<taxonomicName id="0EAF4D3EFFBD7967FBBAFDED60B4FDC4" baseAuthorityName="Ledoyer" baseAuthorityYear="1973" box="[1045,1162,583,609]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="FBDBEAAFFFBD7967FBBAFDED60B4FDC4" box="[1045,1162,583,609]" italics="true" pageId="5" pageNumber="506">H. walkeri</emphasis>
</taxonomicName>
from Mediterranean Sea (
<bibRefCitation id="AD3E4B4CFFBD7967FF30FDC16579FD20" author="Ledoyer, M." box="[159,327,619,645]" pageId="5" pageNumber="506" pagination="881 - 898" refId="ref14362" refString="Ledoyer, M. (1973) Amphipodes gammariens nouveaux ou peu connus de la region de Marseille. Tethys, 4 (4), 881 - 898." type="journal article" year="1973">Ledoyer, 1973</bibRefCitation>
). North Atlantic Ocean,
<taxonomicName id="0EAF4D3EFFBD7967FDF5FDC166D1FD20" authorityName="Thurston" authorityYear="1976" box="[602,751,619,645]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="discoveryi">
<emphasis id="FBDBEAAFFFBD7967FDF5FDC166D1FD20" box="[602,751,619,645]" italics="true" pageId="5" pageNumber="506">H. discoveryi</emphasis>
</taxonomicName>
from the Canary Islands (
<bibRefCitation id="AD3E4B4CFFBD7967FBBAFDC160F8FD20" author="Thurston, M. H." box="[1045,1222,619,645]" pageId="5" pageNumber="506" pagination="143 - 159" refId="ref15144" refString="Thurston, M. H. (1976) New pelagic amphipods (Crustacea: Amphipoda) collected on the Sond Cruise. Journal of the Marine Biological Association of the United Kingdom, 56, 143 - 159. https: // doi. org / 10.1017 / S 0025315400020506" type="journal article" year="1976">Thurston, 1976</bibRefCitation>
),
<taxonomicName id="0EAF4D3EFFBD7967FB75FDC16160FD20" authorityName="Walker" authorityYear="1896" box="[1242,1374,619,645]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="FBDBEAAFFFBD7967FB75FDC16160FD20" box="[1242,1374,619,645]" italics="true" pageId="5" pageNumber="506">H. anomala</emphasis>
</taxonomicName>
from Bay of Biscay, Isle de Yeu (
<bibRefCitation id="AD3E4B4CFFBD7967FE61FD256657FD0C" author="Walker, A. O." box="[462,617,655,681]" pageId="5" pageNumber="506" pagination="343 - 346" refId="ref15195" refString="Walker, A. O. (1896) On two new species of Amphipoda Gammarina. Annals and Magazine of Natural History, Series 6, 17, 343 - 346, pl. 16. https: // doi. org / 10.1080 / 00222939608680377" type="journal article" year="1896">Walker, 1896</bibRefCitation>
),
<taxonomicName id="0EAF4D3EFFBD7967FDD2FD2566E3FD0D" baseAuthorityName="Stephensen" baseAuthorityYear="1931" box="[637,733,655,680]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="FBDBEAAFFFBD7967FDD2FD2566E3FD0D" box="[637,733,655,680]" italics="true" pageId="5" pageNumber="506">H. tertia</emphasis>
</taxonomicName>
from West of
<collectingCountry id="B1B8762DFFBD7967FCD1FD2567CDFD0C" box="[894,1011,655,681]" name="Greenland" pageId="5" pageNumber="506">Greenland</collectingCountry>
(
<bibRefCitation id="AD3E4B4CFFBD7967FBADFD2560F3FD0C" author="Stephensen, K." box="[1026,1229,655,681]" pageId="5" pageNumber="506" pagination="179 - 290" refId="ref15114" refString="Stephensen, K. (1931) Crustacea Malacostraca. VII. (Amphipoda III). Danish Ingolf Expedition, 3 (11), 179 - 290." type="journal article" year="1931">Stephensen, 1931</bibRefCitation>
).
<taxonomicName id="0EAF4D3EFFBD7967FB4EFD256161FD0C" authorityName="Johansen &amp; Vader" authorityYear="2018" box="[1249,1375,655,681]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FBDBEAAFFFBD7967FB4EFD256161FD0C" box="[1249,1375,655,681]" italics="true" pageId="5" pageNumber="506">H. borealis</emphasis>
</taxonomicName>
from Northern North Sea (
<bibRefCitation id="AD3E4B4CFFBD7967FE28FD19669CFD68" author="Johansen, P. O. &amp; Vader, W." box="[391,674,691,717]" pageId="5" pageNumber="506" pagination="1133 - 1145" refId="ref14186" refString="Johansen, P. O. &amp; Vader, W. (2018) Halicoides borealis, a new species of Pardaliscidae (Crustacea: Amphipoda) from the northern North Sea. Journal of Natural History, 52 (17 - 18), 1133 - 1145. https: // doi. org / 10.1080 / 00222933.2018.1453097" type="journal article" year="2018">Johansen &amp; Vader, 2018</bibRefCitation>
),
<taxonomicName id="0EAF4D3EFFBD7967FD19FD196771FD69" authorityName="Do Nascimento &amp; Serejo" authorityYear="2024" box="[694,847,691,716]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="campensis" status="sp. nov">
<emphasis id="FBDBEAAFFFBD7967FD19FD196771FD69" box="[694,847,691,716]" italics="true" pageId="5" pageNumber="506">H. campensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0E857D4FFBD7967FCF8FD196794FD68" box="[855,938,691,717]" pageId="5" pageNumber="506" rank="species">sp. nov</taxonomicNameLabel>
and
<taxonomicName id="0EAF4D3EFFBD7967FC4CFD19605FFD69" authorityName="Do Nascimento &amp; Serejo" authorityYear="2024" box="[995,1121,691,716]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="iemanja" status="sp. nov">
<emphasis id="FBDBEAAFFFBD7967FC4CFD19605FFD69" box="[995,1121,691,716]" italics="true" pageId="5" pageNumber="506">H. iemanja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0E857D4FFBD7967FBC5FD196082FD68" box="[1130,1212,691,717]" pageId="5" pageNumber="506" rank="species">sp. nov</taxonomicNameLabel>
from Southwestern Atlantic (present study,
<figureCitation id="51942A38FFBD7967FE0DFD7D663EFD54" box="[418,512,727,753]" captionStart="FIGURE 1" captionStartId="5.[151,250,1832,1856]" captionTargetBox="[158,1429,928,1801]" captionTargetId="figure-362@5.[151,1436,921,1808]" captionTargetPageId="5" captionText="FIGURE 1. Distribution map of Halicoides campensis sp. nov (orange dots) and H. iemanja sp. nov. (purple dots) along the study area (19o26o S). White stars indicate the type localities of each species as shown in the map. Samples were collected between 293084 m depths, from Espírito Santo, Campos and Santos Basins." figureDoi="http://doi.org/10.5281/zenodo.12782971" httpUri="https://zenodo.org/record/12782971/files/figure.png" pageId="5" pageNumber="506">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="C91036BDFFBD7967FF68FD5165FAFC24" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="506">
The genus seems to be more dominant in shallow waters (
<bibRefCitation id="AD3E4B4CFFBD7967FCDFFD516097FCB0" author="Barnard, J. L. &amp; Karaman, G. S." box="[880,1193,763,789]" pageId="5" pageNumber="506" pagination="1 - 866" refId="ref13224" refString="Barnard, J. L. &amp; Karaman, G. S. (1991) The families and genera of marine gammaridean amphipods (except marine Gammaroidea). Parts 1 and 2. Records of the Australian Museum, Supplement 13, 1 - 866. https: // doi. org / 10.3853 / j. 0812 - 7387.13.1991.367" type="journal article" year="1991">Barnard &amp; Karaman, 1991</bibRefCitation>
), but showed a wide bathymetric range
<quantity id="0E579B58FFBD7967FEDEFCB565CBFC9C" box="[369,501,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.5565000000000002" metricValueMax="3.0839999999999996" metricValueMin="0.028999999999999998" pageId="5" pageNumber="506" unit="m" value="1556.5" valueMax="3084.0" valueMin="29.0">293084 m</quantity>
, based on data of the present study. The present work brings the first record of
<taxonomicName id="0EAF4D3EFFBD7967FF38FCE9652EFCF8" authorityName="Walker" authorityYear="1896" box="[151,272,835,861]" class="Malacostraca" family="Pardaliscidae" genus="Halicoides" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="FBDBEAAFFFBD7967FF38FCE9652EFCF8" box="[151,272,835,861]" italics="true" pageId="5" pageNumber="506">Halicoides</emphasis>
</taxonomicName>
for the Southwest Atlantic Ocean in addition to expand the bathymetry of the genus, which was previously known from
<quantity id="0E579B58FFBD7967FE87FCCD65FEFC24" box="[296,448,871,897]" metricMagnitude="2" metricUnit="m" metricValue="8.754999999999999" metricValueMax="17.2" metricValueMin="0.31" pageId="5" pageNumber="506" unit="m" value="875.5" valueMax="1720.0" valueMin="31.0">31 to 1720 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>