571 lines
71 KiB
XML
571 lines
71 KiB
XML
<document id="7076C7A10EA2B155514D1B691950BD53" ID-CLB-Dataset="8775" ID-DOI="10.11646/zootaxa.4822.4.2" ID-GBIF-Dataset="847ff980-f84b-46a1-9e94-37d1cd058637" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401787" ID-ZooBank="D1AF5AD7-E23A-4121-B6E6-4F31B430D9B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609339126870" checkinUser="plazi" docAuthor="Manuel, Michaël & Ramahandrison, Andriamirado T." docDate="2020" docId="3F78285625078976FF23FD4B88E7F894" docLanguage="en" docName="zootaxa.4822.4.2.pdf" docOrigin="Zootaxa 4822 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccophilus leguyaderi Manuel & Ramahandrison 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="485" masterDocId="C341502E25058975FFB4FFC58C54FFE5" masterDocTitle="Four new species of the diving beetle genus Laccophilus Leach, 1815 from Madagascar (Coleoptera, Dytiscidae, Laccophilini)" masterLastPageNumber="502" masterPageNumber="482" pageNumber="484" updateTime="1698849761501" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="7BB4FAE430D10F0F0A05774BA8DFB663">
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<mods:title id="8518209D213640E831162A3E6C808D49">Four new species of the diving beetle genus Laccophilus Leach, 1815 from Madagascar (Coleoptera, Dytiscidae, Laccophilini)</mods:title>
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</mods:titleInfo>
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<mods:name id="3B9F4C9E037D4368FD59B753DDB9883D" type="personal">
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<mods:roleTerm id="EC30BA0F3B089AB89F694DA32C8314D3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="571025E299D4EFD000AE10D563B51C75">Manuel, Michaël</mods:namePart>
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<mods:affiliation id="AB78FD7D0F3278856ED2DF81692655CB">Sorbonne Université, Institut de Systématique, Evolution, Biodiversité (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, 75005 Paris, France. & michael. manuel @ upmc. fr; http: // orcid. org / 0000 - 0001 - 9728 - 5580</mods:affiliation>
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<mods:nameIdentifier id="3DC635EBA66CB151B53EAF0076E96199" type="email">michael.manuel@upmc.fr</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="480BB4A41DC1DF5997859545DC91743B">Ramahandrison, Andriamirado T.</mods:namePart>
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<mods:affiliation id="0DEE1449922CEF6E9026CAA3ACB80E35">Sorbonne Université, Institut de Systématique, Evolution, Biodiversité (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, 75005 Paris, France. & Département de Biologie et Ecologie Végétales, Faculté des Sciences, BP 906, Université d’Antananarivo, Antananarivo, Madagascar. mirad. 95 @ gmail. com; http: // orcid. org / 0000 - 0002 - 0833 - 8730</mods:affiliation>
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<mods:nameIdentifier id="FE2DDBD77CB01C965C8203025D089B08" type="email">mirad.95@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="F02ADE4CD1ED5AC399057DF59E0572B5">text</mods:typeOfResource>
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<mods:title id="D4764A07D68F8396AD07AADF8D3B010A">Zootaxa</mods:title>
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<mods:part id="864CC268565E86B1C86756062BB10BD1">
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<mods:date id="515B30F9AA96F3F3806DCEE516DCC98A">2020</mods:date>
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<mods:number id="61E0E0660C8A5FC592955C224D97DF5E">2020-08-07</mods:number>
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<mods:number id="7C308EBA18B9584DEE64A8B30776B13E">4822</mods:number>
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<mods:detail id="14562BECAFA18A1EA52D93E7234077C1" type="issue">
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<mods:end id="B819B21EB6368159086281C0312F9F17">502</mods:end>
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<mods:classification id="2415F185CF8CD6C90B4CBA2E826D8649">journal article</mods:classification>
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<mods:identifier id="492BED0A92A8DF910C3B20346DEE4488" type="DOI">10.11646/zootaxa.4822.4.2</mods:identifier>
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<treatment id="3F78285625078976FF23FD4B88E7F894" ID-DOI="http://doi.org/10.5281/zenodo.4401785" ID-GBIF-Taxon="177010028" ID-Zenodo-Dep="4401785" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894" httpUri="http://treatment.plazi.org/id/3F78285625078976FF23FD4B88E7F894" lastPageId="3" lastPageNumber="485" pageId="2" pageNumber="484">
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<subSubSection id="FFCBCACB25078977FF23FD4B8E43FD4C" box="[151,535,654,681]" pageId="2" pageNumber="484" type="nomenclature">
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<paragraph id="B76E994025078977FF23FD4B8E43FD4C" blockId="2.[151,535,654,717]" box="[151,535,654,681]" pageId="2" pageNumber="484">
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<heading id="EC262E2C25078977FF23FD4B8E43FD4C" bold="true" box="[151,535,654,681]" fontSize="11" level="1" pageId="2" pageNumber="484" reason="1">
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<emphasis id="85A5455225078977FF23FD4B8E43FD4C" bold="true" box="[151,535,654,681]" pageId="2" pageNumber="484">
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<taxonomicName id="70D1E2C325078977FF23FD4B8DE6FD4C" ID-CoL="8LZXJ" authority="Manuel & Ramahandrison, 2020" authorityName="Manuel & Ramahandrison" authorityYear="2020" box="[151,434,654,681]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="484" phylum="Arthropoda" rank="species" species="leguyaderi" status="sp. nov.">
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<emphasis id="85A5455225078977FF23FD4B8DE6FD4C" bold="true" box="[151,434,654,681]" italics="true" pageId="2" pageNumber="484">Laccophilus leguyaderi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9E96F82925078977FE0DFD4A8E43FD4C" box="[441,535,655,681]" pageId="2" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FFCBCACB25078977FF23FD768DE9FD28" box="[151,445,690,717]" pageId="2" pageNumber="484" type="description">
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<paragraph id="B76E994025078977FF23FD768DE9FD28" blockId="2.[151,535,654,717]" box="[151,445,690,717]" pageId="2" pageNumber="484">
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<figureCitation id="2FEA85C525078977FF23FD768D54FD28" box="[151,256,691,717]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="2" pageNumber="484">Figures 1</figureCitation>
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,
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<figureCitation id="2FEA85C525078977FEBAFD778D4AFD28" box="[270,286,690,717]" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="2" pageNumber="484">5</figureCitation>
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,
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||
<figureCitation id="2FEA85C525078977FE9EFD768D30FD28" box="[298,356,691,717]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="2" pageNumber="484">9, 16</figureCitation>
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,
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<figureCitation id="2FEA85C525078977FEC4FD768DEDFD28" box="[368,441,691,717]" captionStart="FIGURES 23–30" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[157,1431,561,1885]" captionTargetId="figure-175@10.[157,1431,561,1885]" captionTargetPageId="10" captionText="FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="2" pageNumber="484">23, 24</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="FFCBCACB25078977FF23FD3E887CF97C" pageId="2" pageNumber="484" type="materials_examined">
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<paragraph id="B76E994025078977FF23FD3E8F7DFCDC" blockId="2.[151,1437,762,2013]" pageId="2" pageNumber="484">
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<materialsCitation id="07B9931D25078977FF23FD3E8F70FCDC" ID-GBIF-Occurrence="3019656331" country="Madagascar" elevation="1970" latitude="-22.15" location="Andringitra National Park" longLatPrecision="1261" longitude="46.883335" pageId="2" pageNumber="484" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
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<emphasis id="85A5455225078977FF23FD3E8D60FCF1" bold="true" box="[151,308,762,789]" pageId="2" pageNumber="484">
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<typeStatus id="686A27E225078977FF23FD3E8C86FCF0" box="[151,210,763,789]" pageId="2" pageNumber="484">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="CFC6D9D025078977FE8EFD3E8D97FCF0" box="[314,451,763,789]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
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, former province of
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<collectingRegion id="751557A225078977FD17FD3E8F61FCF0" box="[675,821,763,789]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
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,
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<location id="B20ECF9B25078977FC8AFD3E8836FCF0" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FC8AFD3E8836FCF0" box="[830,1122,763,789]" country="Madagascar" latitude="-22.15" longLatPrecision="1261" longitude="46.883335" name="Andringitra National Park" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Andringitra National Park</location>
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, ca.
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<locationDeviation id="A6DC806225078977FB26FD3E8C84FCDC" location="South of Antanifotsy" pageId="2" pageNumber="484">
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<quantity id="702934A525078977FB26FD3E888BFCF0" box="[1170,1247,763,789]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="484" unit="km" value="4.2">4.2 km</quantity>
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South of Antanifotsy
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||
</locationDeviation>
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, GPS coordinates
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<geoCoordinate id="D2E5FF8725078977FE2AFCDA8DACFCDC" box="[414,504,799,825]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.15">S22°09’</geoCoordinate>
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<geoCoordinate id="D2E5FF8725078977FE4BFCDB8E0BFCDC" box="[511,607,798,825]" degrees="46" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.883335">E46°53’</geoCoordinate>
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, altitude
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<quantity id="702934A525078977FD71FCDA8F70FCDC" box="[709,804,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="484" unit="m" value="1970.0">
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<elevation id="3CFC7E7325078977FD71FCDA8F70FCDC" box="[709,804,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="484" unit="m" value="1970.0">1,970 m</elevation>
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</quantity>
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</materialsCitation>
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.
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||
</paragraph>
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<paragraph id="B76E994025078977FF73FC86887CF97C" blockId="2.[151,1437,762,2013]" pageId="2" pageNumber="484">
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<emphasis id="85A5455225078977FF73FC868D23FCB9" bold="true" box="[199,375,834,861]" pageId="2" pageNumber="484">Type material.</emphasis>
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<materialsCitation id="07B9931D25078977FE34FC868D78FC2C" ID-GBIF-Occurrence="3019656351" collectingDate="2016-05-24" collectionCode="MNHN" collectorName="Manuel" country="Madagascar" elevation="1970" latitude="-22.15" location="Fianarantsoa" longLatPrecision="1261" longitude="46.883335" pageId="2" pageNumber="484" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
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<emphasis id="85A5455225078977FE34FC868DB9FCB8" bold="true" box="[384,493,835,861]" pageId="2" pageNumber="484">
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<typeStatus id="686A27E225078977FE34FC868DB9FCB8" box="[384,493,835,861]" pageId="2" pageNumber="484" type="holotype">Holotype</typeStatus>
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</emphasis>
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(
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<specimenCount id="A1D752C925078977FDB4FC868E4CFCB8" box="[512,536,835,861]" pageId="2" pageNumber="484" type="male">♂</specimenCount>
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): ”
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<collectingCountry id="CFC6D9D025078977FD8DFC868E91FCB8" box="[569,709,835,861]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
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. Ex-prov. /
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<collectingRegion id="751557A225078977FCF9FC868FB4FCB8" box="[845,992,835,861]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
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||
. ca
|
||
<quantity id="702934A525078977FBBAFC86880AFCB8" box="[1038,1118,835,861]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="484" unit="km" value="4.2">4.2 km</quantity>
|
||
S / Antanifotsy.
|
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<date id="C36FBF8025078977FA96FC8689C8FCB8" box="[1314,1436,835,861]" pageId="2" pageNumber="484" value="2016-05-24">
|
||
<collectingDate id="D32B466825078977FA96FC8689C8FCB8" box="[1314,1436,835,861]" pageId="2" pageNumber="484" value="2016-05-24">24 V 2016</collectingDate>
|
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</date>
|
||
/ Ramahandrison &
|
||
<collectorName id="1A24FC9625078977FECAFCA28D80FC64" box="[382,468,871,897]" pageId="2" pageNumber="484">Manuel</collectorName>
|
||
leg. //
|
||
<geoCoordinate id="D2E5FF8725078977FD92FCA28ED5FC64" box="[550,641,871,897]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.15">S22°09’</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FD3DFCA38EB0FC64" box="[649,740,870,897]" degrees="46" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.883335">E46°53’</geoCoordinate>
|
||
/ Alt.
|
||
<quantity id="702934A525078977FC9BFCA28FD3FC64" box="[815,903,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="484" unit="m" value="1970.0">
|
||
<elevation id="3CFC7E7325078977FC9BFCA28FD3FC64" box="[815,903,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="484" unit="m" value="1970.0">1,970m</elevation>
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||
</quantity>
|
||
. Slowly flowing / rather deep brooklet, peaty / bottom. Andringitra NP” [pr], ”
|
||
<typeStatus id="686A27E225078977FE4EFC4E8E3CFC40" box="[506,616,907,933]" pageId="2" pageNumber="484" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<taxonomicName id="70D1E2C325078977FDC7FC4E8F2EFC40" authority="Manuel & Ramahandrison, 2020" authorityName="Manuel & Ramahandrison" authorityYear="2020" box="[627,890,907,933]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="484" phylum="Arthropoda" rank="species" species="leguyaderi" status="sp. nov.">Laccophilus leguyaderi</taxonomicName>
|
||
<taxonomicNameLabel id="9E96F82925078977FC37FC4E8F89FC40" box="[899,989,907,933]" pageId="2" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Manuel & Ramahandrison 2020” [red, pr] (
|
||
<collectionCode id="D1C0018525078977FF7CFC6A8D70FC2C" box="[200,292,943,969]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="484">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FE8CFC6A89CDFC2C" ID-GBIF-Occurrence="3019656326" box="[312,1433,943,969]" collectionCode="BMNH, CMM, DEUA, MNHN, NHRS, NMPC, PBZT, MBC" country="Madagascar" pageId="2" pageNumber="484" specimenCount="95" specimenCount-female="47" specimenCount-male="48" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<emphasis id="85A5455225078977FE8CFC6A8DE4FC2C" bold="true" box="[312,432,943,969]" pageId="2" pageNumber="484">
|
||
<typeStatus id="686A27E225078977FE8CFC6A8DE4FC2C" box="[312,432,943,969]" pageId="2" pageNumber="484" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(in total
|
||
<specimenCount id="A1D752C925078977FDADFC6A8E35FC2C" box="[537,609,943,969]" pageId="2" pageNumber="484" type="male">48♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FDDAFC6A8EE0FC2C" box="[622,692,943,969]" pageId="2" pageNumber="484" type="female">47♀♀</specimenCount>
|
||
) (
|
||
<collectionCode id="D1C0018525078977FD7AFC6A8F72FC2C" box="[718,806,943,969]" pageId="2" pageNumber="484">BMNH</collectionCode>
|
||
,
|
||
<collectionCode id="D1C0018525078977FC86FC6A8F2DFC2C" box="[818,889,943,969]" country="Brazil" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="2" pageNumber="484">CMM</collectionCode>
|
||
,
|
||
<collectionCode id="D1C0018525078977FC32FC6A8F83FC2C" box="[902,983,943,969]" pageId="2" pageNumber="484">DEUA</collectionCode>
|
||
,
|
||
<collectionCode id="D1C0018525078977FC57FC6A886FFC2C" box="[995,1083,943,969]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="484">MNHN</collectionCode>
|
||
,
|
||
<collectionCode id="D1C0018525078977FBFCFC6A88C1FC2C" box="[1096,1173,943,969]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="2" pageNumber="484">NHRS</collectionCode>
|
||
,
|
||
<collectionCode id="D1C0018525078977FB16FC6A88A0FC2C" box="[1186,1268,943,969]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="484">NMPC</collectionCode>
|
||
,
|
||
<collectionCode id="D1C0018525078977FAB5FC6A891DFC2C" box="[1281,1353,943,969]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="2" pageNumber="484">PBZT</collectionCode>
|
||
/
|
||
<collectionCode id="D1C0018525078977FAE5FC6A89C5FC2C" box="[1361,1425,943,969]" country="USA" name="Montgomery Botanical Center" pageId="2" pageNumber="484">MBC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="07B9931D25078977FF23FC168E6EFC08" ID-GBIF-Occurrence="3019656359" box="[151,570,979,1005]" collectingDate="2016-05-24" collectionCode="MNHN" collectorName="Manuel" country="Madagascar" elevation="1970" latitude="-22.15" location="Fianarantsoa" longLatPrecision="1261" longitude="46.883335" pageId="2" pageNumber="484" specimenCount="29" specimenCount-female="13" specimenCount-male="16" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FF23FC168CB4FC08" box="[151,224,979,1005]" pageId="2" pageNumber="484" type="male">16♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FF5FFC168D67FC08" box="[235,307,979,1005]" pageId="2" pageNumber="484" type="female">13♀♀</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FDF0FC178D03FBD0" ID-GBIF-Occurrence="3019656325" collectingDate="2016-05-21" collectorName="Manuel" country="Madagascar" county="Ex-Prov." elevation="1620" latitude="-22.133333" location="Small" longLatPrecision="1261" longitude="46.883335" municipality="Alt." pageId="2" pageNumber="484" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FDF0FC178E29FC08" box="[580,637,978,1005]" pageId="2" pageNumber="484" type="male">5♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FD3DFC168E96FC08" box="[649,706,979,1005]" pageId="2" pageNumber="484" type="female">3♀♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FD6DFC168F31FC08" box="[729,869,979,1005]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
.
|
||
<collectingCounty id="5E0FE1CC25078977FCC4FC168F87FC08" box="[880,979,979,1005]" pageId="2" pageNumber="484">Ex-Prov.</collectingCounty>
|
||
/
|
||
<collectingRegion id="751557A225078977FC5CFC16882FFC08" box="[1000,1147,979,1005]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
. ca
|
||
<quantity id="702934A525078977FB11FC1788A7FC08" box="[1189,1267,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="484" unit="km" value="2.5">2.5 km</quantity>
|
||
/ SSW Antanifotsy.
|
||
<date id="C36FBF8025078977FF6FFC328D1BFBF4" box="[219,335,1015,1041]" pageId="2" pageNumber="484" value="2016-05-21">
|
||
<collectingDate id="D32B466825078977FF6FFC328D1BFBF4" box="[219,335,1015,1041]" pageId="2" pageNumber="484" value="2016-05-21">21 V 2016</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="1A24FC9625078977FED7FC328DEDFBF4" box="[355,441,1015,1041]" pageId="2" pageNumber="484">Manuel</collectorName>
|
||
leg. //
|
||
<geoCoordinate id="D2E5FF8725078977FDB1FC328E0BFBF4" box="[517,607,1015,1041]" degrees="22" direction="south" minutes="08" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.133333">S22°08’</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FDD1FC338E94FBF4" box="[613,704,1014,1041]" degrees="46" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.883335">E46°53’</geoCoordinate>
|
||
/
|
||
<collectingMunicipality id="570A033A25078977FD60FC328EABFBF4" box="[724,767,1015,1041]" pageId="2" pageNumber="484">Alt.</collectingMunicipality>
|
||
<quantity id="702934A525078977FCB1FC328F09FBF4" box="[773,861,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="2" pageNumber="484" unit="m" value="1620.0">
|
||
<elevation id="3CFC7E7325078977FCB1FC328F09FBF4" box="[773,861,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="2" pageNumber="484" unit="m" value="1620.0">1,620m</elevation>
|
||
</quantity>
|
||
.
|
||
<location id="B20ECF9B25078977FCD3FC328FFCFBF4" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FCD3FC328FFCFBF4" box="[871,936,1015,1041]" country="Madagascar" county="Ex-Prov." latitude="-22.133333" longLatPrecision="1261" longitude="46.883335" municipality="Alt." name="Small" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Small</location>
|
||
stream / (just downstream spring) / in forest. Andringitra NP”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FED6FBDE8D88FB98" ID-GBIF-Occurrence="3019656357" collectingDate="2016-05-22" collectorName="Manuel" country="Madagascar" county="Ex-prov." elevation="1970" latitude="-22.15" location="Narrow lake" longLatPrecision="1261" longitude="46.883335" municipality="Antanifotsy." pageId="2" pageNumber="484" specimenCount="3" specimenCount-female="3" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FED6FBDE8DCFFBD0" box="[354,411,1051,1077]" pageId="2" pageNumber="484" type="female">3♀♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FE00FBDE8E14FBD0" box="[436,576,1051,1077]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
.
|
||
<collectingCounty id="5E0FE1CC25078977FDF8FBDE8EFAFBD0" box="[588,686,1051,1077]" pageId="2" pageNumber="484">Ex-prov.</collectingCounty>
|
||
/
|
||
<collectingRegion id="751557A225078977FD72FBDE8F0DFBD0" box="[710,857,1051,1077]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
. ca
|
||
<quantity id="702934A525078977FC32FBDE8F82FBD0" box="[902,982,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="484" unit="km" value="4.2">4.2 km</quantity>
|
||
S /
|
||
<collectingMunicipality id="570A033A25078977FBB1FBDE88C4FBD0" box="[1029,1168,1051,1077]" pageId="2" pageNumber="484">Antanifotsy.</collectingMunicipality>
|
||
<date id="C36FBF8025078977FB23FBDE8944FBD0" box="[1175,1296,1051,1077]" pageId="2" pageNumber="484" value="2016-05-22">
|
||
<collectingDate id="D32B466825078977FB23FBDE8944FBD0" box="[1175,1296,1051,1077]" pageId="2" pageNumber="484" value="2016-05-22">22 V 2016</collectingDate>
|
||
</date>
|
||
/ Ramahandrison &
|
||
<collectorName id="1A24FC9625078977FEB7FBFA8D0DFBBC" box="[259,345,1087,1113]" pageId="2" pageNumber="484">Manuel</collectorName>
|
||
leg. /
|
||
<geoCoordinate id="D2E5FF8725078977FE17FBFA8DA9FBBC" box="[419,509,1087,1113]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.15">S22°09’</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FDB1FBFB8E34FBBC" box="[517,608,1086,1113]" degrees="46" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.883335">E46°53’</geoCoordinate>
|
||
// Alt.
|
||
<quantity id="702934A525078977FD06FBFA8F5EFBBC" box="[690,778,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="484" unit="m" value="1970.0">
|
||
<elevation id="3CFC7E7325078977FD06FBFA8F5EFBBC" box="[690,778,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="484" unit="m" value="1970.0">1,970m</elevation>
|
||
</quantity>
|
||
.
|
||
<location id="B20ECF9B25078977FCA2FBFA8FF6FBBC" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FCA2FBFA8FF6FBBC" box="[790,930,1087,1113]" country="Madagascar" county="Ex-prov." latitude="-22.15" longLatPrecision="1261" longitude="46.883335" municipality="Antanifotsy." name="Narrow lake" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Narrow lake</location>
|
||
/ fed by brook, sandy bottom, / in subalpine grassland. / Andringitra NP”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FE53FBA68D98FB20" ID-GBIF-Occurrence="3019656327" collectingDate="2016-05-23" collectorName="Ramahandrison & Manuel" country="Madagascar" elevation="2000" latitude="-22.15" location="Antanifotsy." longLatPrecision="1261" longitude="46.883335" municipality="Ex-prov." pageId="2" pageNumber="484" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FE53FBA68E5EFB98" box="[487,522,1123,1149]" pageId="2" pageNumber="484" type="female">1♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FD96FBA68EFAFB98" box="[546,686,1123,1149]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
.
|
||
<collectingMunicipality id="570A033A25078977FD0DFBA68F4FFB98" box="[697,795,1123,1149]" pageId="2" pageNumber="484">Ex-prov.</collectingMunicipality>
|
||
/
|
||
<collectingRegion id="751557A225078977FC85FBA68F90FB98" box="[817,964,1123,1149]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
. ca
|
||
<quantity id="702934A525078977FC44FBA6886AFB98" box="[1008,1086,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="484" unit="km" value="4.3">4.3 km</quantity>
|
||
SW /
|
||
<location id="B20ECF9B25078977FB31FBA6895BFB98" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FB31FBA6895BFB98" box="[1157,1295,1123,1149]" country="Madagascar" latitude="-22.15" longLatPrecision="1261" longitude="46.883335" municipality="Ex-prov." name="Antanifotsy." pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Antanifotsy.</location>
|
||
<date id="C36FBF8025078977FAA2FBA689D9FB98" box="[1302,1421,1123,1149]" pageId="2" pageNumber="484" value="2016-05-23">
|
||
<collectingDate id="D32B466825078977FAA2FBA689D9FB98" box="[1302,1421,1123,1149]" pageId="2" pageNumber="484" value="2016-05-23">23 V 2016</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="1A24FC9625078977FF23FB428D13FB44" box="[151,327,1159,1185]" pageId="2" pageNumber="484">Ramahandrison</collectorName>
|
||
&
|
||
<collectorName id="1A24FC9625078977FED9FB428D97FB44" box="[365,451,1159,1185]" pageId="2" pageNumber="484">Manuel</collectorName>
|
||
leg. //
|
||
<geoCoordinate id="D2E5FF8725078977FDA0FB428E3AFB44" box="[532,622,1159,1185]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.15">S22°09’</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FDC2FB438E85FB44" box="[630,721,1158,1185]" degrees="46" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.883335">E46°53’</geoCoordinate>
|
||
/ Alt.
|
||
<quantity id="702934A525078977FCAFFB428F20FB44" box="[795,884,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="484" unit="m" value="2000.0">
|
||
<elevation id="3CFC7E7325078977FCAFFB428F20FB44" box="[795,884,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="484" unit="m" value="2000.0">2,000m</elevation>
|
||
</quantity>
|
||
. Shallow brooklet, / peaty bottom, in subalpine / grassland. Andringitra NP”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FE62FB6E8EB8FAE8" ID-GBIF-Occurrence="3019656363" collectingDate="2016-05-24" collectorName="Ramahandrison & Manuel" country="Madagascar" elevation="2000" latitude="-22.15" location="Antanifotsy." longLatPrecision="1261" longitude="46.9" municipality="Ex-prov." pageId="2" pageNumber="484" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FE62FB6E8DAEFB20" box="[470,506,1195,1221]" pageId="2" pageNumber="484" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FDB3FB6E8E7EFB20" box="[519,554,1195,1221]" pageId="2" pageNumber="484" type="female">1♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FDF6FB6E8E9AFB20" box="[578,718,1195,1221]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
.
|
||
<collectingMunicipality id="570A033A25078977FD6DFB6E8F6EFB20" box="[729,826,1195,1221]" pageId="2" pageNumber="484">Ex-prov.</collectingMunicipality>
|
||
/
|
||
<collectingRegion id="751557A225078977FCE4FB6E8FB7FB20" box="[848,995,1195,1221]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
. ca
|
||
<quantity id="702934A525078977FBB9FB6E8808FB20" box="[1037,1116,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="484" unit="km" value="4.3">4.3 km</quantity>
|
||
S /
|
||
<location id="B20ECF9B25078977FB33FB6E8945FB20" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FB33FB6E8945FB20" box="[1159,1297,1195,1221]" country="Madagascar" latitude="-22.15" longLatPrecision="1261" longitude="46.9" municipality="Ex-prov." name="Antanifotsy." pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Antanifotsy.</location>
|
||
<date id="C36FBF8025078977FAA3FB6E89D9FB20" box="[1303,1421,1195,1221]" pageId="2" pageNumber="484" value="2016-05-24">
|
||
<collectingDate id="D32B466825078977FAA3FB6E89D9FB20" box="[1303,1421,1195,1221]" pageId="2" pageNumber="484" value="2016-05-24">24 V 2016</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="1A24FC9625078977FF23FB0A8D13FB0C" box="[151,327,1231,1257]" pageId="2" pageNumber="484">Ramahandrison</collectorName>
|
||
&
|
||
<collectorName id="1A24FC9625078977FED9FB0A8D97FB0C" box="[365,451,1231,1257]" pageId="2" pageNumber="484">Manuel</collectorName>
|
||
leg. /
|
||
<geoCoordinate id="D2E5FF8725078977FDBFFB0A8E0BFB0C" box="[523,607,1231,1257]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.15">S22°09</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FDD2FB0B8E95FB0C" box="[614,705,1230,1257]" degrees="46" direction="east" minutes="54" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.9">E46°54’</geoCoordinate>
|
||
// Alt.
|
||
<quantity id="702934A525078977FCA7FB0A8F3FFB0C" box="[787,875,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="484" unit="m" value="2000.0">
|
||
<elevation id="3CFC7E7325078977FCA7FB0A8F3FFB0C" box="[787,875,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="484" unit="m" value="2000.0">2,000m</elevation>
|
||
</quantity>
|
||
. Slowly flowing / stream with vertical bank / and peaty bottom. Subalpine / grassland. Andringitra NP”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FD42FB368ED8FAB0" ID-GBIF-Occurrence="3019656314" collectingDate="2016-05-24" collectorName="Ramahandrison & Manuel" country="Madagascar" elevation="2000" latitude="-22.15" location="Antanifotsy." longLatPrecision="1261" longitude="46.9" municipality="Ex-prov." pageId="2" pageNumber="484" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FD42FB368F64FAE8" box="[758,816,1267,1293]" pageId="2" pageNumber="484" type="male">4♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FC8FFB368F27FAE8" box="[827,883,1267,1293]" pageId="2" pageNumber="484" type="female">1♀♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FC3EFB368842FAE8" box="[906,1046,1267,1293]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
.
|
||
<collectingMunicipality id="570A033A25078977FBABFB3688D5FAE8" box="[1055,1153,1267,1293]" pageId="2" pageNumber="484">Ex-prov.</collectingMunicipality>
|
||
/
|
||
<collectingRegion id="751557A225078977FB20FB368973FAE8" box="[1172,1319,1267,1293]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
. ca
|
||
<quantity id="702934A525078977FAFBFB3689C8FAE8" box="[1359,1436,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="484" unit="km" value="4.2">4.2 km</quantity>
|
||
S /
|
||
<location id="B20ECF9B25078977FF09FAD28D13FAD4" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FF09FAD28D13FAD4" box="[189,327,1303,1329]" country="Madagascar" latitude="-22.15" longLatPrecision="1261" longitude="46.9" municipality="Ex-prov." name="Antanifotsy." pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Antanifotsy.</location>
|
||
<date id="C36FBF8025078977FEFAFAD28D91FAD4" box="[334,453,1303,1329]" pageId="2" pageNumber="484" value="2016-05-24">
|
||
<collectingDate id="D32B466825078977FEFAFAD28D91FAD4" box="[334,453,1303,1329]" pageId="2" pageNumber="484" value="2016-05-24">24 V 2016</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="1A24FC9625078977FE69FAD28ED9FAD4" box="[477,653,1303,1329]" pageId="2" pageNumber="484">Ramahandrison</collectorName>
|
||
&
|
||
<collectorName id="1A24FC9625078977FD06FAD28F5CFAD4" box="[690,776,1303,1329]" pageId="2" pageNumber="484">Manuel</collectorName>
|
||
leg. //
|
||
<geoCoordinate id="D2E5FF8725078977FCECFAD38FE4FAD4" box="[856,944,1302,1329]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="2" pageNumber="484" precision="925" value="-22.15">S22°09’</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="D2E5FF8725078977FC0EFAD38843FAD4" box="[954,1047,1302,1329]" degrees="46" direction="east" minutes="54" orientation="longitude" pageId="2" pageNumber="484" precision="925" value="46.9">E46°54’</geoCoordinate>
|
||
/ Alt.
|
||
<quantity id="702934A525078977FBD4FAD288EDFAD4" box="[1120,1209,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="484" unit="m" value="2000.0">
|
||
<elevation id="3CFC7E7325078977FBD4FAD288EDFAD4" box="[1120,1209,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="484" unit="m" value="2000.0">2,000m</elevation>
|
||
</quantity>
|
||
. Margin of / narrow lake, subalpine / grassland. Andringitra NP”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FD22FAFF8E80FA24" ID-GBIF-Occurrence="3019656354" collectingDate="2013-12-03" collectorName="Leg. J. H. Randriamihaja & T. Ranarilalatiana" country="Madagascar" county="Haute" elevation="1542" latitude="-22.141115" location="Amporomahery" longLatPrecision="1" longitude="46.94196" municipality="Matsiara" pageId="2" pageNumber="484" specimenCount="28" specimenCount-female="13" specimenCount-male="15" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FD22FAFF8EB2FAB0" box="[662,742,1338,1365]" pageId="2" pageNumber="484" type="male">15 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FD47FAFE8F15FAB0" box="[755,833,1339,1365]" pageId="2" pageNumber="484" type="female">13 ♀♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FCEDFAFE8FB3FAB0" box="[857,999,1339,1365]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
:
|
||
<collectingRegion id="751557A225078977FC46FAFE88D2FAB0" box="[1010,1158,1339,1365]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
:
|
||
<collectingCounty id="5E0FE1CC25078977FB24FAFE8887FAB0" box="[1168,1235,1339,1365]" pageId="2" pageNumber="484">Haute</collectingCounty>
|
||
/
|
||
<collectingMunicipality id="570A033A25078977FB5EFAFE8904FAB0" box="[1258,1360,1339,1365]" pageId="2" pageNumber="484">Matsiara</collectingMunicipality>
|
||
: Andringitra PN:
|
||
<location id="B20ECF9B25078977FEADFA9A8D9EFA9C" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FEADFA9A8D9EFA9C" box="[281,458,1375,1401]" country="Madagascar" county="Haute" latitude="-22.141115" longLatPrecision="1" longitude="46.94196" municipality="Matsiara" name="Amporomahery" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Amporomahery</location>
|
||
/ (circuit
|
||
<location id="B20ECF9B25078977FD8EFA9A8EC6FA9F" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FD8EFA9A8EC6FA9F" box="[570,658,1375,1402]" country="Madagascar" county="Haute" latitude="-22.141115" longLatPrecision="1" longitude="46.94196" municipality="Matsiara" name="Imaitso" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Imaitso</location>
|
||
),
|
||
<geoCoordinate id="D2E5FF8725078977FD12FA9A8F7EFA9C" box="[678,810,1375,1401]" degrees="22.141116" direction="south" orientation="latitude" pageId="2" pageNumber="484" precision="1" value="-22.141115">S22.141116</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FC86FA9A8FFBFA9C" box="[818,943,1375,1401]" degrees="46.94196" direction="east" orientation="longitude" pageId="2" pageNumber="484" precision="1" value="46.94196">E46.94196</geoCoordinate>
|
||
, /
|
||
<quantity id="702934A525078977FC7DFA9B884EFA9C" box="[969,1050,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.5419999999999998" pageId="2" pageNumber="484" unit="m" value="1542.0">
|
||
<elevation id="3CFC7E7325078977FC7DFA9B884EFA9C" box="[969,1050,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.5419999999999998" pageId="2" pageNumber="484" unit="m" value="1542.0">1542m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="C36FBF8025078977FB92FA9A88E6FA9C" box="[1062,1202,1375,1401]" pageId="2" pageNumber="484" value="2013-12-03">
|
||
<collectingDate id="D32B466825078977FB92FA9A88E6FA9C" box="[1062,1202,1375,1401]" pageId="2" pageNumber="484" value="2013-12-03">03.XII.2013</collectingDate>
|
||
</date>
|
||
, GB nets and sieves: /
|
||
<location id="B20ECF9B25078977FF13FA468DC9FA78" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FF13FA468DC9FA78" box="[167,413,1411,1437]" country="Madagascar" county="Haute" latitude="-22.141115" longLatPrecision="1" longitude="46.94196" municipality="Matsiara" name="River Amporomahery" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">River Amporomahery</location>
|
||
near tourist trail, little / loop (petit boucle),
|
||
<collectorName id="1A24FC9625078977FC27FA4688F4FA78" box="[915,1184,1411,1437]" pageId="2" pageNumber="484">Leg. J.H.Randriamihaja</collectorName>
|
||
/ &
|
||
<collectorName id="1A24FC9625078977FB61FA4689CDFA78" box="[1237,1433,1411,1437]" pageId="2" pageNumber="484">T.Ranarilalatiana</collectorName>
|
||
: Field# MAD13-36 // NHRS-JLKB / 000070609”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FD56FA628DBFF9C8" ID-GBIF-Occurrence="3019656352" collectingDate="2013-12-01" collectorName="Leg. J. H. Randriamihaja & T. Ranarilalatiana" country="Madagascar" county="Haute" elevation="1657" latitude="-22.1394" location="Belambo" longLatPrecision="7" longitude="46.8876" municipality="Matsiara" pageId="2" pageNumber="484" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FD56FA628F72FA24" box="[738,806,1447,1473]" pageId="2" pageNumber="484" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FC82FA628F2EFA24" box="[822,890,1447,1473]" pageId="2" pageNumber="484" type="female">4 ♀♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FC20FA628876FA24" box="[916,1058,1447,1473]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
:
|
||
<collectingRegion id="751557A225078977FB85FA628891FA24" box="[1073,1221,1447,1473]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
:
|
||
<collectingCounty id="5E0FE1CC25078977FB66FA628941FA24" box="[1234,1301,1447,1473]" pageId="2" pageNumber="484">Haute</collectingCounty>
|
||
/
|
||
<collectingMunicipality id="570A033A25078977FA86FA6289CCFA24" box="[1330,1432,1447,1473]" pageId="2" pageNumber="484">Matsiara</collectingMunicipality>
|
||
: Andringitra PN:
|
||
<location id="B20ECF9B25078977FEE0FA0E8DEAFA00" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FEE0FA0E8DEAFA00" box="[340,446,1483,1509]" country="Madagascar" county="Haute" latitude="-22.1394" longLatPrecision="7" longitude="46.8876" municipality="Matsiara" name="Belambo" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Belambo</location>
|
||
: /
|
||
<location id="B20ECF9B25078977FE6CFA0E8EE0FA00" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FE6CFA0E8EE0FA00" box="[472,692,1483,1509]" country="Madagascar" county="Haute" latitude="-22.1394" longLatPrecision="7" longitude="46.8876" municipality="Matsiara" name="Cascade Riambavy" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Cascade Riambavy</location>
|
||
:
|
||
<geoCoordinate id="D2E5FF8725078977FD0BFA0E8F7DFA00" box="[703,809,1483,1509]" degrees="22.1394" direction="south" orientation="latitude" pageId="2" pageNumber="484" precision="5" value="-22.1394">S22.1394</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FC85FA0E8FF4FA00" box="[817,928,1483,1509]" degrees="46.8876" direction="east" orientation="longitude" pageId="2" pageNumber="484" precision="5" value="46.8876">E46.8876</geoCoordinate>
|
||
, /
|
||
<quantity id="702934A525078977FC0FFA0F8858FA00" box="[955,1036,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.657" pageId="2" pageNumber="484" unit="m" value="1657.0">
|
||
<elevation id="3CFC7E7325078977FC0FFA0F8858FA00" box="[955,1036,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.657" pageId="2" pageNumber="484" unit="m" value="1657.0">1657m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="C36FBF8025078977FBA3FA0E88F6FA00" box="[1047,1186,1483,1509]" pageId="2" pageNumber="484" value="2013-12-01">
|
||
<collectingDate id="D32B466825078977FBA3FA0E88F6FA00" box="[1047,1186,1483,1509]" pageId="2" pageNumber="484" value="2013-12-01">01.XII.2013</collectingDate>
|
||
</date>
|
||
, GB nets and sieves: / river from the cascade with pools on the / rocks,
|
||
<collectorName id="1A24FC9625078977FD09FA2A8F9DF9EC" box="[701,969,1519,1545]" pageId="2" pageNumber="484">Leg. J.H.Randriamihaja</collectorName>
|
||
& /
|
||
<collectorName id="1A24FC9625078977FC48FA2A8894F9EC" box="[1020,1216,1519,1545]" pageId="2" pageNumber="484">T.Ranarilalatiana</collectorName>
|
||
: Field# MAD13-30 // NHRS-JLKB / 000070619”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="07B9931D25078977FE41F9D78870F97C" ID-GBIF-Occurrence="3019656332" collectingDate="2013-12-02" collectorName="Leg. J. H. Randriamihaja & T. Ranarilalatiana" country="Madagascar" county="Haute Matsiara" elevation="1965" latitude="-22.151016" location="Riambavy" longLatPrecision="1" longitude="46.89163" municipality="Camp" pageId="2" pageNumber="484" specimenCount="13" specimenCount-female="8" specimenCount-male="5" stateProvince="Fianarantsoa" typeStatus="paratype">
|
||
<specimenCount id="A1D752C925078977FE41F9D78E63F9C8" box="[501,567,1554,1581]" pageId="2" pageNumber="484" type="male">5 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A1D752C925078977FDF7F9D68ED0F9C8" box="[579,644,1555,1581]" pageId="2" pageNumber="484" type="female">8 ♀♀</specimenCount>
|
||
, ”
|
||
<collectingCountry id="CFC6D9D025078977FD2FF9D68F7DF9C8" box="[667,809,1555,1581]" name="Madagascar" pageId="2" pageNumber="484">Madagascar</collectingCountry>
|
||
:
|
||
<collectingRegion id="751557A225078977FC80F9D68F9CF9C8" box="[820,968,1555,1581]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="484">Fianarantsoa</collectingRegion>
|
||
:
|
||
<collectingCounty id="5E0FE1CC25078977FC67F9D688D7F9C8" box="[979,1155,1555,1581]" pageId="2" pageNumber="484">Haute Matsiara</collectingCounty>
|
||
: / Andringitra PN:
|
||
<collectingMunicipality id="570A033A25078977FAEDF9D689C8F9C8" box="[1369,1436,1555,1581]" pageId="2" pageNumber="484">Camp</collectingMunicipality>
|
||
2, river running into / cascade
|
||
<location id="B20ECF9B25078977FE52F9F28E0AF9B4" LSID="urn:lsid:plazi:treatment:3F78285625078976FF23FD4B88E7F894:B20ECF9B25078977FE52F9F28E0AF9B4" box="[486,606,1591,1617]" country="Madagascar" county="Haute Matsiara" latitude="-22.151016" longLatPrecision="1" longitude="46.89163" municipality="Camp" name="Riambavy" pageId="2" pageNumber="484" stateProvince="Fianarantsoa">Riambavy</location>
|
||
:
|
||
<geoCoordinate id="D2E5FF8725078977FDD3F9F38EBAF9B4" box="[615,750,1590,1617]" degrees="22.151016" direction="south" orientation="latitude" pageId="2" pageNumber="484" precision="1" value="-22.151016">S22.151016</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725078977FD40F9F28F25F9B4" box="[756,881,1591,1617]" degrees="46.89163" direction="east" orientation="longitude" pageId="2" pageNumber="484" precision="1" value="46.89163">E46.89163</geoCoordinate>
|
||
, /
|
||
<quantity id="702934A525078977FC3DF9F38F8DF9B4" box="[905,985,1590,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.9649999999999999" pageId="2" pageNumber="484" unit="m" value="1965.0">
|
||
<elevation id="3CFC7E7325078977FC3DF9F38F8DF9B4" box="[905,985,1590,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.9649999999999999" pageId="2" pageNumber="484" unit="m" value="1965.0">1965m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="C36FBF8025078977FC57F9F2883BF9B4" box="[995,1135,1591,1617]" pageId="2" pageNumber="484" value="2013-12-02">
|
||
<collectingDate id="D32B466825078977FC57F9F2883BF9B4" box="[995,1135,1591,1617]" pageId="2" pageNumber="484" value="2013-12-02">02.XII.2013</collectingDate>
|
||
</date>
|
||
, GB nets and sieves: / river with pools surrounded by regular / burning area,
|
||
<collectorName id="1A24FC9625078977FD0AF99E8F9DF990" box="[702,969,1627,1653]" pageId="2" pageNumber="484">Leg. J.H.Randriamihaja</collectorName>
|
||
& /
|
||
<collectorName id="1A24FC9625078977FC48F99E8895F990" box="[1020,1217,1627,1653]" pageId="2" pageNumber="484">T.Ranarilalatiana</collectorName>
|
||
: Field# MAD13-35 // NHRS-JLKB / 000070623”. All
|
||
<typeStatus id="686A27E225078977FDA8F9BA8ED3F97C" box="[540,647,1663,1689]" pageId="2" pageNumber="484" type="paratype">paratypes</typeStatus>
|
||
with the respective printed red label
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FFCBCACB25078976FF73F9678E40FA9C" lastPageId="3" lastPageNumber="485" pageId="2" pageNumber="484" type="description">
|
||
<paragraph id="B76E994025078977FF73F9678F8AF8CC" blockId="2.[151,1437,762,2013]" pageId="2" pageNumber="484">
|
||
<emphasis id="85A5455225078977FF73F9678D8AF958" bold="true" box="[199,478,1698,1725]" pageId="2" pageNumber="484">
|
||
Description of
|
||
<typeStatus id="686A27E225078977FEC3F9668D8AF958" box="[375,478,1699,1725]" pageId="2" pageNumber="484" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="85A5455225078977FE5FF9678E13F958" bold="true" box="[491,583,1698,1725]" italics="true" pageId="2" pageNumber="484">Habitus</emphasis>
|
||
(
|
||
<figureCitation id="2FEA85C525078977FDE2F9668EC2F958" box="[598,662,1699,1725]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="2" pageNumber="484">Fig. 1</figureCitation>
|
||
) oval, with lateral sides widely rounded, posteriorly strongly tapered to rather narrowly rounded apex; lateral outline continuous between pronotum and elytra; maximum body width at about 41% of total body length and about 27% of elytral length; dorsal surface evenly and moderately convex; in lateral view maximum height slightly posterior to first third of total length.
|
||
</paragraph>
|
||
<paragraph id="B76E994025078976FF73F8F78D46FF30" blockId="2.[151,1437,762,2013]" lastBlockId="3.[151,1437,150,1906]" lastPageId="3" lastPageNumber="485" pageId="2" pageNumber="484">
|
||
<emphasis id="85A5455225078977FF73F8F78D0EF8A8" bold="true" box="[199,346,1842,1869]" italics="true" pageId="2" pageNumber="484">Colouration.</emphasis>
|
||
Dorsal surface of head rufo-testaceous. Pronotum rufo-testaceous, narrowly darkened along posterior margin. Elytron with typical
|
||
<emphasis id="85A5455225078977FDADF8928ED4F894" box="[537,640,1879,1905]" italics="true" pageId="2" pageNumber="484">alluaudi-</emphasis>
|
||
group colour pattern as in
|
||
<figureCitation id="2FEA85C525078977FC16F8928FB7F894" box="[930,995,1879,1905]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="2" pageNumber="484">Fig. 1</figureCitation>
|
||
; brown longitudinal vittae well-defined and rather strongly contrasting with testaceous background, leaving wide basal area and humeral region entirely testaceous; pale stripes between dark longitudinal vittae irregularly interrupted. Pronotal epipleura, propleura, lateral parts of prosternum, and diffusely delimited narrow area along lateral margin of metacoxal plate brown; rest of ventral surface rufo-testaceous (
|
||
<figureCitation id="2FEA85C525068976FDB6FF528E13FF54" box="[514,583,150,177]" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="3" pageNumber="485">Fig. 5</figureCitation>
|
||
). Antennae, palps, anterior and median legs testaceous; posterior legs rufotestaceous.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FF1B8E10FE80" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FF1B8D51FF1C" bold="true" box="[199,261,222,249]" italics="true" pageId="3" pageNumber="485">Head</emphasis>
|
||
. Anterior outline of clypeus widely rounded, subtruncated in middle. Surface shiny, with double reticulation; large meshes irregular and of variable sizes, more strongly impressed than small meshes; with sparse micropunctures, on vertex denser and coarser; larger punctures present along inner eye margins and in two irregular transverse rows sub-laterally on frons.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FEAA8D9CFE10" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FEAA8D68FE6C" bold="true" box="[199,316,367,393]" italics="true" pageId="3" pageNumber="485">Pronotum</emphasis>
|
||
. Lateral margins evenly arcuate. Surface shiny, with double reticulation; large meshes irregular and of variable sizes, more strongly impressed than small meshes; small meshes rounded, more strongly impressed than on head; punctation inconspicuous except in anterior quarter and close to posterior angles with sparse, moderately large and coarse punctures.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FE3B886CFD84" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FE3B8D71FDFC" bold="true" box="[199,293,510,537]" italics="true" pageId="3" pageNumber="485">Elytron.</emphasis>
|
||
Surface shiny, with double reticulation; large meshes indistinct except in scutellar region; small meshes irregularly polygonal, of rather homogeneous sizes, larger and much more strongly impressed than on pronotum; punctation inconspicuous beyond rather indistinct discal and lateral puncture rows.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FDAF8FFAFCB8" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FDAF8D23FD61" bold="true" box="[199,375,618,645]" italics="true" pageId="3" pageNumber="485">Ventral surface</emphasis>
|
||
. Prosternal process lanceolate, strongly convex, with apex acutely pointed. Metacoxal lines anteriorly subparallel. Apical abdominal ventrite strongly convex, in posterior half almost tectiform; without protuberance; posterior margin unmodified, straight on each side of slightly produced, rounded apex. Entire ventral surface very shiny; on elytral epipleura, metaventrite, metacoxa and abdominal ventrites with very fine regular microstriae; on abdominal ventrites with oblique striae, more strongly impressed on ventrites I–III; surface impunctate except on antero-medial prosternal region, anterior region of metaventrite and apical abdominal ventrite with shallow punctures of moderate size. Metacoxal plate without stridulatory apparatus.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FCA2897CFBF4" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FCA28D51FC64" bold="true" box="[199,261,871,897]" italics="true" pageId="3" pageNumber="485">Legs.</emphasis>
|
||
Mesotibia strongly modified, very broad with dorsal outline strongly curved, ventral outline straight (
|
||
<figureCitation id="2FEA85C525068976FADBFCA28CF3FC40" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="3" pageNumber="485">Fig. 5</figureCitation>
|
||
); along dorsal edge with row of very long setae; anterior surface mat, with very dense small punctures bearing short setae except in basal quarter and narrowly along apical margin (surface shiny, almost impunctate). Pro- and mesotarsomeres I–III rather strongly broadened, with four ventral transverse dense rows of adhesive setae. Protarsal claws about as long as protarsomere V; length of mesotarsal claws about 3/4 length of mesotarsomere V.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FBDF89C0FBD0" blockId="3.[151,1437,150,1906]" box="[199,1428,1050,1077]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FBDF8D63FBD0" bold="true" box="[199,311,1050,1077]" italics="true" pageId="3" pageNumber="485">Aedeagus</emphasis>
|
||
. Median lobe in lateral view as in
|
||
<figureCitation id="2FEA85C525068976FD0FFBDE8EABFBD0" box="[699,767,1051,1077]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="3" pageNumber="485">Fig. 9</figureCitation>
|
||
, in ventral view as in
|
||
<figureCitation id="2FEA85C525068976FC41FBDE8813FBD0" box="[1013,1095,1051,1077]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="3" pageNumber="485">Fig. 16</figureCitation>
|
||
; parameres as in
|
||
<figureCitation id="2FEA85C525068976FABCFBDE89C4FBD0" box="[1288,1424,1051,1077]" captionStart="FIGURES 23–30" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[157,1431,561,1885]" captionTargetId="figure-175@10.[157,1431,561,1885]" captionTargetPageId="10" captionText="FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="3" pageNumber="485">Figs. 23–24</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FBFA8910FB44" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FBFA8D64FBBC" bold="true" box="[199,304,1087,1113]" pageId="3" pageNumber="485">Females.</emphasis>
|
||
Mesotibia distinctly narrower (yet broader than usual in
|
||
<taxonomicName id="70D1E2C325068976FC02FBFA886BFBBC" authorityName="Leach" authorityYear="1815" box="[950,1087,1087,1113]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="85A5455225068976FC02FBFA886BFBBC" box="[950,1087,1087,1113]" italics="true" pageId="3" pageNumber="485">Laccophilus</emphasis>
|
||
</taxonomicName>
|
||
); with row of long setae along dorsal edge less developed; anterior surface more shiny, with small punctures distinctly larger and sparser and impunctate area along apical margin wider. Pro- and mesotarsi not broadened, without ventral adhesive setae.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FB6F8807FAE8" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FB6F8D19FB21" bold="true" box="[199,333,1194,1220]" pageId="3" pageNumber="485">Variability.</emphasis>
|
||
Specimens vary slightly in habitus (length / width ratio) and dorsal colour pattern mainly with respect to intensity of the elytral brown longitudinal markings and degree of fragmentation of the pale interspaces between these dark bands, the overall elytral pattern being nevertheless highly stable.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73FAD28E40FA9C" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FAD28D2AFAD4" bold="true" box="[199,382,1303,1329]" pageId="3" pageNumber="485">Measurements.</emphasis>
|
||
<typeStatus id="686A27E225068976FE37FAD28DA4FAD4" box="[387,496,1303,1329]" pageId="3" pageNumber="485" type="holotype">Holotype</typeStatus>
|
||
: TL =
|
||
<quantity id="702934A525068976FD80FAD28EDEFAD4" box="[564,650,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="485" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, TL without head =
|
||
<quantity id="702934A525068976FCD1FAD28FE8FAD4" box="[869,956,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="485" unit="mm" value="3.9">3.9 mm</quantity>
|
||
, MW =
|
||
<quantity id="702934A525068976FBA7FAD2883EFAD4" box="[1043,1130,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="485" unit="mm" value="2.3">2.3 mm</quantity>
|
||
, ratio TL/MW = 1.80.
|
||
<typeStatus id="686A27E225068976FAD5FAD28C82FAB0" pageId="3" pageNumber="485" type="paratype">Paratypes</typeStatus>
|
||
: TL =
|
||
<quantity id="702934A525068976FEA9FAFF8DE5FAB0" box="[285,433,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.225" metricValueMax="4.5" metricValueMin="3.95" pageId="3" pageNumber="485" unit="mm" value="4.225" valueMax="4.5" valueMin="3.95">3.95–4.5 mm</quantity>
|
||
(4.28±0.12), TL without head =
|
||
<quantity id="702934A525068976FCAFFAFE8FF5FAB0" box="[795,929,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="3" pageNumber="485" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.7–4.3 mm</quantity>
|
||
(4.06±0.11), MW =
|
||
<quantity id="702934A525068976FB31FAFF895FFAB0" box="[1157,1291,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="3" pageNumber="485" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 mm</quantity>
|
||
(2.37±0.08), TL/MW = 1.73–1.91 (1.81±0.04).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FFCBCACB25068976FF73FA4789C5F8E0" pageId="3" pageNumber="485" type="diagnosis">
|
||
<paragraph id="B76E994025068976FF73FA478DE6F9C8" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73FA478D84FA79" bold="true" box="[199,464,1410,1436]" pageId="3" pageNumber="485">Differential diagnosis.</emphasis>
|
||
This species and the following one differ from all other known
|
||
<taxonomicName id="70D1E2C325068976FB08FA46894DFA78" authorityName="Regimbart" authorityYear="1900" box="[1212,1305,1411,1437]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="485" phylum="Arthropoda" rank="species" species="alluaudi">
|
||
<emphasis id="85A5455225068976FB08FA46894DFA78" box="[1212,1305,1411,1437]" italics="true" pageId="3" pageNumber="485">alluaudi</emphasis>
|
||
</taxonomicName>
|
||
-group species by the combination of elytral colour pattern (notably with particularly broad pale area between anterior elytral margin and anterior extremity of the dark longitudinal markings), brown colouration of the ventral surface of the prothorax, and shape of the median lobe of aedeagus. For differences with
|
||
<taxonomicName id="70D1E2C325068976FC58FA2A8825F9EC" authority="Manuel & Ramahandrison, 2020" authorityName="Manuel & Ramahandrison" authorityYear="2020" box="[1004,1137,1519,1545]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="485" phylum="Arthropoda" rank="species" species="bergsteni" status="sp. nov.">
|
||
<emphasis id="85A5455225068976FC58FA2A8825F9EC" box="[1004,1137,1519,1545]" italics="true" pageId="3" pageNumber="485">L. bergsteni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="85A5455225068976FBCEFA2A8881F9EC" bold="true" box="[1146,1237,1519,1545]" pageId="3" pageNumber="485">
|
||
<taxonomicNameLabel id="9E96F82925068976FBCEFA2A8881F9EC" box="[1146,1237,1519,1545]" pageId="3" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, see below under diagnosis of
|
||
<taxonomicName id="70D1E2C325068976FE92F9D68DFFF9C8" authorityName="Manuel & Ramahandrison" authorityYear="2020" box="[294,427,1555,1581]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="485" phylum="Arthropoda" rank="species" species="bergsteni">
|
||
<emphasis id="85A5455225068976FE92F9D68DFFF9C8" box="[294,427,1555,1581]" italics="true" pageId="3" pageNumber="485">L. bergsteni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B76E994025068976FF73F9F389C5F8E0" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73F9F38D7EF9B5" bold="true" box="[199,298,1590,1616]" pageId="3" pageNumber="485">Habitat.</emphasis>
|
||
This species was collected at an altitude range of
|
||
<quantity id="702934A525068976FCD0F9F38845F9B4" box="[868,1041,1590,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.765" metricValueMax="1.9899999999999998" metricValueMin="1.54" pageId="3" pageNumber="485" unit="m" value="1765.0" valueMax="1990.0" valueMin="1540.0">
|
||
<geoCoordinate id="D2E5FF8725068976FCD0F9F38FF7F9B4" box="[868,931,1590,1617]" degrees="1,540" direction="north" orientation="latitude" pageId="3" pageNumber="485" precision="55" value="1.54">1,540</geoCoordinate>
|
||
<geoCoordinate id="D2E5FF8725068976FC17F9F38FA5F9B4" box="[931,1009,1590,1617]" degrees="1,990" direction="west" orientation="longitude" pageId="3" pageNumber="485" precision="55" value="-1.99">–1,990</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
. It lives at shallow depth along the margins of slowly-flowing brooks, streams or rivers and associated pools, as well as small to large lentic water bodies with inflow and outflow (
|
||
<figureCitation id="2FEA85C525068976FE6FF9BA8E31F97C" box="[475,613,1663,1689]" captionStart="FIGURES 43–46" captionStartId="15.[151,264,1505,1529]" captionTargetBox="[157,1431,524,1481]" captionTargetId="figure-163@15.[157,1431,524,1481]" captionTargetPageId="15" captionText="FIGURES 43–46. Habitats of Laccophilus leguyaderi sp. nov. (43–44) and Laccophilus bergsteni sp. nov. (45–46). (43) Shallow brooklet with peaty bottom in subalpine grassland (Andringitra massif, 2,000 m); (44) Rather deep brook with peaty bottom, slowly flowing among subalpine shrubs (Andringitra massif, 1,970 m; type locality of L. leguyaderi); (45) Slowly flowing stream in natural forest with muddy bottom and lots of tree leaves (Ankaratra massif, Tavolotara, 2,000 m); (46) Slowly flowing stream on flat area in subalpine grassland (Ankaratra massif, 2,400 m; type locality of L. bergsteni)." figureDoi="http://doi.org/10.5281/zenodo.4401801" httpUri="https://zenodo.org/record/4401801/files/figure.png" pageId="3" pageNumber="485">Figs. 43–44</figureCitation>
|
||
). Some collecting sites were located in forest, but the species is also present above tree line in subalpine grassland (e.g., sampling site shown in
|
||
<figureCitation id="2FEA85C525068976FC02F966885EF958" box="[950,1034,1699,1725]" captionStart="FIGURES 43–46" captionStartId="15.[151,264,1505,1529]" captionTargetBox="[157,1431,524,1481]" captionTargetId="figure-163@15.[157,1431,524,1481]" captionTargetPageId="15" captionText="FIGURES 43–46. Habitats of Laccophilus leguyaderi sp. nov. (43–44) and Laccophilus bergsteni sp. nov. (45–46). (43) Shallow brooklet with peaty bottom in subalpine grassland (Andringitra massif, 2,000 m); (44) Rather deep brook with peaty bottom, slowly flowing among subalpine shrubs (Andringitra massif, 1,970 m; type locality of L. leguyaderi); (45) Slowly flowing stream in natural forest with muddy bottom and lots of tree leaves (Ankaratra massif, Tavolotara, 2,000 m); (46) Slowly flowing stream on flat area in subalpine grassland (Ankaratra massif, 2,400 m; type locality of L. bergsteni)." figureDoi="http://doi.org/10.5281/zenodo.4401801" httpUri="https://zenodo.org/record/4401801/files/figure.png" pageId="3" pageNumber="485">Fig. 43</figureCitation>
|
||
). It prefers habitats with substratum moderately to heavily loaded with organic matter (most often including a fine brown organic mud component, together with macroscopic vegetal remnants of trees and/or helophytes) and clear water, with or without vegetation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FFCBCACB25068976FF73F8CB891AF8CC" box="[199,1358,1806,1833]" pageId="3" pageNumber="485" type="distribution">
|
||
<paragraph id="B76E994025068976FF73F8CB891AF8CC" blockId="3.[151,1437,150,1906]" box="[199,1358,1806,1833]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73F8CB8D35F8CD" bold="true" box="[199,353,1806,1832]" pageId="3" pageNumber="485">Distribution.</emphasis>
|
||
So far known only from the Andringitra massif in central-southern
|
||
<collectingCountry id="CFC6D9D025068976FBE3F8CA888AF8CC" box="[1111,1246,1807,1833]" name="Madagascar" pageId="3" pageNumber="485">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="2FEA85C525068976FB59F8CA8915F8CC" box="[1261,1345,1806,1833]" captionStart="FIGURE 53" captionStartId="18.[151,250,1530,1554]" captionTargetBox="[391,1197,597,1507]" captionTargetId="figure-243@18.[391,1197,597,1507]" captionTargetPageId="18" captionText="FIGURE 53. Map showing the distribution of known localities of the newly described species. Red dot: L. leguyaderi; Blue dots: L. bergsteni; Magenta dot: L. bergsteni and L. rakouthae; yellow dot: L. makay. Names of corresponding protected areas or massifs indicated in italics above dots." figureDoi="http://doi.org/10.5281/zenodo.4401805" httpUri="https://zenodo.org/record/4401805/files/figure.png" pageId="3" pageNumber="485">Fig. 53</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FFCBCACB25068976FF73F8F788E7F894" pageId="3" pageNumber="485" type="etymology">
|
||
<paragraph id="B76E994025068976FF73F8F788E7F894" blockId="3.[151,1437,150,1906]" pageId="3" pageNumber="485">
|
||
<emphasis id="85A5455225068976FF73F8F78DF1F8A9" bold="true" box="[199,421,1842,1868]" pageId="3" pageNumber="485">Derivatio nominis.</emphasis>
|
||
This species is dedicated to Prof. Hervé Le Guyader (Institut de Systématique, Evolution, Biodiversité, Sorbonne Université, Paris). The species name is a noun in the genitive singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |