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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.609.6822" ID-PMC="PMC4984416" ID-Pensoft-Pub="1313-2970-609-11" ID-Pensoft-UUID="FFEB90038106FFF4FFB72D58FFA5FFF8" ID-PubMed="27563269" ID-Zenodo-Dep="144490" ID-ZooBank="91E491A407C5448CBA831E4FAB807066" ModsDocID="1313-2970-609-11" checkinTime="1470750541503" checkinUser="pensoft" docAuthor="Haeckel, Martin, Farkac, Jan &amp; Sehnal, Rostislav" docDate="2016" docId="3B7A508A305255EC82AEE715DA8AE8F9" docLanguage="en" docName="ZooKeys 609: 11-28" docOrigin="ZooKeys 609" docPubDate="2016-08-08" docSource="http://dx.doi.org/10.3897/zookeys.609.6822" docTitle="Calosoma (Calosoma) scabrosum subsp. roeschkei Breuning 1927" docType="treatment" docVersion="4" id="FFEB90038106FFF4FFB72D58FFA5FFF8" lastPageNumber="13" masterDocId="FFEB90038106FFF4FFB72D58FFA5FFF8" masterDocTitle="Calosoma aethiops (Jeannel, 1940) as a new synonym of Calosoma imbricatum hottentotum Chaudoir, 1852, a new status of Calosoma roeschkei Breuning, 1927, and a revision of the Calosoma senegalense group sensu Haeckel, 2012 (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="28" masterPageNumber="11" pageNumber="13" updateTime="1668163397258" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Calosoma aethiops (Jeannel, 1940) as a new synonym of Calosoma imbricatum hottentotum Chaudoir, 1852, a new status of Calosoma roeschkei Breuning, 1927, and a revision of the Calosoma senegalense group sensu Haeckel, 2012 (Coleoptera, Carabidae, Carabini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haeckel, Martin</mods:namePart>
<mods:affiliation>Department of Game Management and Wildlife Biology, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamycka 1176, CZ- 165 21 Prague 6 - Suchdol, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Farkac, Jan</mods:namePart>
<mods:affiliation>Department of Game Management and Wildlife Biology, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamycka 1176, CZ- 165 21 Prague 6 - Suchdol, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sehnal, Rostislav</mods:namePart>
<mods:affiliation>V Zahradkach 962, CZ- 273 51 Unhost, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-08-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>609</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>28</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.609.6822</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.609.6822</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-609-11</mods:identifier>
<mods:identifier type="ZooBank">91E491A407C5448CBA831E4FAB807066</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFEB90038106FFF4FFB72D58FFA5FFF8</mods:identifier>
<mods:identifier type="Zenodo-Dep">144490</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182231473" LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9" httpUri="http://treatment.plazi.org/id/3B7A508A305255EC82AEE715DA8AE8F9" lastPageNumber="13" pageId="2" pageNumber="13">
<subSubSection pageId="2" pageNumber="13" type="nomenclature">
<paragraph pageId="2" pageNumber="13">
<taxonomicName LSID="3B7A508A-3052-55EC-82AE-E715DA8AE8F9" authority="Breuning, 1927" authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma (Calosoma) scabrosum subsp. roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="scabrosum" subGenus="Calosoma" subSpecies="roeschkei">Calosoma (Calosoma) scabrosum roeschkei Breuning, 1927</taxonomicName>
<figureCitation captionStart="Plate 1" captionStartId="F1" captionText="Plate 1. Type material (habitus in dorsal aspect, male). Scale bar equals 10 mm. 1 Calosoma scabrosum scabrosum (holotype) 2 Calosoma scabrosum roeschkei (holotype) 3 Calosoma imbricatum hottentottum (holotype of Ctenosta aethiops Jeannel, 1940) 4 Calosoma imbricatum hottentottum (holotype)." figureDoi="10.3897/zookeys.609.6822.plate1" httpUri="https://binary.pensoft.net/fig/98601" pageId="2" pageNumber="13">Plate 1</figureCitation>
<figureCitation captionStart="Plate 2" captionStartId="F2" captionText="Plate 2. Material studied (habitus in dorsal aspect, male). 1 Calosoma scabrosum scabrosum (Djibouti: Obock) 2 Calosoma scabrosum scabrosum (Oman: Dhofar, holotype of Calosoma jakli Haeckel, Farkac &amp; Sehnal, 2005) 3 Calosoma scabrosum roeschkei (Kenya: Voi env.) 4 Calosoma imbricatum hottentottum (Namibia: Okahandja). Scale bar equals 10 mm." figureDoi="10.3897/zookeys.609.6822.plate2" httpUri="https://binary.pensoft.net/fig/98602" pageId="2" pageNumber="13">: Fig. 2. Plate 2</figureCitation>
<figureCitation captionStart="Plate 3" captionStartId="F3" captionText="Plate 3. Aedeagi of Calosoma imbricatum hottentotum (Namibia) and Calosoma scabrosum roeschkei (Kenya) compared with aedeagus of &quot; Calosoma aethiops &quot; (holotype). 1 Calosoma imbricatum hottentottum (Namibia: Okahandja); a - aedeagus with expanded endophalus in right lateral view, b - the same in left lateral view, c - the same in anterior view 2 Calosoma imbricatum hottentottum (holotype of Ctenosta aethiops Jeannel, 1940) aedeagus in left lateral view 3 Calosoma scabrosum roeschkei (Kenya: Voi env.), a - aedeagus with expanded endophalus in right lateral view, b - the same in left lateral view, c - the same in anterior view." figureDoi="10.3897/zookeys.609.6822.plate3" httpUri="https://binary.pensoft.net/fig/98603" pageId="2" pageNumber="13">: Fig. 3. Plate 3</figureCitation>
<figureCitation captionStart="Plate 4" captionStartId="F4" captionText="Plate 4. Aedeagi of Calosoma senegalense species group (sensu Haeckel 2012: 30) and Calosoma imbricatum hottentotum (Namibia). a - aedeagus with expanded endophalus in right lateral view, b - same in left lateral view, c - same in anterior view 1 Calosoma scabrosum scabrosum (Oman: Dhofar) 2 Calosoma scabrosum roeschkei (southeastern Kenya: Voi env.) 3 Ctenosta strandi (northeastern Kenya: El Wak) 4 Calosoma planicolle (Namibia: Kavango) 5 Calosoma senegalense (Zimbabwe: Shamva) 6 Calosoma imbricatum hottentottum (Namibia: Okahandja)." figureDoi="10.3897/zookeys.609.6822.plate4" httpUri="https://binary.pensoft.net/fig/98604" pageId="2" pageNumber="13">: Fig. 3. Plate 4: Fig. 2.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="13" type="reference_group">
<paragraph pageId="2" pageNumber="13">
<taxonomicName authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma (Ctenosta) scabrosum subsp. roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="scabrosum" subGenus="Ctenosta" subSpecies="roeschkei">Calosoma (Ctenosta) scabrosum roeschkei</taxonomicName>
Breuning, 1927: 185 (type loc.
<normalizedToken originalValue="“Usambara”">&quot;Usambara&quot;</normalizedToken>
).
</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName authorityName="Jeannel" authorityYear="1940" class="Insecta" family="Carabidae" genus="Ctenosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenosta aethiops" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="aethiops">Ctenosta (s. str.) aethiops</taxonomicName>
(partim) Jeannel, 1940: 128 (loc.
<normalizedToken originalValue="“Diré-Daoua”">&quot;Dire-Daoua&quot;</normalizedToken>
);
<bibRefCitation author="Rougemont, GM de" journalOrPublisher="Revue de Zoologie africaine" pageId="14" pageNumber="25" pagination="241 - 274" refId="B24" refString="Rougemont, GM de, 1976. Contributions a la connaissance de la faune de l'Ethiopie. I. Les Calosomes (Coleoptera: Carabidae). Revue de Zoologie africaine 90 (2): 241 - 274" title="Contributions a la connaissance de la faune de l'Ethiopie. I. Les Calosomes (Coleoptera: Carabidae)." volume="90" year="1976">Rougemont 1976</bibRefCitation>
: 247.
<bibRefCitation author="Vigna Taglianti, A" journalOrPublisher="Memoire della Societa Entomologica Italiana" pageId="14" pageNumber="25" pagination="3 - 28" refId="B26" refString="Vigna Taglianti, A, Bruschi, S, 1986. Note sui Calosomini dell'Ethiopia. Memoire della Societa Entomologica Italiana 65: 3 - 28" title="Note sui Calosomini dell'Ethiopia." volume="65" year="1986">Vigna Taglianti and Bruschi 1986</bibRefCitation>
: 21.
</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName class="Insecta" family="Carabidae" genus="Ctenosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenosta orientale" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="orientale">Ctenosta (s. str.) orientale</taxonomicName>
(partim) Jeannel, 1940: 129 (loc.
<normalizedToken originalValue="“Érythrée">&quot;Erythree</normalizedToken>
: Tessenei&quot;).
</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName class="Insecta" family="Carabidae" genus="Ctenosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenosta scabrosum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="scabrosum">Ctenosta (s. str.) scabrosum</taxonomicName>
var.
<taxonomicName class="Insecta" family="Carabidae" genus="Ctenosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="roeschkei">roeschkei</taxonomicName>
Jeannel, 1940: 128.
</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName baseAuthorityName="Jeannel" baseAuthorityYear="1940" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma (Ctenosta) aethiops" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="aethiops" subGenus="Ctenosta">Calosoma (Ctenosta) aethiops</taxonomicName>
Culot, 1990: 9.
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="14" pageNumber="25" refId="B20" refString="Lorenz, W, 2005. Systematic List of extant Ground Beetles of the World (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing" title="Systematic List of extant Ground Beetles of the World (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition." year="2005">Lorenz 2005</bibRefCitation>
: 68.
</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName authorityName="Chaudoir" authorityYear="1843" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma scabrosum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="scabrosum">Calosoma (s. str.) scabrosum</taxonomicName>
<normalizedToken originalValue="Häckel">Haeckel</normalizedToken>
, 2012: 57.
<bibRefCitation author="Haeckel, M" journalOrPublisher="Czech University of Life Sciences Prague, Faculty of Environmental Sciences, Department of Ecology, Prague" pageId="14" pageNumber="25" refId="B13" refString="Haeckel, M, 2013. Kriticky nahled na soucasnou infragenerickou systematiku rodu krajnik Calosoma, Carabini, Carabidae) [A critical overview on the recent infrageneric taxonomy of the caterpillar hunter (Calosoma, Carabini, Carabidae)]. Diploma's Work. Czech University of Life Sciences Prague, Faculty of Environmental Sciences, Department of Ecology, Prague" title="Kriticky nahled na soucasnou infragenerickou systematiku rodu krajnik Calosoma, Carabini, Carabidae) [A critical overview on the recent infrageneric taxonomy of the caterpillar hunter (Calosoma, Carabini, Carabidae)]. Diploma's Work." year="2013">
<normalizedToken originalValue="Häckel">Haeckel</normalizedToken>
2013
</bibRefCitation>
: 31.
</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma (Ctenosta) roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="roeschkei" subGenus="Ctenosta">Calosoma (Ctenosta) roeschkei</taxonomicName>
Bruschi, 2013: 129.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="13" type="type material">
<paragraph pageId="2" pageNumber="13">Type material.</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="roeschkei">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma roeschkei</emphasis>
</taxonomicName>
Breuning, 1927. 1♂ labelled
<normalizedToken originalValue="“Usambara”">&quot;Usambara&quot;</normalizedToken>
(cNBCL).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="13" type="materials_examined">
<paragraph pageId="2" pageNumber="13">Other material studied.</paragraph>
<paragraph pageId="2" pageNumber="13">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Kenya" location="Witu" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis bold="true" pageId="2" pageNumber="13">
<collectingCountry name="Kenya">Kenya</collectingCountry>
.
</emphasis>
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
:
<locationDeviation location="Garsen">E of Garsen</locationDeviation>
, W of
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:04B74D5EC1703DCA81C5D096EB2188DE" country="Kenya" name="Witu">Witu</location>
(cSEH)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Kenya" location="Voi" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
: S of
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:3C2E856AD5B6E4D6122168EDED7D85FA" country="Kenya" name="Voi">Voi</location>
(cHAC)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Kenya" location="Voi" municipality="Sagala Hills" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Taita prov.">
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
:
<collectingRegion country="Kenya" name="Taita/Taveta">Taita prov.</collectingRegion>
<collectingMunicipality>Sagala Hills</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:2A894067D932345862AB34FF471FDE6E" country="Kenya" municipality="Sagala Hills" name="Voi" stateProvince="Taita prov.">Voi</location>
env. (cHAC)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Tsavo East" country="Kenya" latitude="-3.23" location="Voi" longLatPrecision="784" longitude="38.34" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
:
<collectorName>Tsavo East</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:8A37B9B53777D8B3D9BE12E1EBA7A05D" country="Kenya" latitude="-3.23" longLatPrecision="784" longitude="38.34" name="Voi">Voi</location>
Lodge,
<geoCoordinate degrees="3.23" direction="south" orientation="latitude" precision="555" value="-3.23">3.23S</geoCoordinate>
/
<geoCoordinate degrees="38.34" direction="east" orientation="longitude" precision="555" value="38.34">38.34E</geoCoordinate>
(cWRA)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Kenya" location="El Wak" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
: NE prov.
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:4D46DB2D88EEF5967EC9DA027E02BFD3" country="Kenya" name="El Wak">El Wak</location>
(cHAC)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Modo Gashi" country="Kenya" location="Wajir" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Wajir">
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
:
<collectorName>Modo Gashi</collectorName>
to
<collectingRegion country="Kenya" name="Wajir">Wajir</collectingRegion>
(cHAC)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Kenya" location="Coast province" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Garissa">
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
:
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:865E93960F34376CC0EFDFA15B883EE8" country="Kenya" name="Coast province" stateProvince="Garissa">Coast province</location>
,
<collectingRegion country="Kenya" name="Garissa">Garissa</collectingRegion>
,
<locationDeviation location="Bura">N of Bura</locationDeviation>
(cHAC, cSEH)
</materialsCitation>
,
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Kenya" location="Amboseli National Park" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
:
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:F6FA2E83CAF2199736573D63F191958A" country="Kenya" name="Amboseli National Park">Amboseli National Park</location>
(cSEH)
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Eastern, Sosoma" country="Kenya" location="202 km E of Thika" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2♀</specimenCount>
:
<collectorName>Eastern</collectorName>
729,
<collectorName>Sosoma</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:CDB0B1238B25AEC815FA433DF2191DDA" country="Kenya" name="202 km E of Thika">
<quantity metricMagnitude="5" metricUnit="m" metricValue="2.02" unit="km" value="202.0">202 km</quantity>
E of Thika
</location>
(cSEH)
</materialsCitation>
.
</paragraph>
<paragraph pageId="2" pageNumber="13">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Sudan" location="Sudan" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis bold="true" pageId="2" pageNumber="13">
<collectingCountry name="Sudan">Sudan</collectingCountry>
.
</emphasis>
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
: Vad Medani (cSEH)
</materialsCitation>
.
</paragraph>
<caption ID-Zenodo-Dep="974719" doi="10.3897/zookeys.609.6822.plate3" httpUri="https://binary.pensoft.net/fig/98603" pageId="2" pageNumber="13" start="Plate 3" startId="F3">
<paragraph pageId="2" pageNumber="13">
<emphasis bold="true" pageId="2" pageNumber="13">Plate 3.</emphasis>
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="Namibia" location="Voi" specimenCount="1" typeStatus="holotype">
Aedeagi of
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma imbricatum subsp. hottentotum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="imbricatum" subSpecies="hottentotum">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma imbricatum hottentotum</emphasis>
</taxonomicName>
(
<collectingCountry name="Namibia">Namibia</collectingCountry>
) and
<taxonomicName authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma scabrosum subsp. roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="scabrosum" subSpecies="roeschkei">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma scabrosum roeschkei</emphasis>
</taxonomicName>
(
<collectingCountry name="Kenya">Kenya</collectingCountry>
) compared with aedeagus of &quot;
<taxonomicName baseAuthorityName="Jeannel" baseAuthorityYear="1940" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma aethiops" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma aethiops</emphasis>
</taxonomicName>
&quot; (
<typeStatus>holotype</typeStatus>
).
<emphasis bold="true" pageId="2" pageNumber="13">1</emphasis>
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma imbricatum subsp. hottentottum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="imbricatum" subSpecies="hottentottum">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma imbricatum hottentottum</emphasis>
</taxonomicName>
(
<collectingCountry name="Namibia">Namibia</collectingCountry>
: Okahandja); a - aedeagus with expanded endophalus in right lateral view, b - the same in left lateral view, c - the same in anterior view
<emphasis bold="true" pageId="2" pageNumber="13">2</emphasis>
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma imbricatum subsp. hottentottum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="imbricatum" subSpecies="hottentottum">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma imbricatum hottentottum</emphasis>
</taxonomicName>
(
<typeStatus>holotype</typeStatus>
of
<taxonomicName authorityName="Jeannel" authorityYear="1940" class="Insecta" family="Carabidae" genus="Ctenosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenosta aethiops" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis italics="true" pageId="2" pageNumber="13">Ctenosta aethiops</emphasis>
</taxonomicName>
Jeannel, 1940) aedeagus in left lateral view
<emphasis bold="true" pageId="2" pageNumber="13">3</emphasis>
<taxonomicName authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma scabrosum subsp. roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="scabrosum" subSpecies="roeschkei">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma scabrosum roeschkei</emphasis>
</taxonomicName>
(
<collectingCountry name="Kenya">Kenya</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:3B7A508A305255EC82AEE715DA8AE8F9:B6762773D1A39BD95271AB6F3B7395D2" country="Namibia" name="Voi">Voi</location>
env.), a - aedeagus with expanded endophalus in right lateral view, b - the same in left lateral view, c - the same in anterior view
</materialsCitation>
.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="974721" doi="10.3897/zookeys.609.6822.plate4" httpUri="https://binary.pensoft.net/fig/98604" pageId="2" pageNumber="13" start="Plate 4" startId="F4">
<paragraph pageId="2" pageNumber="13">
<emphasis bold="true" pageId="2" pageNumber="13">Plate 4.</emphasis>
Aedeagi of
<taxonomicName authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma senegalense" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="senegalense">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma senegalense</emphasis>
</taxonomicName>
species group (sensu
<bibRefCitation author="Haeckel, M" journalOrPublisher="Folia Heyrovskyana (series A)" pageId="14" pageNumber="25" pagination="55 - 65" refId="B12" refString="Haeckel, M, 2012. Three new records of caterpillar hunters of the genus Calosoma Weber 1801 (Coleoptera: Carabidae: Carabini) from Afrotropical and Palearctic regions. Folia Heyrovskyana (series A) 20 (1-2): 55 - 65" title="Three new records of caterpillar hunters of the genus Calosoma Weber 1801 (Coleoptera: Carabidae: Carabini) from Afrotropical and Palearctic regions." volume="20" year="2012">
<normalizedToken originalValue="Häckel">Haeckel</normalizedToken>
2012
</bibRefCitation>
: 30) and
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma imbricatum subsp. hottentotum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="imbricatum" subSpecies="hottentotum">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma imbricatum hottentotum</emphasis>
</taxonomicName>
(
<collectingCountry name="Namibia">Namibia</collectingCountry>
). a - aedeagus with expanded endophalus in right lateral view, b - same in left lateral view, c - same in anterior view
<emphasis bold="true" pageId="2" pageNumber="13">1</emphasis>
<taxonomicName authority="scabrosum" authorityName="scabrosum Chaudoir" authorityYear="1869" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma scabrosum subsp. scabrosum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="scabrosum" subSpecies="scabrosum">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma scabrosum scabrosum</emphasis>
</taxonomicName>
(
<collectingCountry name="Oman">Oman</collectingCountry>
:
<collectingRegion country="Oman" name="Zufar">Dhofar</collectingRegion>
)
<emphasis bold="true" pageId="2" pageNumber="13">2</emphasis>
<taxonomicName authorityName="Breuning" authorityYear="1927" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma scabrosum subsp. roeschkei" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="scabrosum" subSpecies="roeschkei">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma scabrosum roeschkei</emphasis>
</taxonomicName>
(southeastern
<collectingCountry name="Kenya">Kenya</collectingCountry>
: Voi env.)
<emphasis bold="true" pageId="2" pageNumber="13">3</emphasis>
<taxonomicName baseAuthorityName="Breuning" baseAuthorityYear="1934" class="Insecta" family="Carabidae" genus="Ctenosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenosta strandi" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="strandi">
<emphasis italics="true" pageId="2" pageNumber="13">Ctenosta strandi</emphasis>
</taxonomicName>
(northeastern
<collectingCountry name="Kenya">Kenya</collectingCountry>
: El Wak)
<emphasis bold="true" pageId="2" pageNumber="13">4</emphasis>
<taxonomicName authorityName="Chaudoir" authorityYear="1869" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma planicolle" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="planicolle">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma planicolle</emphasis>
</taxonomicName>
(
<collectingCountry name="Namibia">Namibia</collectingCountry>
:
<collectingRegion country="Namibia" name="Kavango West">Kavango</collectingRegion>
)
<emphasis bold="true" pageId="2" pageNumber="13">5</emphasis>
<taxonomicName authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma senegalense" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="senegalense">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma senegalense</emphasis>
</taxonomicName>
(
<collectingCountry name="Zimbabwe">Zimbabwe</collectingCountry>
: Shamva)
<emphasis bold="true" pageId="2" pageNumber="13">6</emphasis>
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma imbricatum subsp. hottentottum" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="imbricatum" subSpecies="hottentottum">
<emphasis italics="true" pageId="2" pageNumber="13">Calosoma imbricatum hottentottum</emphasis>
</taxonomicName>
(
<collectingCountry name="Namibia">Namibia</collectingCountry>
: Okahandja).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="13" type="distribution">
<paragraph pageId="2" pageNumber="13">Geographic distribution.</paragraph>
<paragraph pageId="2" pageNumber="13">Chad, Kenya, Somalia, Sudan, Tanzania.</paragraph>
</subSubSection>
</treatment>
</document>
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