134 lines
13 KiB
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134 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.162.2216" ID-GBIF-Dataset="1d84f4b0-ac89-4089-9a4d-252c2abb8e12" ID-PMC="PMC3253662" ID-Pensoft-Pub="1313-2970-162-43" ID-PubMed="22303125" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-162-43" ModsDocOrigin="ZooKeys 162" ModsDocTitle="Revision of Tipula (Yamatotipula) stackelbergi Alexander (Diptera, Tipulidae), and a short discussion on subspecies among crane flies" checkinTime="1451249403002" checkinUser="pensoft" docAuthor="Salmela, Jukka" docDate="2012" docId="30EBBCA829A92D616339943AC061700D" docLanguage="en" docName="ZooKeys 162: 43-58" docOrigin="ZooKeys 162" docSource="http://dx.doi.org/10.3897/zookeys.162.2216" docTitle="Tipula (Yamatotipula) stackelbergi Alexander, stat. rev." docType="treatment" docVersion="3" lastPageNumber="46" masterDocId="FFE4FFFDFD15FB03FFBAAB0BB1690407" masterDocTitle="Revision of Tipula (Yamatotipula) stackelbergi Alexander (Diptera, Tipulidae), and a short discussion on subspecies among crane flies" masterLastPageNumber="58" masterPageNumber="43" pageNumber="45" updateTime="1668153136816" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Tipula (Yamatotipula) stackelbergi Alexander (Diptera, Tipulidae), and a short discussion on subspecies among crane flies</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salmela, Jukka</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>162</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>58</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.162.2216</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.162.2216</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-162-43</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152032936" LSID="urn:lsid:plazi:treatment:30EBBCA829A92D616339943AC061700D" httpUri="http://treatment.plazi.org/id/30EBBCA829A92D616339943AC061700D" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45">
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<subSubSection pageId="2" pageNumber="45" type="nomenclature">
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<paragraph pageId="2" pageNumber="45">
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<taxonomicName LSID="http://species-id.net/wiki/Tipula_stackelbergi" authority="Alexander" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) stackelbergi" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="stackelbergi" subGenus="Yamatotipula">Tipula (Yamatotipula) stackelbergi Alexander</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="45">stat. rev.</taxonomicNameLabel>
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Figs 123e6a, c, d
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="45" type="reference_group">
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<paragraph pageId="2" pageNumber="45">
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Tipula) stackelbergi" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="stackelbergi" subGenus="Tipula">Tipula (Tipula) stackelbergi</taxonomicName>
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<bibRefCitation author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="10" pageNumber="53" pagination="305 - 348" title="New or little-known Tipulidae from eastern Asia (Diptera). XVI." volume="52" year="1934">Alexander 1934</bibRefCitation>
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: 305.
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</paragraph>
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<paragraph pageId="2" pageNumber="45">
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) pruinosa" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="pruinosa" subGenus="Yamatotipula">Tipula (Yamatotipula) pruinosa</taxonomicName>
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stackelbergi
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<bibRefCitation pageId="2" pageNumber="45">Savchenko 1961</bibRefCitation>
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: 292.
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</paragraph>
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<paragraph pageId="2" pageNumber="45">
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) pruinosa subsp. stackelbergi" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="pruinosa" subGenus="Yamatotipula" subSpecies="stackelbergi">Tipula (Yamatotipula) pruinosa stackelbergi</taxonomicName>
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<bibRefCitation pageId="2" pageNumber="45">Oosterbroek and Theowald 1992</bibRefCitation>
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: 165.
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</paragraph>
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<paragraph pageId="2" pageNumber="45">
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) pruinosa subsp. stackelbergi" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="pruinosa" subGenus="Yamatotipula" subSpecies="stackelbergi">Tipula (Yamatotipula) pruinosa stackelbergi</taxonomicName>
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<bibRefCitation pageId="2" pageNumber="45">Oosterbroek 2011</bibRefCitation>
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: http://nlbif.eti.uva.nl/ccw/
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="45" type="material examined">
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<paragraph pageId="2" pageNumber="45">Material examined.</paragraph>
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<paragraph pageId="2" pageNumber="45">
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Holotype of
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula stackelbergi" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="stackelbergi">Tipula stackelbergi</taxonomicName>
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: male, pinned specimen (ZISP). "Tigrovaja, Suchan./ rn.Uss.kr. 16.VI/ Stackelberg. 927" (white label, partly hand written, in Cyrillic letters).
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<normalizedToken originalValue="“81”">"81"</normalizedToken>
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(white label, handwritten). "HOLOTYPE/ Tipula stackelbergi/ C.P. Alexander" (red label, partly handwritten) (Fig. 1a).
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</paragraph>
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<paragraph pageId="2" pageNumber="45">With except of the male hypopygium, the holotype specimen is in rather good condition (Figs 1b, c). All legs are detached from the specimen, but four legs are glued to the pin below the specimen. Tips of wings are broken. Right antenna is broken, only scape and pedicel are left; left antenna has seven flagellomeres. Tip of abdomen is broken; apparently hypopygium is mounted on a celluloid strip, which is attached on a pin. The surface of this strip is heavily cracked, and the structure of the hypopygium cannot be examined.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="45" type="other material">
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<paragraph pageId="2" pageNumber="45">Other material.</paragraph>
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<paragraph pageId="2" pageNumber="45">Russia, Vladivostok, Nekrutenko leg, 2.VI. 1957, 1 male, 1 female (ZISP).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45" type="redescription">
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<paragraph pageId="2" pageNumber="45">Redescription.</paragraph>
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<paragraph pageId="2" pageNumber="45">
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Male. Head yellowish brown, with grey pruinosity. Rostrum yellowish, nasus distinct, bearing numerous light hairs. Palpi brown. Scape yellowish, elongate, length 387-450
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 126-131
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(n=2). Pedicel yellowish, globular, length 147
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 139
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(n=1). Flagellomere 1 yellowish brown, length 486
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 91
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(n=1). Flagellomere 2 length 464
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 79
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(n=1). Flagellomeres bear erect short hairs, giving silvery appearance. Flagellomeres 2-7 elongate, brown, with dark verticils (Fig. 1c).
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</paragraph>
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<paragraph pageId="2" pageNumber="45">Prescutum with four brown stripes (Fig. 1e). Pronotum, prescutum, scutum, anepisternum, katepisternum and meron brownish, with grey pruinosity. Scutellum, anepimeron and laterotergite yellowish. Anterior part of mediotergite yellowish, more brownish in posterior part, having two weak longitudinal brown stripes. Coxa 1 brown. Anterior part of coxa 2 brown, posterior part yellow. Coxa 3 yellow. Femorae yellowish brown, darkening toward tarsi. Wings without markings, pterostigma brown (Fig. 1f). Wing length 13.8 mm (n=1). Halter yellowish.</paragraph>
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<paragraph pageId="2" pageNumber="45">Abdominal tergites yellowish brown, slightly darkening toward tip of abdomen. 9th tergite with two median projections, densely covered by dark bristles. Lateral corners of 9th tergite glabrous, pointed (Fig. 2a). 9th sternite with median incision, bearing two fleshy and hairy outgrowths in the margin of the incision. Outer gonostylus worm-like, apical half covered by light hairs (Fig. 2b). Inner gonostylus elongate (Figs 2b, c, 3e); beak rounded, with ten stout apical bristles and four subapical weaker bristles; central ridge with few weak bristles along its length; lower beak roundish, not angular. Posterior immovable apodeme of sperm pump almost straight (Fig. 2d). Distal end of compressor apodeme of sperm pump club-shaped, roundish (Fig. 2f). Aedeagal guide as in Fig. 2e.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45">
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Female. In general similar to male. Scutellum brown, abdominal tergites brown. Wing length 18.4 mm (n=1). Female terminalia as in Fig. 6a. Basal part of hypogynial
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<pageBreakToken pageId="3" pageNumber="46" start="start">valves</pageBreakToken>
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with dense black setae. Proximal ends of valves roundish, tapering toward bases (Fig. 6c). Genital fork of vaginal apodeme brown, rather narrow in its whole length (Fig. 6d). Dorsal view of vaginal apodeme as in Fig. 6d.
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</paragraph>
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<caption pageId="3" pageNumber="46">
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<paragraph pageId="3" pageNumber="46">
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Figure 1.
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) stackelbergi" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="stackelbergi" subGenus="Yamatotipula">Tipula (Yamatotipula) stackelbergi</taxonomicName>
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Alexander a Label of the holotype b Holotype male, habitus, lateral view. Yellowish celluloid board is attached below the specimen; most probably C.P. Alexander dissected hypopygium on this board. The surface of the board is heavily cracked, no details of the hypopygium are discernible c Thorax and head, holotype, lateral view d Right wing, holotype e Thorax and head, holotype, dorsal view f Male (Russia, Vladivostok), habitus, lateral view. Scale bars: b, f 2 mm; c & d 1 mm; e 0.5 mm.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="46">
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<paragraph pageId="3" pageNumber="46">
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Figure 2.
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) stackelbergi" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="stackelbergi" subGenus="Yamatotipula">Tipula (Yamatotipula) stackelbergi</taxonomicName>
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Alexander, male (Russia, Vladivostok) a 9th tergite, dorsal view b Outer and inner gonostylus, posterior view; abbreviations: out. gst.= outer gonostylus, in. gst. = inner gonostylus, lo. beak = lower beak c Inner gonostulys, anterior view d Sperm pump, lateral view; abbreviation: p.i.a. = posterior immovable apodeme e Aedeagal guide, dorsal view f Sperm pump, ventral view; abbreviation: c.a. = compressor apodeme. Scale bars: 0.2 mm.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="46">
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<paragraph pageId="3" pageNumber="46">
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Figure 3.
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula usuriensis" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="usuriensis">Tipula usuriensis</taxonomicName>
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Alexander (=syn. of
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<taxonomicName class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Yamatotipula) pruinosa" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="pruinosa" subGenus="Yamatotipula">Tipula (Yamatotipula) pruinosa</taxonomicName>
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Wiedemann), holotype male a Label of the holotype b Slide mounted wing c Habitus, lateral view d Sperm pump (lateral view) and aedeagal guide (dorso-lateral view). Scale bars: c 1mm; d 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |