310 lines
51 KiB
XML
310 lines
51 KiB
XML
<document id="5CCF48DF97F34D5DA67DD738397FA032" ID-CLB-Dataset="22130" ID-DOI="10.5852/ejt.2013.67" ID-GBIF-Dataset="2b629bb4-b2e2-43ba-aff9-2327452d30f6" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827652" ID-ZooBank="2358BCCE-7E9E-445D-B4B6-BBB56453055B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589477989497" checkinUser="carolina" docAuthor="Taverne, Louis & Capasso, Luigi" docDate="2013" docId="1F748A39FFD910335B2BD0ABFB431953" docLanguage="en" docName="ejt-2013-67.pdf.imf" docOrigin="European Journal of Taxonomy 67" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhinopycnodus gabriellae Taverne & Capasso 2013, gen. et sp. nov." docType="treatment" docUuid="D5399E6C-45DF-4D7D-BA77-545D6014BE22" docUuidSource="ZooBank" docVersion="8" lastPageNumber="11" masterDocId="E34DF241FFDB1039590BD65DFFFD1A50" masterDocTitle="Osteology and relationships of Rhinopycnodus gabriellae gen. et sp. nov. (Pycnodontiformes) from the marine Late Cretaceous of Lebanon" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1698839740370" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1CA04027C44BF94769D4A251CADBF7F3">Osteology and relationships of Rhinopycnodus gabriellae gen. et sp. nov. (Pycnodontiformes) from the marine Late Cretaceous of Lebanon</mods:title>
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<mods:roleTerm id="CA45FF35559D9FDEB7E794B6ECF5A762">Author</mods:roleTerm>
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<mods:namePart id="43491C8FB7542ED13696D635508F1AD1">Taverne, Louis</mods:namePart>
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<mods:affiliation id="78CC544018D7BA0286630BD977A65D82">Royal Belgian Institute of Natural Sciences, Department of Paleontology, Vautierstreet 29, B- 1000 Brussels, Belgium. E-mail: louis. taverne @ gmail. com & urn: lsid: zoobank. org: author: 0 CF 81641 - 1 DD 1 - 4 CBD- 9735 - F 1 FE 7 EB 0 BCF 5</mods:affiliation>
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<mods:namePart id="B475FC5A47476FCFAE19E3F5FF7E767E">Capasso, Luigi</mods:namePart>
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<mods:affiliation id="642EBB357331DA478DEA205681B12D17">Museo Universitario dell’Universitá “ G. d’Annunzio ” di Chieti-Pescara, Piazza Trento e Trieste 1, I- 661000 Chieti, Italy. E-mail: lcapasso @ unich. it & urn: lsid: zoobank. org: author: C 79 C 14 CF-C 1 D 7 - 48 E 3 - 9 BA 9 - CAD 8 AA 0909 F 7</mods:affiliation>
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<mods:title id="DA0A8786F5331166D3C3CBA465683F71">European Journal of Taxonomy</mods:title>
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<mods:date id="4E9D730E90554D00F0A399055AE795C3">2013</mods:date>
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<mods:number id="157669DD5A6C0ED2E05EA440198FA771">2013-12-04</mods:number>
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<mods:number id="52EB11C4105B45DF2691F97463D1119F">67</mods:number>
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<treatment id="1F748A39FFD910335B2BD0ABFB431953" ID-DOI="http://doi.org/10.5281/zenodo.3846649" ID-GBIF-Taxon="164152669" ID-Zenodo-Dep="3846649" LSID="urn:lsid:zoobank.org:act:D5399E6C-45DF-4D7D-BA77-545D6014BE22" httpUri="http://treatment.plazi.org/id/1F748A39FFD910335B2BD0ABFB431953" lastPageId="10" lastPageNumber="11" pageId="2" pageNumber="3">
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<subSubSection id="DFC768A4FFD9103B5B2BD0ABFBE91D41" box="[544,1044,1782,1809]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="97623B2FFFD9103B5B2BD0ABFBE91D41" blockId="2.[362,1226,1782,1880]" box="[544,1044,1782,1809]" pageId="2" pageNumber="3">
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<heading id="CC2A8C43FFD9103B5B2BD0ABFBE91D41" box="[544,1044,1782,1809]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="50DD40ACFFD9103B5B2BD0ABFCA01D41" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[544,861,1782,1809]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
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<emphasis id="A5A9E73DFFD9103B5B2BD0ABFCA01D41" bold="true" box="[544,861,1782,1809]" italics="true" pageId="2" pageNumber="3">Rhinopycnodus gabriellae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BE9A5A46FFD9103B5A6FD0AAFBE91D41" box="[868,1044,1783,1809]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="DFC768A4FFD9103B5BE2D147FB371D08" pageId="2" pageNumber="3" type="description">
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<paragraph id="97623B2FFFD9103B5BE2D147FCB61D64" blockId="2.[362,1226,1782,1880]" box="[745,843,1818,1844]" pageId="2" pageNumber="3">
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<figureCitation id="0FE627AAFFD9103B5BE2D147FCB61D64" box="[745,843,1818,1844]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId-0="3.[189,232,1076,1102]" captionStartId-1="3.[189,232,1998,2024]" captionStartId-2="5.[189,232,1964,1990]" captionStartId-3="6.[189,232,1964,1990]" captionStartId-4="7.[189,232,1963,1989]" captionStartId-5="8.[189,232,2000,2026]" captionStartId-6="9.[189,232,1964,1990]" captionStartId-7="10.[189,232,1964,1990]" captionTargetBox-0="[189,1399,333,1036]" captionTargetBox-1="[216,1377,1190,1910]" captionTargetBox-2="[378,1229,591,1922]" captionTargetBox-3="[197,1389,616,1910]" captionTargetBox-4="[495,1093,1443,1916]" captionTargetBox-5="[189,1399,1132,1960]" captionTargetBox-6="[215,1369,869,1923]" captionTargetBox-7="[229,1359,1049,1923]" captionTargetId-0="figure@3.[189,1399,333,1036]" captionTargetId-1="figure@3.[189,1399,1162,1941]" captionTargetId-2="figure@5.[378,1229,591,1922]" captionTargetId-3="figure@6.[189,1399,616,1921]" captionTargetId-4="figure@7.[491,1096,1443,1920]" captionTargetId-5="figure@8.[189,1399,1132,1960]" captionTargetId-6="figure@9.[189,1399,869,1923]" captionTargetId-7="figure@10.[189,1399,1036,1923]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="Fig. 1. Rhinopycnodus gabriellae gen. et sp. nov. Holotype CLC S-725." captionText-1="Fig. 2. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of holotype CLC S-725." captionText-2="Fig. 3. Rhinopycnodus gabriellae gen. et sp. nov. Head and shoulder girdle region of holotype CLC S-725." captionText-3="Fig. 4. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of the skull and shoulder girdle of holotype CLC S-725." captionText-4="Fig. 5. Rhinopycnodus gabriellae gen. et sp. nov. Pelvic girdle of holotype CLC S-725, surrounding pelvic and postcloacal scales and beginning of the anal fin." captionText-5="Fig. 6. Rhinopycnodus gabriellae gen. et sp. nov. Caudal region of holotype CLC S-725." captionText-6="Fig. 7. Rhinopycnodus gabriellae gen. et sp. nov. Caudal skeleton of holotype CLC S-725. The arrows point at the more external principal caudal rays." captionText-7="Fig. 8. Rhinopycnodus gabriellae gen. et sp. nov. A. The first twelve scales of the dorsal ridge. B. Three scales of the ventral keel." figureDoi-0="http://doi.org/10.5281/zenodo.3827654" figureDoi-1="http://doi.org/10.5281/zenodo.3827656" figureDoi-2="http://doi.org/10.5281/zenodo.3827658" figureDoi-3="http://doi.org/10.5281/zenodo.3827662" figureDoi-4="http://doi.org/10.5281/zenodo.3827664" figureDoi-5="http://doi.org/10.5281/zenodo.3827666" figureDoi-6="http://doi.org/10.5281/zenodo.3827668" figureDoi-7="http://doi.org/10.5281/zenodo.3827670" httpUri-0="https://zenodo.org/record/3827654/files/figure.png" httpUri-1="https://zenodo.org/record/3827656/files/figure.png" httpUri-2="https://zenodo.org/record/3827658/files/figure.png" httpUri-3="https://zenodo.org/record/3827662/files/figure.png" httpUri-4="https://zenodo.org/record/3827664/files/figure.png" httpUri-5="https://zenodo.org/record/3827666/files/figure.png" httpUri-6="https://zenodo.org/record/3827668/files/figure.png" httpUri-7="https://zenodo.org/record/3827670/files/figure.png" pageId="2" pageNumber="3">Figs 1-8</figureCitation>
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</paragraph>
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<paragraph id="97623B2FFFD9103B5861D160FB371D08" blockId="2.[362,1226,1782,1880]" box="[362,1226,1853,1880]" pageId="2" pageNumber="3">
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<uri id="E34C372DFFD9103B5861D160FB371D08" box="[362,1226,1853,1880]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid id="E37B01FAFFD9103B5B8DD160FB371D08" box="[646,1226,1853,1880]" pageId="2" pageNumber="3">D5399E6C-45DF-4D7D-BA77-545D6014BE22</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="DFC768A4FFD9103D59B6D122FB5F18F8" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="97623B2FFFD9103B59B6D122FECB1DC9" blockId="2.[189,310,1919,1945]" box="[189,310,1919,1945]" pageId="2" pageNumber="3">
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<heading id="CC2A8C43FFD9103B59B6D122FECB1DC9" bold="true" box="[189,310,1919,1945]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="A5A9E73DFFD9103B59B6D122FECB1DC9" bold="true" box="[189,310,1919,1945]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="97623B2FFFD9103D59B6D1F0FB5F18F8" blockId="2.[189,1399,1965,2027]" lastBlockId="4.[189,1399,267,681]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">Primitive pycnodontid characterized by a long and broad premaxilla bearing one short and very broad tooth. Elongated head with a long preorbital region. Dentary bearing 2 small incisiform teeth. No temporal fossa. Exoccipital-basioccipital region well visible behind the dermopterotic. Mouth gape obliquely oriented. Preopercle larger than the exposed region of the hyomandibula-dermohyomandibula. Maximum body depth: 67.6 % of the SL. Pectoral fin with 9 rays. Ventral fin with 3 rays. Dorsal fin with 49 pterygiophores. Origin of the dorsal fin behind the dorsal apex and at 76.2 % of the SL. Anal fin with 47 rays and 45 pterygiophores. Origin of the anal fin on the ventral apex and at 63.8 % of the SL. Neural and haemal arches almost completely surrounding the notochord. 32 vertebral elements (neural spines) before the epichordal series. Neural spines 1-8 autogenous. 11 pairs of ribs. Postcoelomic bone reaching the axial skeleton and the ventral margin. 15 haemal spines before the hypochordal series. 7 epichordal and 11 hypochordal elements in the caudal skeleton. Caudal fin with 30 principal rays. Body scales only in the abdominal region. Complete scales ventrally and scale bars dorsally. 19 dorsal ridge scales. First dorsal ridge scale small, triangular in shape and not sutured to the dermosupraoccipital. Some spiny scales in the dorsal ridge and in the ventral keel. 3 pelvic scales. 2 postcloacal scales.</paragraph>
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</subSubSection>
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<caption id="C3A26BA7FFD8103A59B6D269FBFA1E1F" ID-DOI="http://doi.org/10.5281/zenodo.3827654" ID-Zenodo-Dep="3827654" box="[189,1031,1076,1103]" httpUri="https://zenodo.org/record/3827654/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1076,1102]" targetBox="[189,1399,333,1036]" targetPageId="3">
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<paragraph id="97623B2FFFD8103A59B6D269FBFA1E1F" blockId="3.[189,1031,1076,1103]" box="[189,1031,1076,1103]" pageId="3" pageNumber="4">
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<emphasis id="A5A9E73DFFD8103A59B6D269FEF71E1E" bold="true" box="[189,266,1076,1102]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
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<taxonomicName id="50DD40ACFFD8103A5819D268FDBA1E1F" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[274,583,1077,1103]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
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<emphasis id="A5A9E73DFFD8103A5819D268FDBA1E1F" box="[274,583,1077,1103]" italics="true" pageId="3" pageNumber="4">Rhinopycnodus gabriellae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BE9A5A46FFD8103A5B45D268FD031E1F" box="[590,766,1077,1103]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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Holotype CLC S-725.
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</paragraph>
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</caption>
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<caption id="C3A26BA7FFD8103A59B6D193FB271DB9" ID-DOI="http://doi.org/10.5281/zenodo.3827656" ID-Zenodo-Dep="3827656" box="[189,1242,1998,2025]" httpUri="https://zenodo.org/record/3827656/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1998,2024]" targetBox="[216,1377,1190,1910]" targetPageId="3">
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<paragraph id="97623B2FFFD8103A59B6D193FB271DB9" blockId="3.[189,1242,1998,2025]" box="[189,1242,1998,2025]" pageId="3" pageNumber="4">
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<emphasis id="A5A9E73DFFD8103A59B6D193FEF71DB8" bold="true" box="[189,266,1998,2024]" pageId="3" pageNumber="4">Fig. 2.</emphasis>
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<taxonomicName id="50DD40ACFFD8103A5819D193FDBA1DB8" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[274,583,1998,2024]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
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<emphasis id="A5A9E73DFFD8103A5819D193FDBA1DB8" box="[274,583,1998,2024]" italics="true" pageId="3" pageNumber="4">Rhinopycnodus gabriellae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BE9A5A46FFD8103A5B45D192FD031DB9" box="[590,766,1999,2025]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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Reconstruction of holotype CLC S-725.
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</paragraph>
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</caption>
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<subSubSection id="DFC768A4FFDF103D59B6D48DFF04196C" pageId="4" pageNumber="5" type="etymology">
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<paragraph id="97623B2FFFDF103D59B6D48DFEB918BA" blockId="4.[189,324,720,746]" box="[189,324,720,746]" pageId="4" pageNumber="5">
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<heading id="CC2A8C43FFDF103D59B6D48DFEB918BA" bold="true" box="[189,324,720,746]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
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<emphasis id="A5A9E73DFFDF103D59B6D48DFEB918BA" bold="true" box="[189,324,720,746]" pageId="4" pageNumber="5">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="97623B2FFFDF103D59B6D4A3FF04196C" blockId="4.[189,1398,766,828]" pageId="4" pageNumber="5">The species name of the new Lebanese fossil fish is dedicated to Mrs. Gabriella di Tota, the co-author’s wife.</paragraph>
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</subSubSection>
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<subSubSection id="DFC768A4FFDF103D59B6D53EFD2D19FC" pageId="4" pageNumber="5" type="multiple">
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<paragraph id="97623B2FFFDF103D59B6D53EFE21192D" blockId="4.[189,476,867,893]" box="[189,476,867,893]" pageId="4" pageNumber="5">
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<heading id="CC2A8C43FFDF103D59B6D53EFE21192D" bold="true" box="[189,476,867,893]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
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<emphasis id="A5A9E73DFFDF103D59B6D53EFE21192D" bold="true" box="[189,476,867,893]" pageId="4" pageNumber="5">Formation and locality</emphasis>
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</heading>
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</paragraph>
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<paragraph id="97623B2FFFDF103D59B6D5CFFD2D19FC" blockId="4.[189,720,914,940]" box="[189,720,914,940]" pageId="4" pageNumber="5">
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Marine Upper Cenomanian, Haqel,
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<collectingCountry id="EFCA7BBFFFDF103D5B6AD5CFFD3119FC" box="[609,716,914,940]" name="Lebanon" pageId="4" pageNumber="5">Lebanon</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="DFC768A4FFDF103D59B6D58EFE821E6F" pageId="4" pageNumber="5" type="materials_examined">
|
||
<materialsCitation id="27B53172FFDF103D59B6D58EFE821E6F" ID-GBIF-Occurrence="2619829301" collectionCode="CLC" country="Lebanon" location="Total" pageId="4" pageNumber="5" specimenCode="CLC S-725" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="97623B2FFFDF103D59B6D58EFDC219BD" blockId="4.[189,575,979,1005]" box="[189,575,979,1005]" pageId="4" pageNumber="5">
|
||
<heading id="CC2A8C43FFDF103D59B6D58EFDC219BD" bold="true" box="[189,575,979,1005]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="A5A9E73DFFDF103D59B6D58EFDC219BD" bold="true" box="[189,575,979,1005]" pageId="4" pageNumber="5">
|
||
<typeStatus id="4866858DFFDF103D59B6D58EFED219BD" box="[189,303,979,1005]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
and unique
|
||
<specimenCount id="81DBF0A6FFDF103D58C0D58EFDC219BD" box="[459,575,979,1005]" pageId="4" pageNumber="5" type="generic">specimen</specimenCount>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDF103D59B6D25FFE821E6F" blockId="4.[189,1399,1025,1087]" pageId="4" pageNumber="5">
|
||
Sample
|
||
<specimenCode id="C77B9354FFDF103D5816D25FFE5A1E4C" box="[285,423,1025,1052]" pageId="4" pageNumber="5">CLC S-725</specimenCode>
|
||
, a complete specimen seen by its right side (
|
||
<figureCitation id="0FE627AAFFDF103D5ABED25FFBFD1E4C" box="[949,1024,1026,1052]" captionStart="Fig" captionStartId="3.[189,232,1076,1102]" captionTargetBox="[189,1399,333,1036]" captionTargetId="figure@3.[189,1399,333,1036]" captionTargetPageId="3" captionText="Fig. 1. Rhinopycnodus gabriellae gen. et sp. nov. Holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827654" httpUri="https://zenodo.org/record/3827654/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="0FE627AAFFDF103D5D04D25FFBE21E4C" box="[1039,1055,1026,1052]" captionStart="Fig" captionStartId="3.[189,232,1998,2024]" captionTargetBox="[216,1377,1190,1910]" captionTargetId="figure@3.[189,1399,1162,1941]" captionTargetPageId="3" captionText="Fig. 2. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827656" httpUri="https://zenodo.org/record/3827656/files/figure.png" pageId="4" pageNumber="5">2</figureCitation>
|
||
) from Haqel,
|
||
<collectingCountry id="EFCA7BBFFFDF103D5DCED25FFACD1E4C" box="[1221,1328,1026,1052]" name="Lebanon" pageId="4" pageNumber="5">Lebanon</collectingCountry>
|
||
.
|
||
<location id="92026DF4FFDF103D5C30D25FFA8A1E4C" LSID="urn:lsid:plazi:treatment:1F748A39FFD910335B2BD0ABFB431953:92026DF4FFDF103D5C30D25FFA8A1E4C" box="[1339,1399,1026,1052]" country="Lebanon" name="Total" pageId="4" pageNumber="5">Total</location>
|
||
length:
|
||
<quantity id="502596CAFFDF103D581DD278FE871E6F" box="[278,378,1061,1087]" metricMagnitude="-1" metricUnit="m" metricValue="2.23" pageId="4" pageNumber="5" unit="mm" value="223.0">223 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="DFC768A4FFDF103359B6D23AFB431953" lastPageId="10" lastPageNumber="11" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="97623B2FFFDF103D59B6D23AFDD71ED1" blockId="4.[189,554,1126,1153]" box="[189,554,1126,1153]" pageId="4" pageNumber="5">
|
||
<heading id="CC2A8C43FFDF103D59B6D23AFDD71ED1" bold="true" box="[189,554,1126,1153]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="A5A9E73DFFDF103D59B6D23AFDD71ED1" bold="true" box="[189,554,1126,1153]" pageId="4" pageNumber="5">
|
||
<typeStatus id="4866858DFFDF103D59B6D23AFED21ED1" box="[189,303,1127,1153]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
morphometric data
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDF103D59B6D2C8FB381EFF" blockId="4.[189,1221,1173,1199]" box="[189,1221,1173,1199]" pageId="4" pageNumber="5">
|
||
The morphometric data are given in % of the standard length (
|
||
<quantity id="502596CAFFDF103D5A93D2C8FC001EFF" box="[920,1021,1173,1199]" metricMagnitude="-1" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="5" unit="mm" value="183.0">183 mm</quantity>
|
||
) of the
|
||
<typeStatus id="4866858DFFDF103D5D5DD2C8FB3C1EFF" box="[1110,1217,1173,1199]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDF103D59B6D28BFA8B1FB6" blockId="4.[189,1398,1237,1511]" pageId="4" pageNumber="5">Length of the head (opercle included) ……………………………………………………………45.2 % Depth of the head (in the occipital region) ………………………………………………………36.7 % Maximum depth of the body (at the anal fin origin level) ………………………………………67.6 % Prepelvic length …………………………………………………………………………………52.4 % Predorsal length …………………………………………………………………………………76.2 % Basal length of the dorsal fin ……………………………………………………………………39.0 % Preanal length ……………………………………………………………………………………63.8 % Basal length of the anal fin ………………………………………………………………………40.0 %</paragraph>
|
||
<paragraph id="97623B2FFFDF103D59B6D053FEC41C78" blockId="4.[189,313,1550,1576]" box="[189,313,1550,1576]" pageId="4" pageNumber="5">
|
||
<heading id="CC2A8C43FFDF103D59B6D053FEC41C78" bold="true" box="[189,313,1550,1576]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="A5A9E73DFFDF103D59B6D053FEC41C78" bold="true" box="[189,313,1550,1576]" pageId="4" pageNumber="5">Osteology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDF103D59E1D060FE001C07" blockId="4.[189,1399,1597,1694]" box="[234,509,1597,1623]" pageId="4" pageNumber="5">
|
||
<emphasis id="A5A9E73DFFDF103D59E1D060FE861C07" bold="true" box="[234,379,1597,1623]" pageId="4" pageNumber="5">1. The skull</emphasis>
|
||
(
|
||
<figureCitation id="0FE627AAFFDF103D5880D060FE2A1C07" box="[395,471,1597,1623]" captionStart="Fig" captionStartId="5.[189,232,1964,1990]" captionTargetBox="[378,1229,591,1922]" captionTargetId="figure@5.[378,1229,591,1922]" captionTargetPageId="5" captionText="Fig. 3. Rhinopycnodus gabriellae gen. et sp. nov. Head and shoulder girdle region of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827658" httpUri="https://zenodo.org/record/3827658/files/figure.png" pageId="4" pageNumber="5">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="0FE627AAFFDF103D58EFD060FE091C07" box="[484,500,1597,1623]" captionStart="Fig" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[197,1389,616,1910]" captionTargetId="figure@6.[189,1399,616,1921]" captionTargetPageId="6" captionText="Fig. 4. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of the skull and shoulder girdle of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827662" httpUri="https://zenodo.org/record/3827662/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDF103D59B6D03DFA8A1CCD" blockId="4.[189,1399,1597,1694]" pageId="4" pageNumber="5">The head is high, with the preorbital region much longer than the orbital-postorbital region. The dermal bones of the skull are slightly ornamented with small granulations. The mouth gape is inclined ventrally.</paragraph>
|
||
<paragraph id="97623B2FFFDF103C59B6D099FE1C1B3C" blockId="4.[189,1399,1732,2005]" lastBlockId="5.[189,1399,267,364]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">The mesethmoid is broad, very long and its upper margin is covered by a pair of long and very narrow prefrontals. The vomer is seen in profile and only six rounded molariform teeth ranged in one rank are visible. The frontal is rather short and not very broad. The posterior margin of the bone is a little enlarged and meets the dermosupraocciptal, the parietal and the small autosphenotic, but not the dermopterotic. Posteriorly, the dermosupraoccipital ends in a short pointed tip. The parietal bears a large posterior brush-like process. There is no temporal fenestra. The dermopterotic is not deepened and is located at the level of the upper border of the orbit. The opisthotic, intercalar, basioccipital and exoccipital are visible behind the dermopterotic and the hyomandibula. A small orbitosphenoid is pressed against the posterior border of the mesethmoid. The parasphenoid is very long and its trabecular region is obliquely oriented. Posteriorly, the parasphenoid reaches the level of the basioccipital. The sensory canals on the braincase are not visible.</paragraph>
|
||
<paragraph id="97623B2FFFDE103C59B6D7CEFAFD1B80" blockId="5.[189,1398,403,465]" pageId="5" pageNumber="6">The quadratic arch contains a well developed quadrate, a large metapterygoid, a large entopterygoid and a small ectopterygoid. The quadrate and the symplectic are both articulated on the lower jaw.</paragraph>
|
||
<caption id="C3A26BA7FFDE103C59B6D1F1FEF61DBA" ID-DOI="http://doi.org/10.5281/zenodo.3827658" ID-Zenodo-Dep="3827658" httpUri="https://zenodo.org/record/3827658/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1964,1990]" targetBox="[378,1229,591,1922]" targetPageId="5">
|
||
<paragraph id="97623B2FFFDE103C59B6D1F1FEF61DBA" blockId="5.[189,1398,1964,2026]" pageId="5" pageNumber="6">
|
||
<emphasis id="A5A9E73DFFDE103C59B6D1F1FEF01D96" bold="true" box="[189,269,1964,1990]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
|
||
<taxonomicName id="50DD40ACFFDE103C581CD1F1FDB21D96" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[279,591,1964,1990]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
|
||
<emphasis id="A5A9E73DFFDE103C581CD1F1FDB21D96" box="[279,591,1964,1990]" italics="true" pageId="5" pageNumber="6">Rhinopycnodus gabriellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BE9A5A46FFDE103C5B52D1F0FCEC1D97" box="[601,785,1965,1991]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Head and shoulder girdle region of holotype CLC S-725.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97623B2FFFDD103F59B6D756FAD91BA8" blockId="6.[189,1399,267,505]" pageId="6" pageNumber="7">The premaxilla is long and very broad. It bears only one short but very broad tooth. The maxilla is small, deeper than long, reniform, toothless and pressed against the premaxilla by its upper margin. When seen in profile, the upper jaw is hog snout-like because of the broadening of the premaxilla and of its tooth. The lower jaw is small and triangular in shape. The dentary is reduced to its ventral branch and it bears two small incisiform teeth. The angular covers a great part of the external face of the mandible. The small articular and the dentary are articulated together. The coronoid process of the prearticular is well marked. Three deep molariform teeth fallen from the prearticular are visible just behind the jaw.</paragraph>
|
||
<caption id="C3A26BA7FFDD103F59B6D1F1FE441DBA" ID-DOI="http://doi.org/10.5281/zenodo.3827662" ID-Zenodo-Dep="3827662" httpUri="https://zenodo.org/record/3827662/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1964,1990]" targetBox="[197,1389,616,1910]" targetPageId="6">
|
||
<paragraph id="97623B2FFFDD103F59B6D1F1FE441DBA" blockId="6.[189,1398,1964,2026]" pageId="6" pageNumber="7">
|
||
<emphasis id="A5A9E73DFFDD103F59B6D1F1FEF01D96" bold="true" box="[189,269,1964,1990]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
|
||
<taxonomicName id="50DD40ACFFDD103F581CD1F1FDAD1D96" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[279,592,1964,1990]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
|
||
<emphasis id="A5A9E73DFFDD103F581CD1F1FDAD1D96" box="[279,592,1964,1990]" italics="true" pageId="6" pageNumber="7">Rhinopycnodus gabriellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BE9A5A46FFDD103F5B52D1F0FCEF1D97" box="[601,786,1965,1991]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Reconstruction of the skull and shoulder girdle of holotype CLC S-725.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97623B2FFFDC103E59B6D756FC961B18" blockId="7.[189,1399,267,328]" pageId="7" pageNumber="8">The last infraorbital is long, well ossified and forms the posterior and ventral margins of the orbit. No other infraorbital is preserved. The dermosphenotic is lost.</paragraph>
|
||
<paragraph id="97623B2FFFDC103E59B6D72DFDCC1BA3" blockId="7.[189,1399,368,500]" pageId="7" pageNumber="8">The preopercle is the largest bone of the skull, all together deep and broad. The hyomandibula and dermohyomandibula are fused and their exposed part is important but, however, much smaller than the preopercle. The opercle is well developed, with a pointed ventral tip and a broader upper part. No branchiostegal ray is preserved.</paragraph>
|
||
<paragraph id="97623B2FFFDC103E59B6D446FAF31808" blockId="7.[189,1398,539,601]" pageId="7" pageNumber="8">The anterior ceratohyal is the only preserved part of the hyoid bar. A few hook-like branchial teeth and some branchial filaments are visible under some broken regions of the opercle and preopercle.</paragraph>
|
||
<paragraph id="97623B2FFFDC103E59E1D4DDFDED18CA" blockId="7.[189,1399,639,807]" box="[234,528,639,666]" pageId="7" pageNumber="8">
|
||
<emphasis id="A5A9E73DFFDC103E59E1D4DDFE6918C9" bold="true" box="[234,404,639,666]" pageId="7" pageNumber="8">2. The girdles</emphasis>
|
||
(
|
||
<figureCitation id="0FE627AAFFDC103E58AFD4DDFDFB18CA" box="[420,518,639,666]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="5.[189,232,1964,1990]" captionStartId-1="6.[189,232,1964,1990]" captionStartId-2="7.[189,232,1963,1989]" captionTargetBox-0="[378,1229,591,1922]" captionTargetBox-1="[197,1389,616,1910]" captionTargetBox-2="[495,1093,1443,1916]" captionTargetId-0="figure@5.[378,1229,591,1922]" captionTargetId-1="figure@6.[189,1399,616,1921]" captionTargetId-2="figure@7.[491,1096,1443,1920]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Fig. 3. Rhinopycnodus gabriellae gen. et sp. nov. Head and shoulder girdle region of holotype CLC S-725." captionText-1="Fig. 4. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of the skull and shoulder girdle of holotype CLC S-725." captionText-2="Fig. 5. Rhinopycnodus gabriellae gen. et sp. nov. Pelvic girdle of holotype CLC S-725, surrounding pelvic and postcloacal scales and beginning of the anal fin." figureDoi-0="http://doi.org/10.5281/zenodo.3827658" figureDoi-1="http://doi.org/10.5281/zenodo.3827662" figureDoi-2="http://doi.org/10.5281/zenodo.3827664" httpUri-0="https://zenodo.org/record/3827658/files/figure.png" httpUri-1="https://zenodo.org/record/3827662/files/figure.png" httpUri-2="https://zenodo.org/record/3827664/files/figure.png" pageId="7" pageNumber="8">Figs 3-5</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDC103E59B6D4FEFE3B1977" blockId="7.[189,1399,639,807]" pageId="7" pageNumber="8">
|
||
Only the ventral part of the posttemporal is preserved. The hypercleithrum (= supracleithrum) is deep, rather broad and not splint-like as usual in
|
||
<taxonomicName id="50DD40ACFFDC103E5BA1D49BFC7818B0" authorityName="Berg, 1937 sensu Nursall" authorityYear="2010" box="[682,901,710,736]" class="Actinopterygii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="order">Pycnodontiformes</taxonomicName>
|
||
. The cleithrum is a large bone with a broad palaform ventral branch and a sinus in its posterior margin to house the pectoral fin, which is short and contains a least 9 rays.
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDC103E59B6D513FAF319DC" blockId="7.[189,1399,846,908]" pageId="7" pageNumber="8">The two pelvic bones are vertically oriented. The ventral fins are rather long. Each of them contains 3 rays. The origin of the ventral fins is located a little before the midpoint of the ventral contour.</paragraph>
|
||
<paragraph id="97623B2FFFDC103E59E1D5EFFDB4199D" blockId="7.[189,1399,946,1395]" box="[234,585,946,973]" pageId="7" pageNumber="8">
|
||
<emphasis id="A5A9E73DFFDC103E59E1D5EFFE17199D" bold="true" box="[234,490,946,973]" pageId="7" pageNumber="8">3. The axial skeleton</emphasis>
|
||
(
|
||
<figureCitation id="0FE627AAFFDC103E58F1D5EFFDBC199D" box="[506,577,946,973]" captionStart="Fig" captionStartId="3.[189,232,1998,2024]" captionTargetBox="[216,1377,1190,1910]" captionTargetId="figure@3.[189,1399,1162,1941]" captionTargetPageId="3" captionText="Fig. 2. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827656" httpUri="https://zenodo.org/record/3827656/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFDC103159B6D58BFCA81B3C" blockId="7.[189,1399,946,1395]" lastBlockId="8.[189,1399,267,364]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">Starting from the caudal region, the vertebral axis progressively elevates to reach the level of the orbit anteriorly. The vertebrae are constituted by only the dorsal and ventral arcocentra. No chordacentrum or autocentrum is visible. The neural and haemal arches surround the notochord almost completely. There are 32 neural spines before the epichordal series and thus 32 vertebral segments before the elements involved in the caudal fin support. The first 17 vertebral segments are abdominal and the following 15 caudal. The anteriormost 8 neural spines are autogenous. The first seven lean on the occipital region of the braincase and the eighth spine is located just above the first ossified but small basidorsal. The basiventrals are strongly reduced in the abdominal region but well developed in the caudal region. There are 11 pairs of long alate ribs and 15 haemal spines before the hypochordal series. The two last pairs of ribs are noticeably shorter than the nine preceding ribs. The first three haemal spines are pressed together and against the postcoelomic bone. The first haemal spine is incomplete and does not reach the axial skeleton. The neural and haemal spines bear an anterior sagittal flange, except for the first five neural and the first three haemal spines. A well developed postzygaphophysis links each neural and haemal arch with the following one. The postcoelomic bone is a long and robust bone dorsally reaching the axial skeleton and ventrally the lower margin of the abdomen.</paragraph>
|
||
<caption id="C3A26BA7FFDC103E59B6D1F6FC921DB9" ID-DOI="http://doi.org/10.5281/zenodo.3827664" ID-Zenodo-Dep="3827664" httpUri="https://zenodo.org/record/3827664/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1963,1989]" targetBox="[495,1093,1443,1916]" targetPageId="7">
|
||
<paragraph id="97623B2FFFDC103E59B6D1F6FC921DB9" blockId="7.[189,1400,1963,2025]" pageId="7" pageNumber="8">
|
||
<emphasis id="A5A9E73DFFDC103E59B6D1F6FEF01D96" bold="true" box="[189,269,1963,1990]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
|
||
<taxonomicName id="50DD40ACFFDC103E581CD1F1FDB21D96" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[279,591,1964,1990]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
|
||
<emphasis id="A5A9E73DFFDC103E581CD1F1FDB21D96" box="[279,591,1964,1990]" italics="true" pageId="7" pageNumber="8">Rhinopycnodus gabriellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BE9A5A46FFDC103E5B52D1F1FCEC1D96" box="[601,785,1964,1990]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Pelvic girdle of holotype CLC S-725, surrounding pelvic and postcloacal scales and beginning of the anal fin.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97623B2FFFD3103159E1D7CEFD691BFD" blockId="8.[189,1399,403,570]" box="[234,660,403,429]" pageId="8" pageNumber="9">
|
||
<emphasis id="A5A9E73DFFD3103159E1D7CEFDC81BFD" bold="true" box="[234,565,403,429]" pageId="8" pageNumber="9">4. The dorsal and anal fins</emphasis>
|
||
(
|
||
<figureCitation id="0FE627AAFFD310315B4ED7CEFD711BFD" box="[581,652,403,429]" captionStart="Fig" captionStartId="3.[189,232,1998,2024]" captionTargetBox="[216,1377,1190,1910]" captionTargetId="figure@3.[189,1399,1162,1941]" captionTargetPageId="3" captionText="Fig. 2. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827656" httpUri="https://zenodo.org/record/3827656/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFD3103159B6D7EBFE62186A" blockId="8.[189,1399,403,570]" pageId="8" pageNumber="9">The dorsal fin begins a little behind the dorsal apex. The fin is supported by 49 long and strong pterygiophores. The first seventeen of them have lost the corresponding rays. The last thirty two pterygiophores bear short segmented and branched rays. The first two pterygiophores are broader than the following ones.</paragraph>
|
||
<paragraph id="97623B2FFFD3103159B6D43CFA8A1959" blockId="8.[189,1399,609,777]" pageId="8" pageNumber="9">
|
||
The anal fin is strip-like in shape (
|
||
<typeStatus id="4866858DFFD310315B50D43FFD72182C" box="[603,655,610,636]" pageId="8" pageNumber="9">type</typeStatus>
|
||
A2 of
|
||
<bibRefCitation id="F34C46DEFFD310315BEDD43CFBCB182C" author="Poyato-Ariza F. J. & Wenz S." box="[742,1078,609,636]" pageId="8" pageNumber="9" pagination="139 - 248" refId="ref5156" refString="Poyato-Ariza F. J. & Wenz S. 2002. A new insight into pycnodontiform fishes. Geodiversitas 24 (1): 139 - 248." type="journal article" year="2002">Poyato-Ariza & Wenz 2002</bibRefCitation>
|
||
: fig. 34). The origin of the fin is located at the ventral apex. There are 45 strong pterygiophores bearing 47 rays. The first five pterygiophores abut against the postcoelomic bone and are progressively lengthened from the first to the fifth. The first three rays are reduced to short spines. The other rays are segmented and branched. The very short first pterygiophore supports two little spiny rays and the last pterygiophore two soft rays.
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFD3103159E1D56DFD7C191A" blockId="8.[189,1399,816,1053]" box="[234,641,816,842]" pageId="8" pageNumber="9">
|
||
<emphasis id="A5A9E73DFFD3103159E1D56DFDFC191A" bold="true" box="[234,513,816,842]" pageId="8" pageNumber="9">5. The caudal skeleton</emphasis>
|
||
(
|
||
<figureCitation id="0FE627AAFFD310315B1BD56DFDA1191A" box="[528,604,816,842]" captionStart="Fig" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1399,1132,1960]" captionTargetId="figure@8.[189,1399,1132,1960]" captionTargetPageId="8" captionText="Fig. 6. Rhinopycnodus gabriellae gen. et sp. nov. Caudal region of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827666" httpUri="https://zenodo.org/record/3827666/files/figure.png" pageId="8" pageNumber="9">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="0FE627AAFFD310315B62D56DFD85191A" box="[617,632,816,842]" captionStart="Fig" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[215,1369,869,1923]" captionTargetId="figure@9.[189,1399,869,1923]" captionTargetPageId="9" captionText="Fig. 7. Rhinopycnodus gabriellae gen. et sp. nov. Caudal skeleton of holotype CLC S-725. The arrows point at the more external principal caudal rays." figureDoi="http://doi.org/10.5281/zenodo.3827668" httpUri="https://zenodo.org/record/3827668/files/figure.png" pageId="8" pageNumber="9">7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFD3103159B6D50EFB791E4D" blockId="8.[189,1399,816,1053]" pageId="8" pageNumber="9">There is no caudal peduncle because the dorsal and anal fins reach the caudal fin.The caudal endoskeleton contains 1 urodermal, 7 epichordal and 11 hypochordal elements. The first epichordal neural arch bears a long and narrow neural spine but the length of the neural spines progressively decreases from the first to the seventh epichordal element, which has only a very short neural spine. Some elements in the hypochordal series are broadened but there is no real hypertrophy. This broadening is more important on the eighth and the tenth hypochordal elements than on the other parts of the series.</paragraph>
|
||
<caption id="C3A26BA7FFD3103159B6D18DFB301DBB" ID-DOI="http://doi.org/10.5281/zenodo.3827666" ID-Zenodo-Dep="3827666" box="[189,1229,2000,2027]" httpUri="https://zenodo.org/record/3827666/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,2000,2026]" targetBox="[189,1399,1132,1960]" targetPageId="8">
|
||
<paragraph id="97623B2FFFD3103159B6D18DFB301DBB" blockId="8.[189,1229,2000,2027]" box="[189,1229,2000,2027]" pageId="8" pageNumber="9">
|
||
<emphasis id="A5A9E73DFFD3103159B6D18DFEF71DBA" bold="true" box="[189,266,2000,2026]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
|
||
<taxonomicName id="50DD40ACFFD310315819D18DFDBA1DBA" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[274,583,2000,2026]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
|
||
<emphasis id="A5A9E73DFFD310315819D18DFDBA1DBA" box="[274,583,2000,2026]" italics="true" pageId="8" pageNumber="9">Rhinopycnodus gabriellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BE9A5A46FFD310315B45D18CFD031DBB" box="[590,766,2001,2027]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Caudal region of holotype CLC S-725.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97623B2FFFD2103059B6D756FE8A1B3C" blockId="9.[189,1399,267,364]" pageId="9" pageNumber="10">
|
||
The contour of the caudal fin is double emarginated (
|
||
<bibRefCitation id="F34C46DEFFD210305A48D756FB6A1B75" author="Poyato-Ariza F. J. & Wenz S." box="[835,1175,267,294]" pageId="9" pageNumber="10" pagination="139 - 248" refId="ref5156" refString="Poyato-Ariza F. J. & Wenz S. 2002. A new insight into pycnodontiform fishes. Geodiversitas 24 (1): 139 - 248." type="journal article" year="2002">Poyato-Ariza & Wenz 2002</bibRefCitation>
|
||
: fig. 36 E) but the median convex part of the fin is greatly enlarged. There are 30 principal rays, 3 dorsal and 4 ventral procurrent rays.
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFD2103059E1D7CEFDA61BFD" blockId="9.[189,1215,403,465]" box="[234,603,403,430]" pageId="9" pageNumber="10">
|
||
<emphasis id="A5A9E73DFFD2103059E1D7CEFE5D1BFD" bold="true" box="[234,416,403,429]" pageId="9" pageNumber="10">6. Squamation</emphasis>
|
||
(
|
||
<figureCitation id="0FE627AAFFD2103058A4D7CEFE061BFD" box="[431,507,403,429]" captionStart="Fig" captionStartId="3.[189,232,1998,2024]" captionTargetBox="[216,1377,1190,1910]" captionTargetId="figure@3.[189,1399,1162,1941]" captionTargetPageId="3" captionText="Fig. 2. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827656" httpUri="https://zenodo.org/record/3827656/files/figure.png" pageId="9" pageNumber="10">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="0FE627AAFFD210305B03D7CEFDE41BFD" box="[520,537,403,429]" captionStart="Fig" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[197,1389,616,1910]" captionTargetId="figure@6.[189,1399,616,1921]" captionTargetPageId="6" captionText="Fig. 4. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of the skull and shoulder girdle of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827662" httpUri="https://zenodo.org/record/3827662/files/figure.png" pageId="9" pageNumber="10">4</figureCitation>
|
||
,
|
||
<figureCitation id="0FE627AAFFD210305B2ED7CEFDC81BFE" box="[549,565,403,430]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[495,1093,1443,1916]" captionTargetId="figure@7.[491,1096,1443,1920]" captionTargetPageId="7" captionText="Fig. 5. Rhinopycnodus gabriellae gen. et sp. nov. Pelvic girdle of holotype CLC S-725, surrounding pelvic and postcloacal scales and beginning of the anal fin." figureDoi="http://doi.org/10.5281/zenodo.3827664" httpUri="https://zenodo.org/record/3827664/files/figure.png" pageId="9" pageNumber="10">5</figureCitation>
|
||
,
|
||
<figureCitation id="0FE627AAFFD210305B49D7CEFDAC1BFD" box="[578,593,403,429]" captionStart="Fig" captionStartId="10.[189,232,1964,1990]" captionTargetBox="[229,1359,1049,1923]" captionTargetId="figure@10.[189,1399,1036,1923]" captionTargetPageId="10" captionText="Fig. 8. Rhinopycnodus gabriellae gen. et sp. nov. A. The first twelve scales of the dorsal ridge. B. Three scales of the ventral keel." figureDoi="http://doi.org/10.5281/zenodo.3827670" httpUri="https://zenodo.org/record/3827670/files/figure.png" pageId="9" pageNumber="10">8</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFD2103059B6D7EBFB421B80" blockId="9.[189,1215,403,465]" box="[189,1215,438,465]" pageId="9" pageNumber="10">The dorsal ridge and ventral keel scales are notably differentiated from the flank scales.</paragraph>
|
||
<paragraph id="97623B2FFFD2103059B6D7A5FBAE18CF" blockId="9.[189,1399,504,671]" pageId="9" pageNumber="10">There are 19 dorsal ridge scales between the dermosupraoccipital and the origin of the dorsal fin but only the first and the seventh to the tenth are well preserved. The first dorsal scale is small, triangular in shape and located just behind the dermosupraoccipital. The five following scales are badly crushed. Only fragments of the last nine are visible in a fissure of the substratum at the dorsal apex level. Each upper margin of the seventh to tenth dorsal scales bears up to six small spines.</paragraph>
|
||
<paragraph id="97623B2FFFD2103059B6D49BFA8A1976" blockId="9.[189,1399,710,807]" pageId="9" pageNumber="10">
|
||
The total number of ventral ke el scales is unknown. The ventral keel begins with 3 scales located under the cleithrum (
|
||
<figureCitation id="0FE627AAFFD210305863D4B4FE4C1953" box="[360,433,745,771]" captionStart="Fig" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[197,1389,616,1910]" captionTargetId="figure@6.[189,1399,616,1921]" captionTargetPageId="6" captionText="Fig. 4. Rhinopycnodus gabriellae gen. et sp. nov. Reconstruction of the skull and shoulder girdle of holotype CLC S-725." figureDoi="http://doi.org/10.5281/zenodo.3827662" httpUri="https://zenodo.org/record/3827662/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
|
||
). The first one bears a few very small spines. The second one has a large spine and the third one two large spines. A few ventral keel scales bearing very small spines are visible on the
|
||
</paragraph>
|
||
<caption id="C3A26BA7FFD2103059B6D1F1FD131DBA" ID-DOI="http://doi.org/10.5281/zenodo.3827668" ID-Zenodo-Dep="3827668" httpUri="https://zenodo.org/record/3827668/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1964,1990]" targetBox="[215,1369,869,1923]" targetPageId="9">
|
||
<paragraph id="97623B2FFFD2103059B6D1F1FD131DBA" blockId="9.[189,1399,1964,2026]" pageId="9" pageNumber="10">
|
||
<emphasis id="A5A9E73DFFD2103059B6D1F1FEF61D97" bold="true" box="[189,267,1964,1991]" pageId="9" pageNumber="10">Fig. 7.</emphasis>
|
||
<taxonomicName id="50DD40ACFFD210305818D1F1FDB51D96" authority="Taverne & Capasso, 2013" authorityName="Taverne & Capasso" authorityYear="2013" box="[275,584,1964,1990]" class="Actinopterygii" family="Pycnodontes" genus="Rhinopycnodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="gabriellae" status="gen. et sp. nov.">
|
||
<emphasis id="A5A9E73DFFD210305818D1F1FDB51D96" box="[275,584,1964,1990]" italics="true" pageId="9" pageNumber="10">Rhinopycnodus gabriellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BE9A5A46FFD210305B5BD1F0FCFC1D97" box="[592,769,1965,1991]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Caudal skeleton of holotype CLC S-725. The arrows point at the more external principal caudal rays.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97623B2FFFD1103359B6D756FDAC1B19" blockId="10.[189,1398,267,329]" pageId="10" pageNumber="11">ventral contour between the cleithrum and the pelvic girdle, but some elements of this series are lost because of the taphonomic events.</paragraph>
|
||
<paragraph id="97623B2FFFD1103359B6D72DFE301BFD" blockId="10.[189,1398,368,429]" pageId="10" pageNumber="11">There are 3 pelvic scales associated with the pelvic bones and 2 postcloacal scales are located just before the postcoelomic bone.</paragraph>
|
||
<paragraph id="97623B2FFFD1103359B6D788FAF81892" blockId="10.[189,1399,469,706]" pageId="10" pageNumber="11">
|
||
There are flank scales only in the abdominal region of the body, anterior to the origin of the dorsal and anal fins. In the ventralmost area of the
|
||
<emphasis id="A5A9E73DFFD110335B8AD7A4FCDB1842" box="[641,806,505,530]" italics="true" pageId="10" pageNumber="11">situs viscerum</emphasis>
|
||
, between the cleithrum and the postcoelomic bone, the scales are complete, thick, deep, broad, slightly ornamented with small tubercules and articulated together. There are 11 rows of these large ventral flank scales. The other body scales are reduced to scale bars. In the dorsal area of the abdominal region the scale bars are badly preserved and only fragments are visible between the neural spines. Scale bars also are associated with the first eight dorsal ridge scales. The scales linked to the eleven other dorsal ridge scales are progressively broader and longer.
|
||
</paragraph>
|
||
<paragraph id="97623B2FFFD1103359B6D4B4FB431953" blockId="10.[189,1214,745,772]" box="[189,1214,745,772]" pageId="10" pageNumber="11">The first scale of the lateral line is visible beneath the brush-like process of the parietal.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |