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<document id="EFBEB264BACC6CE30A908FAFC9BBA0DD" ID-CLB-Dataset="53658" ID-DOI="10.11646/zootaxa.5231.5.1" ID-GBIF-Dataset="30c14449-ed8b-4f00-b877-9ef2c72adb08" ID-ISSN="1175-5326" ID-Zenodo-Dep="7609508" ID-ZooBank="22143C8D-9639-4B87-8D8C-C3BB421534A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1675673990683" checkinUser="plazi" docAuthor="Strümpher, Werner P. &amp; Kalawate, Aparna S." docDate="2023" docId="14216F6BFF9FA134FF2DFBEE9755C287" docLanguage="en" docName="zootaxa.5231.5.1.pdf" docOrigin="Zootaxa 5231 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omorgus (Omorgus) khandesh Strumpher &amp; Kalawate, new 2023, new" docType="treatment" docUuid="3CB1C743-5D9B-4547-B64A-B3CBB3860D80" docUuidSource="ZooBank" docVersion="6" lastPageNumber="506" masterDocId="E8181713FF9DA131FFBAFFD19662C642" masterDocTitle="Omorgus (Omorgus) khandesh (Coleoptera: Scarabaeoidea: Trogidae), a new species from India, along with an annotated catalogue of the Oriental and Palearctic species of the subgenus Omorgus Erichson, 1847" masterLastPageNumber="522" masterPageNumber="501" pageNumber="503" updateTime="1699474605112" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BD8AA76A2EE87FB7240DD1EEEAC58682">Omorgus (Omorgus) khandesh (Coleoptera: Scarabaeoidea: Trogidae), a new species from India, along with an annotated catalogue of the Oriental and Palearctic species of the subgenus Omorgus Erichson, 1847</mods:title>
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<mods:namePart id="165AD56259DF64AC390AF38D252DF50F">Strümpher, Werner P.</mods:namePart>
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<mods:namePart id="DA309B73D88D0D9762953A97975AA814">Kalawate, Aparna S.</mods:namePart>
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<mods:affiliation id="B730AFF66B004D4ED824CE0B56A51142">Zoological Survey of India, Western Regional Centre, Vidhya Nagar, Sector- 29, P. C. N. T. (PO), Rawet Road, Akurdi, Pune, Maharashtra 411044, India. aparna _ ent @ yahoo. co. in; https: // orcid. org / 0000 - 0001 - 6595 - 6749</mods:affiliation>
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<treatment id="14216F6BFF9FA134FF2DFBEE9755C287" ID-DOI="http://doi.org/10.5281/zenodo.7614724" ID-GBIF-Taxon="209281823" ID-Zenodo-Dep="7614724" LSID="urn:lsid:zoobank.org:act:3CB1C743-5D9B-4547-B64A-B3CBB3860D80" httpUri="http://treatment.plazi.org/id/14216F6BFF9FA134FF2DFBEE9755C287" lastPageId="5" lastPageNumber="506" pageId="2" pageNumber="503">
<subSubSection id="D4928DF6FF9FA133FF2DFBEE95BCC21A" box="[151,990,1086,1113]" pageId="2" pageNumber="503" type="nomenclature">
<paragraph id="9C37DE7DFF9FA133FF2DFBEE95BCC21A" blockId="2.[151,990,1086,1149]" box="[151,990,1086,1113]" pageId="2" pageNumber="503">
<heading id="C77F6911FF9FA133FF2DFBEE95BCC21A" bold="true" box="[151,990,1086,1113]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="AEFC026FFF9FA133FF2DFBEE95BCC21A" bold="true" box="[151,990,1086,1113]" pageId="2" pageNumber="503">
<taxonomicName id="5B88A5FEFF9FA133FF2DFBEE95E3C21B" ID-CoL="B3FND" authority="Strumpher &amp; Kalawate" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[151,897,1086,1113]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="khandesh" status="new" subGenus="Omorgus">
<emphasis id="AEFC026FFF9FA133FF2DFBEE9765C21B" bold="true" box="[151,263,1087,1113]" italics="true" pageId="2" pageNumber="503">Omorgus</emphasis>
(
<emphasis id="AEFC026FFF9FA133FEA3FBEE97EBC21B" bold="true" box="[281,393,1087,1113]" italics="true" pageId="2" pageNumber="503">Omorgus</emphasis>
)
<emphasis id="AEFC026FFF9FA133FE20FBEF946DC21B" bold="true" box="[410,527,1086,1113]" italics="true" pageId="2" pageNumber="503">khandesh</emphasis>
Strümpher &amp; Kalawate,
<taxonomicNameLabel id="B5CFBF14FF9FA133FCF4FBEE95E3C21B" box="[846,897,1087,1113]" pageId="2" pageNumber="503" rank="species">new</taxonomicNameLabel>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D4928DF6FF9FA133FF2DFBB29707C23F" box="[151,357,1123,1149]" pageId="2" pageNumber="503" type="description">
<paragraph id="9C37DE7DFF9FA133FF2DFBB29707C23F" blockId="2.[151,990,1086,1149]" box="[151,357,1123,1149]" pageId="2" pageNumber="503">
(
<figureCitation id="04B3C2F8FF9FA133FF25FBB2976EC23F" box="[159,268,1123,1149]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="2" pageNumber="503">Figs. 36</figureCitation>
,
<figureCitation id="04B3C2F8FF9FA133FEADFBB2973EC23F" box="[279,348,1123,1149]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="2" pageNumber="503">1113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4928DF6FF9FA133FF2DFB7A957DC33B" pageId="2" pageNumber="503" type="materials_examined">
<paragraph id="9C37DE7DFF9FA133FF2DFB7A9680C2AB" blockId="2.[151,1436,1194,2013]" pageId="2" pageNumber="503">
<materialsCitation id="2CE0D420FF9FA133FF2DFB7A96BCC2AB" ID-GBIF-Occurrence="4026640305" country="India" county="Nandurbar" location="Jalgaon" municipality="Dhule" pageId="2" pageNumber="503" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<emphasis id="AEFC026FFF9FA133FF2DFB7A9756C286" bold="true" box="[151,308,1194,1221]" pageId="2" pageNumber="503">
<typeStatus id="433360DFFF9FA133FF2DFB7A96B0C287" box="[151,210,1195,1221]" pageId="2" pageNumber="503">Type</typeStatus>
locality.
</emphasis>
“Khandesh” [= approximately the present-day
<location id="995788A6FF9FA133FC80FB7A95F0C287" LSID="urn:lsid:plazi:treatment:14216F6BFF9FA134FF2DFBEE9755C287:995788A6FF9FA133FC80FB7A95F0C287" box="[826,914,1195,1221]" country="India" county="Nandurbar" municipality="Dhule" name="Jalgaon" pageId="2" pageNumber="503" stateProvince="Maharashtra">Jalgaon</location>
,
<collectingMunicipality id="7C534407FF9FA133FC26FB7A9582C287" box="[924,992,1195,1221]" pageId="2" pageNumber="503">Dhule</collectingMunicipality>
and
<collectingCounty id="7556A6F1FF9FA133FBA8FB7A92E8C287" box="[1042,1162,1195,1221]" pageId="2" pageNumber="503">Nandurbar</collectingCounty>
districts of
<collectingRegion id="5E4C109FFF9FA133FAB3FB7A93FAC287" box="[1289,1432,1195,1221]" country="India" name="Maharashtra" pageId="2" pageNumber="503">Maharashtra</collectingRegion>
,
<collectingCountry id="E49F9EEDFF9FA133FF2DFB1E96B5C2AB" box="[151,215,1231,1257]" name="India" pageId="2" pageNumber="503">India</collectingCountry>
]
</materialsCitation>
.
</paragraph>
<paragraph id="9C37DE7DFF9FA133FF7DFB22957DC33B" blockId="2.[151,1436,1194,2013]" pageId="2" pageNumber="503">
<emphasis id="AEFC026FFF9FA133FF7DFB229794C34E" bold="true" box="[199,502,1266,1293]" pageId="2" pageNumber="503">Type material examined.</emphasis>
<materialsCitation id="2CE0D420FF9FA133FDBAFB23957DC33B" ID-GBIF-Occurrence="4026640316" collectionCode="ZSI-WRC" collectorName="T. R. Bell" country="India" location="Maharashtra province" pageId="2" pageNumber="503" specimenCount="2" stateProvince="Maharashtra" typeStatus="holotype">
<emphasis id="AEFC026FFF9FA133FDBAFB23959CC352" bold="true" box="[512,1022,1266,1296]" pageId="2" pageNumber="503">
<collectingCountry id="E49F9EEDFF9FA133FDBAFB239423C34E" box="[512,577,1266,1292]" name="India" pageId="2" pageNumber="503">India</collectingCountry>
,
<collectingRegion id="5E4C109FFF9FA133FDEBFB229501C34E" box="[593,867,1266,1293]" country="India" name="Maharashtra" pageId="2" pageNumber="503">Maharashtra province</collectingRegion>
.
<typeStatus id="433360DFFF9FA133FCC8FB2295BDC34F" box="[882,991,1267,1293]" pageId="2" pageNumber="503" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="FA9946B8FF9FA133FBAAFB2292EAC34F" box="[1040,1160,1267,1293]" pageId="2" pageNumber="503">ZSI-WRC</collectionCode>
, ENT-1/3217, specimen cleaned, aedeagus extracted) (
<figureCitation id="04B3C2F8FF9FA133FE55FAC6943CC373" box="[495,606,1302,1329]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="2" pageNumber="503">Figs. 35</figureCitation>
): “
<collectorName id="317DBBABFF9FA133FD3BFAC69497C373" box="[641,757,1303,1329]" pageId="2" pageNumber="503">T. R. Bell</collectorName>
, | Khandesh. | [back, handwriting in pencil]: May [19]03 || Andrews | Bequest. | B.M.1922221. [rectangle, faded white cardstock] ||
<typeStatus id="433360DFFF9FA133FC01FAEA9253C317" box="[955,1073,1339,1365]" pageId="2" pageNumber="503" type="holotype">HOLOTYPE</typeStatus>
:
<taxonomicName id="5B88A5FEFF9FA133FB84FAEA92C7C317" authorityName="Erichson" authorityYear="1847" box="[1086,1189,1339,1365]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="AEFC026FFF9FA133FB84FAEA92C7C317" box="[1086,1189,1339,1365]" italics="true" pageId="2" pageNumber="503">Omorgus</emphasis>
</taxonomicName>
|
<taxonomicName id="5B88A5FEFF9FA133FB0EFAEA93FEC317" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[1204,1436,1339,1365]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="khandesh" subGenus="Omorgus">
(
<emphasis id="AEFC026FFF9FA133FB07FAEA9346C317" box="[1213,1316,1339,1365]" italics="true" pageId="2" pageNumber="503">Omorgus</emphasis>
)
<emphasis id="AEFC026FFF9FA133FA88FAEA93FEC317" box="[1330,1436,1339,1365]" italics="true" pageId="2" pageNumber="503">khandesh</emphasis>
</taxonomicName>
| Strümpher &amp; Kalawate 2022 [rectangle, red cardstock]”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4928DF6FF9FA134FF7DFA5397C0C557" lastPageId="5" lastPageNumber="506" pageId="2" pageNumber="503" type="description">
<paragraph id="9C37DE7DFF9FA133FF7DFA5395B6C3DF" blockId="2.[151,1436,1194,2013]" box="[199,980,1410,1439]" pageId="2" pageNumber="503">
<emphasis id="AEFC026FFF9FA133FF7DFA53978CC3DE" bold="true" box="[199,494,1410,1437]" pageId="2" pageNumber="503">
Description of
<typeStatus id="433360DFFF9FA133FECDFA5297BCC3DF" box="[375,478,1411,1437]" pageId="2" pageNumber="503" type="holotype">holotype</typeStatus>
(
</emphasis>
<emphasis id="AEFC026FFF9FA133FDB9FA539471C3DE" bold="true" box="[515,531,1410,1436]" pageId="2" pageNumber="503">).</emphasis>
<emphasis id="AEFC026FFF9FA133FDA0FA529425C3DF" box="[538,583,1411,1437]" italics="true" pageId="2" pageNumber="503">Size</emphasis>
. Length: 11.9 mm. Width: 6.1 mm.
</paragraph>
<paragraph id="9C37DE7DFF9FA133FF7DFA7695EBC3A7" blockId="2.[151,1436,1194,2013]" pageId="2" pageNumber="503">
<emphasis id="AEFC026FFF9FA133FF7DFA769725C383" box="[199,327,1447,1473]" italics="true" pageId="2" pageNumber="503">Body shape</emphasis>
. Strongly convex, elongate oval with almost parallel elytral margins, elytral profile convex, attaining maximum height behind the middle, strongly declivous posteriorly.
</paragraph>
<paragraph id="9C37DE7DFF9FA133FF7DFA3E92E5C06F" blockId="2.[151,1436,1194,2013]" pageId="2" pageNumber="503">
<emphasis id="AEFC026FFF9FA133FF7DFA3E9774C04B" box="[199,278,1519,1545]" italics="true" pageId="2" pageNumber="503">Colour</emphasis>
(
<figureCitation id="04B3C2F8FF9FA133FE9CFA3E97F0C04B" box="[294,402,1518,1545]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="2" pageNumber="503">Figs. 35</figureCitation>
). Matt black, surface of head, pronotum, legs, elytral margins and tubercles, and parts on the ventral side with light brown tomentose coating. Body setae mostly brown, to rust-brown.
</paragraph>
<paragraph id="9C37DE7DFF9FA133FF7DF9E69287C10F" blockId="2.[151,1436,1194,2013]" pageId="2" pageNumber="503">
<emphasis id="AEFC026FFF9FA133FF7DF9E69766C013" box="[199,260,1591,1617]" italics="true" pageId="2" pageNumber="503">Head</emphasis>
(
<figureCitation id="04B3C2F8FF9FA133FEAEF9E697E3C013" box="[276,385,1590,1617]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="2" pageNumber="503">Figs. 35</figureCitation>
). Clypeus obtusely triangular, with one raised oval tubercle, apex of clypeus pointed, margin reflexed, entire; surface sculpted with discrete and irregularly shaped punctures, these punctures close, each with a short stout seta, surface between punctures faintly tomentose. Frons raised with two large central tubercles, mediobasally with large flat nitid patch; surface and tubercles sculpted with discrete round punctures, punctures close, surface between punctures tomentose. Genal angles acute, recurved, deeply concave, surface sculpted with discrete and irregularly shaped punctures; gena and genal margin with scattered, straight setae. Eyes large and bulbous, barely visible in dorsal view. Antennal scape elongate, stout, with long straight brown setae; pedicel attached subapically; antennal club and setae fulvous. Setae around mouthparts dense, rust-brown.
</paragraph>
<paragraph id="9C37DE7DFF9FA132FF7DF8869466C47F" blockId="2.[151,1436,1194,2013]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<emphasis id="AEFC026FFF9FA133FF7DF8869754C132" box="[199,310,1879,1904]" italics="true" pageId="2" pageNumber="503">Pronotum</emphasis>
(
<figureCitation id="04B3C2F8FF9FA133FEF2F88697DAC133" box="[328,440,1878,1905]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="2" pageNumber="503">Figs. 35</figureCitation>
): Length of pronotum almost one third of elytral length, total pronotum width slightly narrower than elytron width, pronotum widest behind middle. Surface tomentose with discrete, rounded punctures, uniformly spaced, rather close, space between punctures less than diameter of punctures, each puncture bearing a short stout seta. Lateral margins broad and flat, explanate, attenuated anteriorly, margin evenly rounded over anterior-half towards anterior angle, anterior angle broadly obtuse, lateral margin emarginate in middle anterior to greatest width, there angle subobtuse, feebly extended laterally, then obliquely converging backwards, sinuate basally, posterior angles more distinct, almost right angled. Base of pronotum margined laterally, obliquely truncate behind posterior angles, base slightly projecting backward with distinctly produced posteromedian lobe, base slightly sinuated on either side of lobe. All margins smooth with short straight setae, edges thickly tomentose. Median discal area raised, stretching over anterior half of pronotum, flattened anteriorly, strongly declivous on sides. Pronotal disc with 2 broad elevated median longitudinal ridges. Ridges high, parallel, and not converging posteriorly. Median pronotal depression distinct, entire, fairly evenly wide, shallow anteriorly, deepest at posteromedian lobe. Medial basal tubercles distinct, slightly laterally offset, partially fused to median ridges, connected with lateral basal tubercles by faint marginal transverse ridge; anterior and lateral basal tubercles round; basal (larger) and apical (smaller) lateral tubercles large, fused forming distinct sinuous ridge on either side; short lateral carina present below lateral anterolateral ridge.
</paragraph>
<paragraph id="9C37DE7DFF9EA132FF7DFD969224C4C7" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="504">
<taxonomicName id="5B88A5FEFF9EA132FF7DFD969757C423" box="[199,309,583,609]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="AEFC026FFF9EA132FF7DFD969757C423" box="[199,309,583,609]" italics="true" pageId="3" pageNumber="504">Scutellum</emphasis>
</taxonomicName>
(
<figureCitation id="04B3C2F8FF9EA132FEFFFD9697EBC423" box="[325,393,583,609]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="3" pageNumber="504">Fig. 3</figureCitation>
): Hastate, slightly longer than wide, surface finely sculpted, mediobasally depressed, surface finely sculpted, apex slightly raised, distal half of scutellum and margins tomentose.
</paragraph>
<paragraph id="9C37DE7DFF9EA132FF7DFD5E95D8C373" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="504">
<emphasis id="AEFC026FFF9EA132FF7DFD5E976FC4EB" box="[199,269,655,681]" italics="true" pageId="3" pageNumber="504">Elytra</emphasis>
(
<figureCitation id="04B3C2F8FF9EA132FEA6FD5E97EBC4EB" box="[284,393,655,681]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="3" pageNumber="504">Figs. 34</figureCitation>
): Elongate, length of elytra equals of 2.4 pronotum length, lateral margins subparallel, smooth with tomentosity bearing row of fine short setae; sides wide, with low small round tubercles, these tubercles closely and crudely evenly spaced along elytral length, each with weak posterior tomentosity and 23 short stout setae. Humeral calli prominent bearing small tubercles with tomentosity and short setae, humeral angles obtuse, blunt. Sutural margin raised, surface with tomentosity, margin with raised elongate to round tubercles, separated, evenly distributed along margin; these tubercles smooth, shiny, each with posterior tomentosity and short setae, tubercles elongate and roughly of similar size for 0.75 elytral length, thereafter, becoming rounder and decreasing in size until barely discernible. Elytral costae distinct, even numbered costae more prominent than odd-numbered. Apical callosity present on fourth costal interval at top of elytral declivity. Even-numbered costae prominent, with distinctly raised, oval to round shiny tubercles (on intervals 2, 4 and 6), these tubercles close and regularly spaced along elytral length, roughly equal in size and height for 0.75 elytral length thereafter decreasing in size and height; costae 2 and 4 with basal 0.25 fused into partial ridges; tubercles on even-numbered costae each with tomentose posterior patches and short setae, anterior half of tubercle smooth and shiny; costae 8 with smaller, widely spaced, round tubercles, slightly raised, each tomentose with short setae. Odd-numbered costae marked by row of small round to oval tubercles and irregular shiny flat nitid areas (especially noticeable on intervals 1, 3 and 5; interval 2 without nitid patches); tubercles fairly evenly spaced along elytral length, size variable, each with tomentose patch; nitid areas slightly raised, irregular in shape. Intercostae marked by row of small round shiny punctures, puncture regularly spaced over length of elytra. Elytral surface matt, mostly smooth and without undulations and grooves; elytral profile convex, attaining maximum height approximately in the middle.
</paragraph>
<paragraph id="9C37DE7DFF9EA132FF7DFAEA9448C317" blockId="3.[151,1437,151,2013]" box="[199,554,1339,1365]" pageId="3" pageNumber="504">
<emphasis id="AEFC026FFF9EA132FF7DFAEA97C4C316" box="[199,422,1339,1365]" italics="true" pageId="3" pageNumber="504">Metathoracic wings</emphasis>
: Complete.
</paragraph>
<paragraph id="9C37DE7DFF9EA132FF7DFA8E9263C19F" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="504">
<emphasis id="AEFC026FFF9EA132FF7DFA8E969EC33A" box="[199,252,1375,1400]" italics="true" pageId="3" pageNumber="504">Legs</emphasis>
(
<figureCitation id="04B3C2F8FF9EA132FEB4FA8E971EC33B" box="[270,380,1374,1401]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="3" pageNumber="504">Figs. 35</figureCitation>
). Protibia dorsally keeled, ending with small apical process, apex rounded; tibial spur stout, pointed forward, as long as four basal tarsomeres combined; tarsal segments smooth, without setae, fifth tarsomere with two claws, simple, equal; lateral tibial margin smooth, without teeth or indentations, dorsally outer tibial surface rugose and coarsely punctate; inner tibial surface (top lateral) smooth, glabrous, with round punctures running closely parallel to the outer and inner margins, punctures close, each with thin long seta; inner margin (lower half) punctate, punctures with long, dense setae; ventrally semi-flattened, submedially marginate, inner section smooth, glabrous without seate, outer section indumentose. Profemur large, widest basally, marginate on front face with row of straight forward pointing setae; ventral face with regularly spaced round punctures, each with single seta, surface indumentose; dorsal surfaces smooth. Procoxa narrow, elongate, flattened in front, surface rugose and punctuate. Meso- and metafemora slender, fore margins rounded without rows of setae; ventral faces with evenly spaced round punctures, each with a seta, space between punctures more than diameter of punctures; surface indumentose; dorsal surfaces smooth. Mesotibial and metatibial surfaces rugose, covered with thick indument, coarsely punctate, each puncture with a short seta, superomedial longitudinal carinae with row of stout setae; inner carinae with row of long slender setae; outer margin on apex of both meso- and metatibia with row of closely spaced stout backward pointing setae. Apex of mesotibia with two short tooth-like projections on inner margin beneath apical spurs. Mesotibia and metatibia with two apical spurs on inner margin; upper spur slightly longer and lower spur shorter than 1
<superScript id="6BFD7335FF9EA132FAACF8AA9343C1CB" attach="left" box="[1302,1313,1915,1929]" fontSize="6" pageId="3" pageNumber="504">st</superScript>
tarsomere. Mesotarsi and metatarsi indumentose with long setal brushes, tarsomeres 14 subequal in length, 5
<superScript id="6BFD7335FF9EA132FB43F84F9364C1EE" attach="left" box="[1273,1286,1950,1964]" fontSize="6" pageId="3" pageNumber="504">th</superScript>
tarsomere as long as tarsomers 3 and 4 combined; fifth tarsal with two claws simple, equal.
</paragraph>
<caption id="C8F78EF5FF99A135FF2DF8B8950CC1E4" ID-DOI="http://doi.org/10.5281/zenodo.7609514" ID-Zenodo-Dep="7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="4" pageNumber="505" startId="4.[151,264,1897,1921]" targetBox="[155,1416,193,1874]" targetPageId="4">
<paragraph id="9C37DE7DFF99A135FF2DF8B8950CC1E4" blockId="4.[151,1436,1897,1958]" pageId="4" pageNumber="505">
<emphasis id="AEFC026FFF99A135FF2DF8B89759C1C3" bold="true" box="[151,315,1897,1921]" pageId="4" pageNumber="505">FIGURES 36.</emphasis>
<taxonomicName id="5B88A5FEFF99A135FEF8F8BB95CBC1C0" authority="Strumpher &amp; Kalawate" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[322,937,1898,1922]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="506" phylum="Arthropoda" rank="species" species="khandesh" status="new" subGenus="Omorgus">
<emphasis id="AEFC026FFF99A135FEF8F8BB97FDC1C3" box="[322,415,1898,1921]" italics="true" pageId="4" pageNumber="505">Omorgus</emphasis>
(
<emphasis id="AEFC026FFF99A135FE15F8BB946EC1C3" box="[431,524,1898,1921]" italics="true" pageId="4" pageNumber="505">Omorgus</emphasis>
)
<emphasis id="AEFC026FFF99A135FDA6F8BB941EC1C0" box="[540,636,1898,1922]" italics="true" pageId="4" pageNumber="505">khandesh</emphasis>
Strümpher &amp; Kalawate,
<taxonomicNameLabel id="B5CFBF14FF99A135FCC7F8BB95CBC1C0" box="[893,937,1898,1922]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEFC026FFF99A135FCC7F8BB95CBC1C0" bold="true" box="[893,937,1898,1922]" pageId="4" pageNumber="505">new</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="AEFC026FFF99A135FC15F8B89598C1C3" bold="true" box="[943,1018,1897,1921]" pageId="4" pageNumber="505">species</emphasis>
, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels.
</paragraph>
</caption>
<paragraph id="9C37DE7DFF98A134FF7DFF469706C7B7" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="506">
<emphasis id="AEFC026FFF98A134FF7DFF469716C6F2" box="[199,372,151,177]" italics="true" pageId="5" pageNumber="506">Ventral aspects</emphasis>
:
<emphasis id="AEFC026FFF98A134FE3FFF469474C6F2" box="[389,534,151,176]" italics="true" pageId="5" pageNumber="506">Proepisterna</emphasis>
: Surfaces rugose and coarsely punctate, thickly indumentose, punctures round to oval.
<taxonomicName id="5B88A5FEFF98A134FF4CFF6A9704C697" box="[246,358,187,213]" class="Insecta" family="Uraniidae" genus="Epipleura" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="AEFC026FFF98A134FF4CFF6A9704C697" box="[246,358,187,213]" italics="true" pageId="5" pageNumber="506">Epipleura</emphasis>
</taxonomicName>
broad basally narrowing apically, concave along entire length, surface mostly smooth, weakly undulating, sparsely setose. Setae fine; inner margin with thick tomentosity.
<emphasis id="AEFC026FFF98A134FB8BFF0E92ABC6BA" box="[1073,1225,223,248]" italics="true" pageId="5" pageNumber="506">Mesoventrum</emphasis>
medially densely punctate, slightly depressed, punctures large, medial area covered with thick tomentosity, laterally surface smooth, without puncture.
<emphasis id="AEFC026FFF98A134FED2FEF6979DC702" box="[360,511,295,320]" italics="true" pageId="5" pageNumber="506">Mesepisterna</emphasis>
: Surface tomentose, punctate, punctures with seta.
<emphasis id="AEFC026FFF98A134FBF4FEF69280C702" box="[1102,1250,295,320]" italics="true" pageId="5" pageNumber="506">Metepisterna</emphasis>
: Surface rugose and covered with indument, coarsely punctate, punctures irregular in shape.
<emphasis id="AEFC026FFF98A134FBACFE9A92C9C726" box="[1046,1195,331,356]" italics="true" pageId="5" pageNumber="506">Metaventrum</emphasis>
: Surface rugose and coarsely punctate, covered with indument, disc rhomboidal, flat, declivous behind without covering of indument.
<emphasis id="AEFC026FFF98A134FF2DFE429718C7EE" box="[151,378,403,429]" italics="true" pageId="5" pageNumber="506">Abdominal ventrites</emphasis>
: Five visible ventrites (IVVIII); ventrite IV with rough surface, coarsely punctate; ventrites VVII smooth, finely punctate, distal potion ridged; ventrite VIII course densely punctate, punctures shallow, distal margin marginate.
</paragraph>
<paragraph id="9C37DE7DFF98A134FF7DFE2E9464C4B3" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="506">
<emphasis id="AEFC026FFF98A134FF7DFE2E970DC45B" box="[199,367,511,537]" italics="true" pageId="5" pageNumber="506">Male genitalia</emphasis>
(
<figureCitation id="04B3C2F8FF98A134FE3BFE2E946FC45B" box="[385,525,511,537]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Figs. 1113</figureCitation>
): Symmetrical, in profile slightly arched. Phallobasis divided and membranous dorsally (
<figureCitation id="04B3C2F8FF98A134FEB8FDF29737C47F" box="[258,341,547,573]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 11</figureCitation>
); phallobasis short, length 3.4 aedeagus length (in profile) (
<figureCitation id="04B3C2F8FF98A134FC44FDF29202C47F" box="[1022,1120,547,573]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 12c</figureCitation>
). Parameres applied closely to phallus and extended somewhat beyond phallus, parameres narrower than in
<taxonomicName id="5B88A5FEFF98A134FBA2FD969314C423" authority="(Haaf, 1957)" authorityName="Kalawate" authorityYear="2023" baseAuthorityName="Haaf" baseAuthorityYear="1957" box="[1048,1398,582,609]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rimulosus" subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FBA2FD969251C423" box="[1048,1075,583,609]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
(
<emphasis id="AEFC026FFF98A134FBFEFD96923DC423" box="[1092,1119,583,609]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FBCAFD9692BCC423" box="[1136,1246,583,609]" italics="true" pageId="5" pageNumber="506">rimulosus</emphasis>
(
<bibRefCitation id="F819A38CFF98A134FB54FD96930FC423" author="Haaf, E." box="[1262,1389,582,609]" pageId="5" pageNumber="506" pagination="692 - 695" refId="ref9103" refString="Haaf, E. (1957) Drei neue Trox - Arten (Col. Scarab). Entomologische Arbeiten aus dem Museum G. Frey, 8, 692 - 695." type="journal article" year="1957">Haaf, 1957</bibRefCitation>
)
</taxonomicName>
(in profile; compare
<figureCitation id="04B3C2F8FF98A134FEEFFDBA97D0C4C7" box="[341,434,619,645]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 12b</figureCitation>
with
<figureCitation id="04B3C2F8FF98A134FE54FDBA942AC4C7" box="[494,584,618,645]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 15f</figureCitation>
), apex of each sclerotised without membranous projection (compare
<figureCitation id="04B3C2F8FF98A134FAF8FDBA93FEC4C7" box="[1346,1436,619,645]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 12a</figureCitation>
with
<figureCitation id="04B3C2F8FF98A134FF75FD5E974FC4EB" box="[207,301,654,681]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 15e</figureCitation>
). Phallus dorsally clearly concave anteriorly and projecting dorsolaterally forming two distinct humps (distinct in lateral view) (
<figureCitation id="04B3C2F8FF98A134FE04FD629442C48F" box="[446,544,691,717]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 12d</figureCitation>
) thereafter attenuated anteriorly, apex pointed, margin straight (not recurved) (compare
<figureCitation id="04B3C2F8FF98A134FEB2FD06973DC4B3" box="[264,351,727,753]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 13i</figureCitation>
with
<figureCitation id="04B3C2F8FF98A134FE25FD06979BC4B3" box="[415,505,727,753]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Fig. 16j</figureCitation>
).
</paragraph>
<paragraph id="9C37DE7DFF98A134FF7DFD2A97C0C557" blockId="5.[151,1437,151,1221]" box="[199,418,763,789]" pageId="5" pageNumber="506">
<emphasis id="AEFC026FFF98A134FF7DFD2A9747C557" bold="true" box="[199,293,763,789]" pageId="5" pageNumber="506">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4928DF6FF98A134FF7DFCCF9276C21B" pageId="5" pageNumber="506" type="diagnosis">
<paragraph id="9C37DE7DFF98A134FF7DFCCF9276C21B" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="506">
<emphasis id="AEFC026FFF98A134FF7DFCCF9720C57A" bold="true" box="[199,322,798,824]" pageId="5" pageNumber="506">Diagnosis.</emphasis>
<taxonomicName id="5B88A5FEFF98A134FEF3FCCE950AC57B" authority="Strumpher &amp; Kalawate" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[329,872,799,825]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="khandesh" status="sp. nov." subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FEF3FCCE97D2C57B" box="[329,432,799,825]" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis>
(
<emphasis id="AEFC026FFF98A134FE78FCCE97BFC57B" box="[450,477,799,825]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FE54FCCE943AC57B" box="[494,600,799,825]" italics="true" pageId="5" pageNumber="506">khandesh</emphasis>
Strümpher &amp; Kalawate
</taxonomicName>
,
<emphasis id="AEFC026FFF98A134FCCEFCCE959CC57A" bold="true" box="[884,1022,798,825]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="B5CFBF14FF98A134FCCEFCCE959CC57A" box="[884,1022,798,825]" pageId="5" pageNumber="506" rank="species">new species</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="5B88A5FEFF98A134FB8CFCCE93F5C57B" authority="(Haaf, 1957)" authorityName="Kalawate" authorityYear="2023" baseAuthorityName="Haaf" baseAuthorityYear="1957" box="[1078,1431,798,825]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rimulosus" subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FB8CFCCE9233C57B" box="[1078,1105,799,825]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
(
<emphasis id="AEFC026FFF98A134FBD9FCCE921CC57B" box="[1123,1150,799,825]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FB35FCCE929FC57B" box="[1167,1277,799,825]" italics="true" pageId="5" pageNumber="506">rimulosus</emphasis>
(
<bibRefCitation id="F819A38CFF98A134FAB7FCCE93F2C57B" author="Haaf, E." box="[1293,1424,798,825]" pageId="5" pageNumber="506" pagination="692 - 695" refId="ref9103" refString="Haaf, E. (1957) Drei neue Trox - Arten (Col. Scarab). Entomologische Arbeiten aus dem Museum G. Frey, 8, 692 - 695." type="journal article" year="1957">Haaf, 1957</bibRefCitation>
)
</taxonomicName>
, are morphologically very similar, but the former can be distinguished from the latter in having the tubercles on the even numbered costae close together, smaller, and not as highly elevated (
<figureCitation id="04B3C2F8FF98A134FBBDFCB69217C5C3" box="[1031,1141,871,897]" captionStart="FIGURES 36" captionStartId="4.[151,264,1897,1921]" captionTargetBox="[155,1416,193,1874]" captionTargetId="figure-22@4.[151,1436,181,1874]" captionTargetPageId="4" captionText="FIGURES 36. Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC). 3, habitus, dorsal view; 4, habitus, lateral view; 5, head and pronotum; 6, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609514" httpUri="https://zenodo.org/record/7609514/files/figure.png" pageId="5" pageNumber="506">Figs. 34</figureCitation>
).
<taxonomicName id="5B88A5FEFF98A134FB33FCB69744C5E7" authority="(Haaf, 1957)" authorityName="Kalawate" authorityYear="2023" baseAuthorityName="Haaf" baseAuthorityYear="1957" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rimulosus" subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FB33FCB69292C5C3" box="[1161,1264,871,897]" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis>
(
<emphasis id="AEFC026FFF98A134FABBFCB6937FC5C3" box="[1281,1309,871,897]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FA94FCB693FEC5C3" box="[1326,1436,871,897]" italics="true" pageId="5" pageNumber="506">rimulosus</emphasis>
(
<bibRefCitation id="F819A38CFF98A134FF25FC5A977CC5E7" author="Haaf, E." box="[159,286,906,933]" pageId="5" pageNumber="506" pagination="692 - 695" refId="ref9103" refString="Haaf, E. (1957) Drei neue Trox - Arten (Col. Scarab). Entomologische Arbeiten aus dem Museum G. Frey, 8, 692 - 695." type="journal article" year="1957">Haaf, 1957</bibRefCitation>
)
</taxonomicName>
have these tubercles considerably larger, widely spaced along elytral length and more raised (
<figureCitation id="04B3C2F8FF98A134FAFBFC5A96C5C58B" captionStart="FIGURES 710" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[216,1364,202,1903]" captionTargetId="figure-19@7.[199,1389,181,1920]" captionTargetPageId="7" captionText="FIGURES 710. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype NHMB. 7, habitus, dorsal view; 8, habitus, lateral view; 9, habitus, ventral view 10, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.7609516" httpUri="https://zenodo.org/record/7609516/files/figure.png" pageId="5" pageNumber="506">Figs. 7 8</figureCitation>
). The male genitalia of the two species are distinctive with the most important differences found on the phalluses, parameres, and phallobases as described in the text (and as shown in
<figureCitation id="04B3C2F8FF98A134FC2DFC029278C5AF" box="[919,1050,979,1005]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Figs. 1113</figureCitation>
versus
<figureCitation id="04B3C2F8FF98A134FBD5FC029294C5AF" box="[1135,1270,979,1005]" captionStart="FIGURES 1116" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 1116. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 1113. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 1416. aj, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" pageId="5" pageNumber="506">Figs. 1416</figureCitation>
; see also
<tableCitation id="D10AEBC6FF98A134FAE4FC0296C6C253" captionStart="TABLE 1" captionStartId="5.[151,239,1268,1292]" captionTargetBox="[166,1422,1345,1947]" captionText="TABLE 1. Summary of morphological distinction between Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species and Omorgus (Omorgus) rimulosus (Haaf, 1954)." httpUri="http://table.plazi.org/id/C8F78EF5FF98A134FF2DFB25949EC373" pageId="5" pageNumber="506" tableUuid="C8F78EF5FF98A134FF2DFB25949EC373">Table 1</tableCitation>
). Both
<taxonomicName id="5B88A5FEFF98A134FF42FC2694A0C253" authority="Strumpher &amp; Kalawate" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[248,706,1015,1041]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="khandesh" status="sp. nov." subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FF42FC269771C253" box="[248,275,1015,1041]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
(
<emphasis id="AEFC026FFF98A134FE99FC26975CC253" box="[291,318,1015,1041]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FEF7FC2697D5C253" box="[333,439,1015,1041]" italics="true" pageId="5" pageNumber="506">khandesh</emphasis>
Strümpher &amp; Kalawate
</taxonomicName>
,
<emphasis id="AEFC026FFF98A134FD76FC269536C252" bold="true" box="[716,852,1014,1041]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="B5CFBF14FF98A134FD76FC269536C252" box="[716,852,1014,1041]" pageId="5" pageNumber="506" rank="species">new species</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="5B88A5FEFF98A134FC33FC269283C253" authority="(Haaf, 1957)" authorityName="Kalawate" authorityYear="2023" baseAuthorityName="Haaf" baseAuthorityYear="1957" box="[905,1249,1014,1041]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rimulosus" subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FC33FC2695C6C253" box="[905,932,1015,1041]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
(
<emphasis id="AEFC026FFF98A134FC0EFC2695ADC253" box="[948,975,1015,1041]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FC64FC26922EC253" box="[990,1100,1015,1041]" italics="true" pageId="5" pageNumber="506">rimulosus</emphasis>
(
<bibRefCitation id="F819A38CFF98A134FBE0FC2692BAC253" author="Haaf, E." box="[1114,1240,1014,1041]" pageId="5" pageNumber="506" pagination="692 - 695" refId="ref9103" refString="Haaf, E. (1957) Drei neue Trox - Arten (Col. Scarab). Entomologische Arbeiten aus dem Museum G. Frey, 8, 692 - 695." type="journal article" year="1957">Haaf, 1957</bibRefCitation>
)
</taxonomicName>
have an external appearance similar to
<taxonomicName id="5B88A5FEFF98A134FE36FBCA94BCC277" authority="(Haaf 1954)" baseAuthorityName="Haaf" baseAuthorityYear="1954" box="[396,734,1050,1077]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="maissouri" subGenus="Omorgus">
<emphasis id="AEFC026FFF98A134FE36FBCA97C5C277" box="[396,423,1051,1077]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
(
<emphasis id="AEFC026FFF98A134FE0DFBCA97B0C277" box="[439,466,1051,1077]" italics="true" pageId="5" pageNumber="506">O.</emphasis>
)
<emphasis id="AEFC026FFF98A134FE58FBCA9432C276" box="[482,592,1051,1076]" italics="true" pageId="5" pageNumber="506">maissouri</emphasis>
(Haaf 1954)
</taxonomicName>
(
<figureCitation id="04B3C2F8FF98A134FD57FBCA95FFC277" box="[749,925,1051,1077]" captionStart="FIGURES 2831" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[191,1400,201,1910]" captionTargetId="figure-22@14.[174,1414,181,1919]" captionTargetPageId="14" captionText="FIGURES 2831. Omorgus (Omorgus) maissouri (Haaf, 1954), holotype (MNHN). 28, habitus, dorsal view; 29, habitus, lateral view; 30, holotype labels; 31, aedeagus, dorsal and lateral view. Photos by Antoine Mantilleri, copyright MNHN." figureDoi="http://doi.org/10.5281/zenodo.7609540" httpUri="https://zenodo.org/record/7609540/files/figure.png" pageId="5" pageNumber="506">Figs. 2829, 31</figureCitation>
), but both can be distinguished from the latter by the appearance of the pronotum and elytra and the distinctive male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="D4928DF6FF98A134FF7DFBB394B8C23F" box="[199,730,1122,1149]" pageId="5" pageNumber="506" type="distribution">
<paragraph id="9C37DE7DFF98A134FF7DFBB394B8C23F" blockId="5.[151,1437,151,1221]" box="[199,730,1122,1149]" pageId="5" pageNumber="506">
<emphasis id="AEFC026FFF98A134FF7DFBB39703C23E" bold="true" box="[199,353,1122,1148]" pageId="5" pageNumber="506">Distribution.</emphasis>
Known only from Western
<collectingCountry id="E49F9EEDFF98A134FD23FBB294B4C23F" box="[665,726,1123,1149]" name="India" pageId="5" pageNumber="506">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D4928DF6FF98A134FF7DFB569755C287" pageId="5" pageNumber="506" type="etymology">
<paragraph id="9C37DE7DFF98A134FF7DFB569755C287" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="506">
<emphasis id="AEFC026FFF98A134FF7DFB56972CC2E3" bold="true" box="[199,334,1159,1185]" pageId="5" pageNumber="506">Etymology.</emphasis>
The species name is a noun in apposition referring to the Khandesh Region where the
<typeStatus id="433360DFFF98A134FA80FB5693FEC2E3" box="[1338,1436,1159,1185]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>