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<mods:title id="EDCB12C3BCA5E910C26A3BD0A33A7C62">Taxonomic review of the genera Micridium Motschulsky, 1869 and Micridina Johnson, 1969 (Coleoptera: Ptiliidae) with eleven new species including the first records from South America and Madagascar</mods:title>
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<treatment id="03E64B08E339FFE685855F74FCFAEA08" ID-DOI="http://doi.org/10.5281/zenodo.6045225" ID-GBIF-Taxon="127843543" ID-Zenodo-Dep="6045225" LSID="urn:lsid:plazi:treatment:03E64B08E339FFE685855F74FCFAEA08" httpUri="http://treatment.plazi.org/id/03E64B08E339FFE685855F74FCFAEA08" lastPageId="18" lastPageNumber="251" pageId="17" pageNumber="250">
<subSubSection id="C355A995E339FFE585855F74FDB7EE60" pageId="17" pageNumber="250" type="nomenclature">
<paragraph id="8BF0FA1EE339FFE585855F74FC39EE5F" blockId="17.[151,898,1589,1648]" box="[151,898,1589,1615]" pageId="17" pageNumber="250">
<heading id="D0B84D72E339FFE585855F74FC39EE5F" bold="true" box="[151,898,1589,1615]" fontSize="11" level="1" pageId="17" pageNumber="250" reason="1">
<emphasis id="B93B260CE339FFE585855F74FC39EE5F" bold="true" box="[151,898,1589,1615]" pageId="17" pageNumber="250">
<taxonomicName id="4C4F819DE339FFE585855F74FDE3EE5F" ID-CoL="73FYB" authority="Casey, 1924" authorityName="Casey" authorityYear="1924" box="[151,600,1589,1615]" class="Insecta" family="Ptiliidae" genus="Micridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="250" phylum="Arthropoda" rank="species" species="rhodeanum">
<emphasis id="B93B260CE339FFE585855F74FE10EE5F" bold="true" box="[151,427,1589,1615]" italics="true" pageId="17" pageNumber="250">Micridium rhodeanum</emphasis>
(
<bibRefCitation id="EFDE87EFE339FFE584A85F74FDEBEE5F" author="Casey" box="[442,592,1589,1615]" pageId="17" pageNumber="250" refString="Casey, T. L. (1924) Additions to the known Coleoptera of North America. In Memoirs on the Coleoptera. Uol. 11. Lancaster Press, Lancaster, Pa., 347 pp." type="book" year="1924">Casey, 1924</bibRefCitation>
)
</taxonomicName>
as
<taxonomicName id="4C4F819DE339FFE587935F74FC39EE5F" box="[641,898,1589,1615]" class="Insecta" family="Ptiliidae" genus="Dilinium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="250" phylum="Arthropoda" rank="species" species="rhodeanum">
<emphasis id="B93B260CE339FFE587935F74FC39EE5F" bold="true" box="[641,898,1589,1615]" italics="true" pageId="17" pageNumber="250">Dilinium rhodeanum</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF0FA1EE339FFE585855F16FDB7EE60" blockId="17.[151,898,1589,1648]" box="[151,524,1623,1648]" pageId="17" pageNumber="250">
(
<figureCitation id="1374E69BE339FFE5858D5F16FF56EE60" box="[159,237,1623,1648]" captionStart="FIGURES 1 14" captionStartId="5.[151,264,1856,1879]" captionTargetBox="[208,1396,215,1756]" captionTargetId="figure@5.[151,1436,195,1835]" captionTargetPageId="5" captionText="FIGURES 1 14. Micridium habitus. 1. M. angulicolle (Fairmaire) 2. M. halidaii (Matthews) 3. M. vittatum Motschulsky 4. M. lineatum Le Conte 5. M. rhodeanum (Casey) 6. M. hilli (Johnson) 7. M. johnsoni sp. n. 8. M. elegans sp. n. 9. M. oweni sp. n. 10. M. attenboroughi sp. n. 11. M. boliviense sp. n. 12. M. newtoni sp. n. 13. M. foveatum sp. n. 14. M. thayerae sp. n. 15. M. proprium sp. n. 16. M. quadridens sp. n." httpUri="https://zenodo.org/record/376375/files/figure.png" pageId="17" pageNumber="250">Figs. 5</figureCitation>
,
<figureCitation id="1374E69BE339FFE585E85F19FEA2EE60" box="[250,281,1624,1648]" captionStart="FIGURES 17 32" captionStartId="6.[151,264,1880,1903]" captionTargetBox="[238,1353,189,1847]" captionTargetId="figure@6.[205,1371,157,1869]" captionTargetPageId="6" captionText="FIGURES 17 32. Micridium pronotums. 17. M. quadridens sp. n. x 605. 18. M. inornatum sp. n. x 590. 19. M. boliviense sp. n. x 625. 20. M. newtoni sp. n. x 695. 21. M. foveatum sp. n. x 830. 22. M. johnsoni sp. n. x 790. 23. M. elegans sp. n. x 810. 24. M. thayerae sp. n. x 650. 25. M. halidaii (Matthews) x 770. 26. M. hilli (Johnson) x 740. 27. M. oweni sp. n. x 575. 28. M. attenboroughi sp. n. x 675. 29. M. angulicolle (Fairmaire). 30. M. rhodeanum (Casey) 31. M. vittatum Motschulsky 32. M. lineatum Le Conte." httpUri="https://zenodo.org/record/376376/files/figure.png" pageId="17" pageNumber="250">30</figureCitation>
,
<figureCitation id="1374E69BE339FFE584375F16FEF8EE60" box="[293,323,1623,1648]" captionStart="FIGURES 47 60" captionStartId="10.[151,264,1643,1666]" captionTargetBox="[163,1408,218,1606]" captionTargetId="figure@10.[151,1436,193,1622]" captionTargetPageId="10" captionText="FIGURES 47 60. Micridium mesoventra. 47. M. inornatum sp. n. x 540. 48. M. quadridens sp. n. x 710. 49. M. boliviense sp. n. x 695. 50. M. attenboroughi sp. n. x 870. 51. M. proprium sp. n. x 920. 52. M. newtoni sp. n. 53. M. johnsoni sp. n. x 655. 54. M. thayerae sp. n. x 860. 55. M. halidaii (Matthews) x 615. 56. M. oweni sp. n. x 585. 57. M. elegans sp. n. x 705. 58. M. hilli (Johnson) x 760. 59. M. foveatum sp. n. x 1300. 60. M. rhodeanum (Casey) x 545." httpUri="https://zenodo.org/record/376379/files/figure.png" pageId="17" pageNumber="250">60</figureCitation>
,
<figureCitation id="1374E69BE339FFE5845C5F19FED7EE60" box="[334,364,1624,1648]" captionStart="FIGURES 61 71" captionStartId="12.[151,264,1919,1942]" captionTargetBox="[328,1250,191,1869]" captionTargetId="figure@12.[311,1262,171,1888]" captionTargetPageId="12" captionText="FIGURES 61 71. Micridium metepiventral sutures. 61. M. johnsoni sp. n. x 530. 62. M. thayerae sp. n. x 505. 63. M. attenboroughi sp. n. x 670. 64. M. foveatum sp. n. x 735. 65. M. quadridens sp. n. x 710. 66. M. elegans sp. n. x 730. 67. M. oweni sp. n. x 585. 68. M. hilli (Johnson) x 760. 69. M. halidaii (Matthews) x 615. 70. M. newtoni sp. n. x 710. 71. M. rhodeanum (Casey) x 545." httpUri="https://zenodo.org/record/376380/files/figure.png" pageId="17" pageNumber="250">71</figureCitation>
, 90a, 90b, 99)
</paragraph>
</subSubSection>
<subSubSection id="C355A995E339FFE685855FE1FDBAE8E8" lastPageId="18" lastPageNumber="251" pageId="17" pageNumber="250" type="description">
<paragraph id="8BF0FA1EE339FFE585855FE1FDC2EEA6" blockId="17.[151,633,1696,1718]" box="[151,633,1696,1718]" pageId="17" pageNumber="250">
<taxonomicName id="4C4F819DE339FFE585855FE1FE69EEA6" authority="Casey" authorityName="Casey" box="[151,466,1696,1718]" class="Insecta" family="Ptiliidae" genus="Micridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="250" phylum="Arthropoda" rank="species" species="rhodeanum">
<emphasis id="B93B260CE339FFE585855FE1FEC0EEA6" box="[151,379,1696,1718]" italics="true" pageId="17" pageNumber="250">Micridium rhodeanum</emphasis>
(Casey)
</taxonomicName>
;
<bibRefCitation id="EFDE87EFE339FFE584CF5FE1FDC2EEA6" author="Johnson" box="[477,633,1696,1718]" pageId="17" pageNumber="250" refString="Johnson, C. (2001) Notes of Palearctic Ptiliidae (Coleoptera). Entomologists's Gazette, 52, 129 - 137." type="journal article" year="2001">Johnson (2001)</bibRefCitation>
</paragraph>
<paragraph id="8BF0FA1EE339FFE685855FA3FDDFE8C5" blockId="17.[151,1437,1762,2003]" lastBlockId="18.[151,1436,151,536]" lastPageId="18" lastPageNumber="251" pageId="17" pageNumber="250">
Habitus
<figureCitation id="1374E69BE339FFE585E75FA3FE81EEEB" box="[245,314,1762,1787]" captionStart="FIGURES 1 14" captionStartId="5.[151,264,1856,1879]" captionTargetBox="[208,1396,215,1756]" captionTargetId="figure@5.[151,1436,195,1835]" captionTargetPageId="5" captionText="FIGURES 1 14. Micridium habitus. 1. M. angulicolle (Fairmaire) 2. M. halidaii (Matthews) 3. M. vittatum Motschulsky 4. M. lineatum Le Conte 5. M. rhodeanum (Casey) 6. M. hilli (Johnson) 7. M. johnsoni sp. n. 8. M. elegans sp. n. 9. M. oweni sp. n. 10. M. attenboroughi sp. n. 11. M. boliviense sp. n. 12. M. newtoni sp. n. 13. M. foveatum sp. n. 14. M. thayerae sp. n. 15. M. proprium sp. n. 16. M. quadridens sp. n." httpUri="https://zenodo.org/record/376375/files/figure.png" pageId="17" pageNumber="250">Fig. 5</figureCitation>
. Length
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. Colour yellow brown, legs and antennae yellow. Antennomeres 311 length
<quantity id="4CB757FBE339FFE580795FA2FF7DEF0E" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="250" unit="mm" value="0.22">0.22 mm</quantity>
. Width across eyes
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. Mentum narrowed towards base, lateral margins straight. Pronotum
<quantity id="4CB757FBE339FFE580245E47FA26EF0E" box="[1334,1437,1798,1823]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="250" unit="mm" value="0.25">0.25 mm</quantity>
wide,
<quantity id="4CB757FBE339FFE585CC5E6AFEFEEF52" box="[222,325,1835,1859]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="17" pageNumber="250" unit="mm" value="0.19">0.19 mm</quantity>
long, with two distinct elongate depressions reaching from the posterior margin to just past the middle, the distance between them narrowing anteriorly; sides slightly sinuate before hind angles,
<figureCitation id="1374E69BE339FFE581EA5E0FFAF0EF77" box="[1272,1355,1870,1895]" captionStart="FIGURES 17 32" captionStartId="6.[151,264,1880,1903]" captionTargetBox="[238,1353,189,1847]" captionTargetId="figure@6.[205,1371,157,1869]" captionTargetPageId="6" captionText="FIGURES 17 32. Micridium pronotums. 17. M. quadridens sp. n. x 605. 18. M. inornatum sp. n. x 590. 19. M. boliviense sp. n. x 625. 20. M. newtoni sp. n. x 695. 21. M. foveatum sp. n. x 830. 22. M. johnsoni sp. n. x 790. 23. M. elegans sp. n. x 810. 24. M. thayerae sp. n. x 650. 25. M. halidaii (Matthews) x 770. 26. M. hilli (Johnson) x 740. 27. M. oweni sp. n. x 575. 28. M. attenboroughi sp. n. x 675. 29. M. angulicolle (Fairmaire). 30. M. rhodeanum (Casey) 31. M. vittatum Motschulsky 32. M. lineatum Le Conte." httpUri="https://zenodo.org/record/376376/files/figure.png" pageId="17" pageNumber="250">Fig. 30</figureCitation>
. Elytra
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long,
<quantity id="4CB757FBE339FFE5845E5E32FE0FEF9A" box="[332,436,1907,1931]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="250" unit="mm" value="0.3">0.30 mm</quantity>
wide, widest just in front of middle. Mesoventrum: keel narrow between mesocoxae, collar with two short ridges extending posteriorly between the keel and the mesepiventral ridges which are faint, mesepiventra without obvious reticulation
<figureCitation id="1374E69BE339FFE5876E5EFBFD74EFC3" box="[636,719,1978,2003]" captionStart="FIGURES 47 60" captionStartId="10.[151,264,1643,1666]" captionTargetBox="[163,1408,218,1606]" captionTargetId="figure@10.[151,1436,193,1622]" captionTargetPageId="10" captionText="FIGURES 47 60. Micridium mesoventra. 47. M. inornatum sp. n. x 540. 48. M. quadridens sp. n. x 710. 49. M. boliviense sp. n. x 695. 50. M. attenboroughi sp. n. x 870. 51. M. proprium sp. n. x 920. 52. M. newtoni sp. n. 53. M. johnsoni sp. n. x 655. 54. M. thayerae sp. n. x 860. 55. M. halidaii (Matthews) x 615. 56. M. oweni sp. n. x 585. 57. M. elegans sp. n. x 705. 58. M. hilli (Johnson) x 760. 59. M. foveatum sp. n. x 1300. 60. M. rhodeanum (Casey) x 545." httpUri="https://zenodo.org/record/376379/files/figure.png" pageId="17" pageNumber="250">Fig. 60</figureCitation>
. Metaventrum: metepiventral sutures faint, angled inwards and extending half way to posterior margin
<figureCitation id="1374E69BE33AFFE6877059D6FD0CE8A0" box="[610,695,151,176]" captionStart="FIGURES 61 71" captionStartId="12.[151,264,1919,1942]" captionTargetBox="[328,1250,191,1869]" captionTargetId="figure@12.[311,1262,171,1888]" captionTargetPageId="12" captionText="FIGURES 61 71. Micridium metepiventral sutures. 61. M. johnsoni sp. n. x 530. 62. M. thayerae sp. n. x 505. 63. M. attenboroughi sp. n. x 670. 64. M. foveatum sp. n. x 735. 65. M. quadridens sp. n. x 710. 66. M. elegans sp. n. x 730. 67. M. oweni sp. n. x 585. 68. M. hilli (Johnson) x 760. 69. M. halidaii (Matthews) x 615. 70. M. newtoni sp. n. x 710. 71. M. rhodeanum (Casey) x 545." httpUri="https://zenodo.org/record/376380/files/figure.png" pageId="18" pageNumber="251">Fig. 71</figureCitation>
; intercoxal process evenly rounded with comb of +/- 24 narrow setae Fig. 99. Wings of usual ptiliid
<typeStatus id="54F444BCE33AFFE6873F59FCFDDAE8C5" box="[557,609,189,213]" pageId="18" pageNumber="251">type</typeStatus>
.
</paragraph>
<paragraph id="8BF0FA1EE33AFFE685D5599EFDBAE8E8" blockId="18.[151,1436,151,536]" box="[199,513,223,248]" pageId="18" pageNumber="251">Male: aedeagus Figs. 90a/b.</paragraph>
</subSubSection>
<subSubSection id="C355A995E33AFFE685D55845FCFAEA08" pageId="18" pageNumber="251" type="discussion">
<paragraph id="8BF0FA1EE33AFFE685D55845FCDFE9BD" blockId="18.[151,1436,151,536]" pageId="18" pageNumber="251">
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<emphasis id="B93B260CE33AFFE685D55845FE80E90D" bold="true" box="[199,315,260,285]" pageId="18" pageNumber="251">Remarks.</emphasis>
Five Casey
<typeStatus id="54F444BCE33AFFE684DE5844FD95E90D" box="[460,558,261,285]" pageId="18" pageNumber="251" type="syntype">syntypes</typeStatus>
(labelled as paratypes; designated as
<typeStatus id="54F444BCE33AFFE686E35845FB28E90D" box="[1009,1171,260,285]" pageId="18" pageNumber="251" type="paralectotype">paralectotypes</typeStatus>
by Sörensson) (
<figureCitation id="1374E69BE33AFFE6805D5845FA28E90D" box="[1359,1427,260,285]" captionStart="FIGURES 1 14" captionStartId="5.[151,264,1856,1879]" captionTargetBox="[208,1396,215,1756]" captionTargetId="figure@5.[151,1436,195,1835]" captionTargetPageId="5" captionText="FIGURES 1 14. Micridium habitus. 1. M. angulicolle (Fairmaire) 2. M. halidaii (Matthews) 3. M. vittatum Motschulsky 4. M. lineatum Le Conte 5. M. rhodeanum (Casey) 6. M. hilli (Johnson) 7. M. johnsoni sp. n. 8. M. elegans sp. n. 9. M. oweni sp. n. 10. M. attenboroughi sp. n. 11. M. boliviense sp. n. 12. M. newtoni sp. n. 13. M. foveatum sp. n. 14. M. thayerae sp. n. 15. M. proprium sp. n. 16. M. quadridens sp. n." httpUri="https://zenodo.org/record/376375/files/figure.png" pageId="18" pageNumber="251">Fig. 5</figureCitation>
) were borrowed for study from the
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,
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,
<collectingCountry id="F358BA8EE33AFFE681355869FBD8E950" box="[1063,1123,296,320]" name="United States of America" pageId="18" pageNumber="251">USA</collectingCountry>
.
<location id="8E90ACC5E33AFFE681615869FB1AE950" LSID="urn:lsid:plazi:treatment:03E64B08E339FFE685855F74FCFAEA08:8E90ACC5E33AFFE681615869FB1AE950" box="[1139,1185,296,320]" country="United States of America" county="No" municipality="Euparal" name="One" pageId="18" pageNumber="251" stateProvince="Washington DC">One</location>
was remounted as a disarticulated slide in
<collectingMunicipality id="6B946064E33AFFE68482580DFE5CE975" box="[400,487,332,357]" pageId="18" pageNumber="251">Euparal</collectingMunicipality>
on a piece of acrylic attached to the pin and a male and female remounted on cards for examination in the SEM.
<collectingCounty id="62918292E33AFFE684F35831FDB8E997" box="[481,515,368,391]" pageId="18" pageNumber="251">No</collectingCounty>
genitalia were detected in the female and further dissections were not carried out on the other very fragile specimens for the reasons given above.
</materialsCitation>
</paragraph>
<paragraph id="8BF0FA1EE33AFFE685D558F9FCFAEA08" blockId="18.[151,1436,151,536]" pageId="18" pageNumber="251">
<materialsCitation id="3B27F043E33AFFE685D558F9FCFAEA08" ID-GBIF-Occurrence="1438637131" collectingDate="1953-10-30" collectorName="R. Coleman" country="United States of America" county="Sank Co." location="Fairfield Township" municipality="Johnson'" pageId="18" pageNumber="251" specimenCount="1" stateProvince="Wisconsin">
I have also seen what appear to be two further examples of this species mounted on a single card, collected by
<collectorName id="26BA9FC8E33AFFE68585589DFEAEE9E5" box="[151,277,476,501]" pageId="18" pageNumber="251">R.Coleman</collectorName>
in
<location id="8E90ACC5E33AFFE68457589DFD98E9E5" LSID="urn:lsid:plazi:treatment:03E64B08E339FFE685855F74FCFAEA08:8E90ACC5E33AFFE68457589DFD98E9E5" box="[325,547,476,501]" country="United States of America" county="Sank Co." municipality="Johnson'" name="Fairfield Township" pageId="18" pageNumber="251" stateProvince="Wisconsin">Fairfield Township</location>
,
<collectingCounty id="62918292E33AFFE68721589DFD19E9E4" box="[563,674,476,501]" pageId="18" pageNumber="251">Sank Co.</collectingCounty>
,
<collectingRegion id="498B34FCE33AFFE687A1589DFC90E9E5" box="[691,811,476,501]" country="United States of America" name="Wisconsin" pageId="18" pageNumber="251">Wisconsin</collectingRegion>
,
<collectingCountry id="F358BA8EE33AFFE6862E589DFCCCE9E4" box="[828,887,476,500]" name="United States of America" pageId="18" pageNumber="251">USA</collectingCountry>
,
<date id="FFF1DCDEE33AFFE68695589DFB9CE9E5" box="[903,1063,476,501]" pageId="18" pageNumber="251" value="1953-10-30">
<collectingDate id="EFB52536E33AFFE68695589DFB9CE9E5" box="[903,1063,476,501]" pageId="18" pageNumber="251" value="1953-10-30">30 Oct. 1953</collectingDate>
</date>
, in leaf mould and bearing the determination label '
<taxonomicName id="4C4F819DE33AFFE6846B58BEFE48EA08" box="[377,499,511,536]" class="Insecta" family="Ptiliidae" genus="Micridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="251" phylum="Arthropoda" rank="genus">Micridium</taxonomicName>
sp.
<location id="8E90ACC5E33AFFE6873358BEFDD2EA08" LSID="urn:lsid:plazi:treatment:03E64B08E339FFE685855F74FCFAEA08:8E90ACC5E33AFFE6873358BEFDD2EA08" box="[545,617,511,536]" country="United States of America" county="Sank Co." municipality="Johnson'" name="Dybas" pageId="18" pageNumber="251" stateProvince="Wisconsin">Dybas</location>
&amp;
<collectingMunicipality id="6B946064E33AFFE6879F58BEFD55EA08" box="[653,750,511,536]" pageId="18" pageNumber="251">Johnson'</collectingMunicipality>
(MM).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>