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<document id="86FA3AFBFF81251DFC11E3379752CAF9" ID-DOI="10.1080/00222930400014189" ID-ISSN="1464-5262" ID-Zenodo-Dep="5214952" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629262137941" checkinUser="felipe" docAuthor="Chen, Yifeng &amp; Chen, Yongxia" docDate="2005" docId="03E2A563FFED5B63FE7AFC44FB48FBE0" docLanguage="en" docName="JNatHist.39.19.1641-1651.pdf" docOrigin="Journal of Natural History 39 (19)" docSource="http://dx.doi.org/10.1080/00222930400014189" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Niwaella longibarba Chen &amp; Chen 2005, sp. n." docType="treatment" docUuid="1A6C7B3C-6CDC-4B62-8152-A2DDE8B5289C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="1649" masterDocId="FFDBDD1BFFE55B69FF84FFEAFFE9FFCF" masterDocTitle="Revision of the genus Niwaella in China (Pisces, Cobitidae), with description of two new species" masterLastPageNumber="1651" masterPageNumber="1641" pageNumber="1647" updateTime="1699065556629" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="F52F9C0108E5F87DB59B2DD3ABAD819C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6EB9B23DE5D7A6F9BFCF08FC25FFF55B">
<mods:title id="C9307DCA040EFD6B93114C2E50D07791">Revision of the genus Niwaella in China (Pisces, Cobitidae), with description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="91650B0278B393163322E9E99FCEC175" type="personal">
<mods:role id="331424C53D16E6E986605FDD05D0DC33">
<mods:roleTerm id="78865FC191FB03A1A8AF058FFBEA0885">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="085EC83EA1E7E518BAF259193DADF342">Chen, Yifeng</mods:namePart>
</mods:name>
<mods:name id="47226A2D6BF41153233CFBEB03B3007C" type="personal">
<mods:role id="6951617DDA19D820CD29ABA6B3013B42">
<mods:roleTerm id="92071EFAA6F47456BCFD257D8CF28419">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="917914396072DA0D33858CBAE8A0ECB1">Chen, Yongxia</mods:namePart>
</mods:name>
<mods:typeOfResource id="35DA074965F0555EB3CF9F43E41BCB81">text</mods:typeOfResource>
<mods:relatedItem id="286EE53B39B63D737738BD4D48C5F63F" type="host">
<mods:titleInfo id="FFD308B5DED4E8113F698B9793AEBE5C">
<mods:title id="87CF901A45D2B9D10134C1CBAFD6BF0E">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="3E2A8D06FBB7A7B1968288F5F59B4877">
<mods:date id="AB638D1F6853BEC080A9A8256FBE37FC">2005</mods:date>
<mods:detail id="8D3D3A83E9DD4F04C7EC9FA13BCC59E9" type="pubDate">
<mods:number id="C081984D053926DE2FF8BD4F18D604E1">2005-04-18</mods:number>
</mods:detail>
<mods:detail id="766AADF4B393E48A4C9D272FDCB77BBE" type="volume">
<mods:number id="899C4A8F8F15C3847D1BD3169567C6E2">39</mods:number>
</mods:detail>
<mods:detail id="B598E73741DB73EE77B664F691082223" type="issue">
<mods:number id="0B0D960723837A3ADA80F76B64BCD0BF">19</mods:number>
</mods:detail>
<mods:extent id="3A7B598962E94DBE2973E4624F612518" unit="page">
<mods:start id="360A368172F3ECF0BD59BDFF02CCCE21">1641</mods:start>
<mods:end id="3A85D068B27DE66A6A18CA29F32808A8">1651</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="86D8670D70CAE6020E5E7D51C97526FC">
<mods:url id="E88F9A218109AD85D1CBA3687B70FD9C">http://dx.doi.org/10.1080/00222930400014189</mods:url>
</mods:location>
<mods:classification id="CB296DB988D300F592062018D84927DF">journal article</mods:classification>
<mods:identifier id="A739A8F816B07646E26A71A40EF2C3C1" type="DOI">10.1080/00222930400014189</mods:identifier>
<mods:identifier id="85B7CE72570849020D306C18CFF6385D" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="CAF59C00EF134A54B84BF4A7321AD6A4" type="Zenodo-Dep">5214952</mods:identifier>
<mods:identifier id="A40C360A1EA7C65C2B46DF546E3CA86A" type="ZooBank">BF9465DF-9F9E-469D-BB9F-571458B59427</mods:identifier>
</mods:mods>
<treatment id="03E2A563FFED5B63FE7AFC44FB48FBE0" LSID="urn:lsid:zoobank.org:act:1A6C7B3C-6CDC-4B62-8152-A2DDE8B5289C" httpUri="http://treatment.plazi.org/id/03E2A563FFED5B63FE7AFC44FB48FBE0" lastPageId="10" lastPageNumber="1649" pageId="8" pageNumber="1647">
<subSubSection id="C35147FEFFED5B61FE7AFC44FCB8FC09" box="[510,849,942,966]" pageId="8" pageNumber="1647" type="nomenclature">
<paragraph id="8BF41475FFED5B61FE7AFC44FCB8FC09" blockId="8.[510,849,942,999]" box="[510,849,942,966]" pageId="8" pageNumber="1647">
<heading id="D0BCA319FFED5B61FE7AFC44FCB8FC09" box="[510,849,942,966]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="1647" reason="4">
<taxonomicName id="4C4B6FF6FFED5B61FE7AFC44FCEEFC09" ID-CoL="47K96" authority="Chen &amp; Chen, 2005" authorityName="Chen &amp; Chen" authorityYear="2005" box="[510,775,942,966]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="1647" phylum="Chordata" rank="species" species="longibarba" status="sp. nov.">
<emphasis id="B93FC867FFED5B61FE7AFC44FCEEFC09" bold="true" box="[510,775,942,966]" italics="true" pageId="8" pageNumber="1647">Niwaella longibarba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C751CFFED5B61FC94FC44FCB8FC09" box="[784,849,942,966]" pageId="8" pageNumber="1647" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35147FEFFED5B63FE7AFC25FB48FBE0" lastPageId="10" lastPageNumber="1649" pageId="8" pageNumber="1647" type="description">
<paragraph id="8BF41475FFED5B61FE7AFC25FCAEFC28" blockId="8.[510,849,942,999]" box="[510,839,975,999]" pageId="8" pageNumber="1647">
(
<figureCitation id="137008F0FFED5B61FD82FC25FD63FC28" box="[518,650,975,999]" captionStart="Figure 1" captionStartId="3.[125,186,1638,1657]" captionTargetBox="[330,966,832,1602]" captionTargetId="figure-266@3.[330,967,831,1602]" captionTargetPageId="3" captionText="Figure 1. Loaches of the genus Niwaella. (A) N. delicata; (B) N. multifasciata; (C) N. laterimaculata; (D) N. longibarba sp. n.; (E) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214954" httpUri="https://zenodo.org/record/5214954/files/figure.png" pageId="8" pageNumber="1647">Figures 1D</figureCitation>
,
<figureCitation id="137008F0FFED5B61FD1DFC25FD54FC28" box="[665,701,975,999]" captionStart="Figure 2" captionStartId="6.[138,198,1634,1653]" captionTargetBox="[267,1055,1234,1598]" captionTargetId="figure-354@6.[267,1056,1234,1598]" captionTargetPageId="6" captionText="Figure 2. Ventral view of the mouth with barbels and mental lobes. (A) N. laterimaculata; (B) N. longibarba sp. n.; (C) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214956" httpUri="https://zenodo.org/record/5214956/files/figure.png" pageId="8" pageNumber="1647">2B</figureCitation>
,
<figureCitation id="137008F0FFED5B61FD4EFC25FD07FC28" box="[714,750,975,999]" captionStart="Figure 3" captionStartId="7.[180,240,425,444]" captionTargetBox="[249,1046,134,389]" captionTargetId="figure-219@7.[249,1047,134,389]" captionTargetPageId="7" captionText="Figure 3. Subdorsal scales. (A) N. laterimaculata; (B) N. longibarba sp. n.; (C) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214958" httpUri="https://zenodo.org/record/5214958/files/figure.png" pageId="8" pageNumber="1647">3B</figureCitation>
,
<figureCitation id="137008F0FFED5B61FD78FC25FCC9FC28" box="[764,800,975,999]" captionStart="Figure 4" captionStartId="7.[180,240,925,944]" captionTargetBox="[254,1041,550,886]" captionTargetId="figure-196@7.[250,1047,549,889]" captionTargetPageId="7" captionText="Figure 4. Suborbital spine. (A) N. laterimaculata; (B) N. longibarba sp. n.; (C) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214960" httpUri="https://zenodo.org/record/5214960/files/figure.png" pageId="8" pageNumber="1647">4B</figureCitation>
,
<figureCitation id="137008F0FFED5B61FCAAFC25FCD4FC28" box="[814,829,975,999]" captionStart="Figure 5" captionStartId="8.[346,406,1664,1683]" captionTargetBox="[265,1057,1401,1625]" captionTargetId="figure-256@8.[262,1060,1397,1627]" captionTargetPageId="8" captionText="Figure 5. Niwaella longibarba sp. n., holotype, female, no.9607011." figureDoi="http://doi.org/10.5281/zenodo.5214962" httpUri="https://zenodo.org/record/5214962/files/figure.png" pageId="8" pageNumber="1647">5</figureCitation>
)
</paragraph>
<paragraph id="8BF41475FFED5B61FF0EFBFBFD9DFB86" blockId="8.[138,1184,1041,1097]" pageId="8" pageNumber="1647">
<taxonomicName id="4C4B6FF6FFED5B61FF0EFBFBFDDFFBE6" authority=": Son and He" authorityName="Son and He" box="[138,566,1041,1065]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="1647" phylum="Chordata" rank="species" species="laterimaculata">
<emphasis id="B93FC867FFED5B61FF0EFBFBFE7BFBE6" box="[138,402,1041,1065]" italics="true" pageId="8" pageNumber="1647">Niwaella laterimaculata</emphasis>
:
<materialsCitation id="3B231E28FFED5B61FE21FBFBFDDFFBE6" box="[421,566,1041,1065]" county="Shengxian County" location="Shengxian County" pageId="8" pageNumber="1647" specimenCount="1" stateProvince="Zhejiang">Son and He</materialsCitation>
</taxonomicName>
(not Yan and Zheng) 2001, p 1–5 (Huangzhezhen,
<collectingCounty id="62956CF9FFED5B61FF21FBDBFE68FB86" box="[165,385,1073,1097]" pageId="8" pageNumber="1647">Shengxian County</collectingCounty>
,
<collectingRegion id="498FDA97FFED5B61FE0AFBDBFD81FB86" box="[398,616,1073,1097]" country="China" name="Zhejiang" pageId="8" pageNumber="1647">Zhejiang Province</collectingRegion>
).
</paragraph>
<paragraph id="8BF41475FFED5B61FF0EFB90FEB1FB5D" blockId="8.[138,344,1146,1170]" box="[138,344,1146,1170]" pageId="8" pageNumber="1647">
<emphasis id="B93FC867FFED5B61FF0EFB90FEB1FB5D" box="[138,344,1146,1170]" italics="true" pageId="8" pageNumber="1647">Material examined</emphasis>
</paragraph>
<paragraph id="8BF41475FFED5B61FF0EFB41FEEBFACB" blockId="8.[138,1185,1195,1349]" pageId="8" pageNumber="1647">
<materialsCitation id="3B231E28FFED5B61FF0EFB41FEEBFACB" collectingDate="1996-07-10" county="Chengxian County" latitude="29.6065" location="Huangzezhen" longLatPrecision="12" longitude="120.908165" municipality="The Cao'ejiang River" pageId="8" pageNumber="1647" specimenCount="1" stateProvince="Zhejiang" typeStatus="holotype">
<typeStatus id="54F0AAD7FFED5B61FF0EFB41FF13FB0C" box="[138,250,1195,1219]" pageId="8" pageNumber="1647" type="holotype">Holotype</typeStatus>
: no. 9607011, female,
<quantity id="4CB3B990FFED5B61FD86FB41FD80FB0C" box="[514,617,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="5.77" pageId="8" pageNumber="1647" unit="mm" value="57.7">57.7 mm</quantity>
TL,
<quantity id="4CB3B990FFED5B61FD22FB41FCE4FB0C" box="[678,781,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="4.87" pageId="8" pageNumber="1647" unit="mm" value="48.7">48.7 mm</quantity>
SL. Type locality:
<collectingMunicipality id="6B908E0FFFED5B61FC69FB41FF22FB2C" pageId="8" pageNumber="1647">The Cao’ejiang River</collectingMunicipality>
,
<location id="8E9442AEFFED5B61FF5EFB21FE92FB2C" LSID="urn:lsid:plazi:treatment:03E2A563FFED5B63FE7AFC44FB48FBE0:8E9442AEFFED5B61FF5EFB21FE92FB2C" box="[218,379,1227,1251]" county="Chengxian County" latitude="29.6065" longLatPrecision="12" longitude="120.908165" municipality="The Cao'ejiang River" name="Huangzezhen" pageId="8" pageNumber="1647" stateProvince="Zhejiang">Huangzezhen</location>
,
<geoCoordinate id="EE7F72B2FFED5B61FE0DFB21FDEFFB2C" box="[393,518,1227,1251]" degrees="29" direction="north" minutes="36.39" orientation="latitude" pageId="8" pageNumber="1647" precision="9" value="29.6065">
29
<emphasis id="B93FC867FFED5B61FE23FB21FE58FB2C" bold="true" box="[423,433,1227,1251]" italics="true" pageId="8" pageNumber="1647">°</emphasis>
36.39N
</geoCoordinate>
,
<geoCoordinate id="EE7F72B2FFED5B61FD92FB21FD77FB2C" box="[534,670,1227,1251]" degrees="120" direction="east" minutes="54.49" orientation="longitude" pageId="8" pageNumber="1647" precision="9" value="120.908165">
120
<emphasis id="B93FC867FFED5B61FDC6FB21FDA5FB2C" bold="true" box="[578,588,1227,1251]" italics="true" pageId="8" pageNumber="1647">°</emphasis>
54.49E
</geoCoordinate>
,
<collectingCounty id="62956CF9FFED5B61FD29FB21FC66FB2C" box="[685,911,1227,1251]" pageId="8" pageNumber="1647">Chengxian County</collectingCounty>
,
<collectingRegion id="498FDA97FFED5B61FC19FB21FB9CFB2C" box="[925,1141,1227,1251]" country="China" name="Zhejiang" pageId="8" pageNumber="1647">Zhejiang Province</collectingRegion>
,
<date id="FFF532B5FFED5B61FB07FB21FF16FACB" pageId="8" pageNumber="1647" value="1996-07-10">
<collectingDate id="EFB1CB5DFFED5B61FB07FB21FF16FACB" pageId="8" pageNumber="1647" value="1996-07-10">10 July 1996</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF41475FFED5B61FF21FAE7FE09FA8A" blockId="8.[138,1185,1195,1349]" pageId="8" pageNumber="1647">
<typeStatus id="54F0AAD7FFED5B61FF21FAE7FEF0FAEA" box="[165,281,1293,1317]" pageId="8" pageNumber="1647" type="paratype">Paratypes</typeStatus>
: nos 9607012–55, male and female,
<quantity id="4CB3B990FFED5B61FD3BFAE7FC8FFAEA" box="[703,870,1293,1317]" metricMagnitude="-2" metricUnit="m" metricValue="6.34" metricValueMax="7.56" metricValueMin="5.12" pageId="8" pageNumber="1647" unit="mm" value="63.4" valueMax="75.6" valueMin="51.2">51.2–75.6 mm</quantity>
TL,
<quantity id="4CB3B990FFED5B61FC27FAE7FBA2FAEA" box="[931,1099,1293,1317]" metricMagnitude="-2" metricUnit="m" metricValue="5.43" metricValueMax="6.55" metricValueMin="4.31" pageId="8" pageNumber="1647" unit="mm" value="54.3" valueMax="65.5" valueMin="43.1">43.1–65.5 mm</quantity>
SL,
<specimenCount id="9D4DDFFCFFED5B61FB07FAE7FEE8FA8A" pageId="8" pageNumber="1647" type="generic">34 specimens</specimenCount>
from type locality.
</paragraph>
<caption id="DF3444FDFFED5B61FEDEF96AFC3AF95C" ID-DOI="http://doi.org/10.5281/zenodo.5214962" ID-Zenodo-Dep="5214962" box="[346,979,1664,1683]" httpUri="https://zenodo.org/record/5214962/files/figure.png" pageId="8" pageNumber="1647" startId="8.[346,406,1664,1683]" targetBox="[265,1057,1401,1625]" targetPageId="8">
<paragraph id="8BF41475FFED5B61FEDEF96AFC3AF95C" blockId="8.[346,979,1664,1683]" box="[346,979,1664,1683]" pageId="8" pageNumber="1647">
Figure 5.
<taxonomicName id="4C4B6FF6FFED5B61FE3DF96AFD80F95C" authority="Chen &amp; Chen, 2005" authorityName="Chen &amp; Chen" authorityYear="2005" box="[441,617,1664,1683]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="1647" phylum="Chordata" rank="species" species="longibarba" status="sp. nov.">
<emphasis id="B93FC867FFED5B61FE3DF96AFD80F95C" box="[441,617,1664,1683]" italics="true" pageId="8" pageNumber="1647">Niwaella longibarba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C751CFFED5B61FDF4F96AFD4CF95C" box="[624,677,1664,1683]" pageId="8" pageNumber="1647" rank="species">sp. n.</taxonomicNameLabel>
, holotype, female, no.9607011.
</paragraph>
</caption>
<paragraph id="8BF41475FFEC5B60FFF9FF62FD2CFF2F" blockId="9.[125,1172,136,256]" pageId="9" pageNumber="1648">
<emphasis id="B93FC867FFEC5B60FFF9FF62FE73FF6F" box="[125,410,136,160]" italics="true" pageId="9" pageNumber="1648">Other material examined.</emphasis>
<materialsCitation id="3B231E28FFEC5B60FE23FF62FD9CFF2F" county="Chengxian County" latitude="33.749832" location="Cao'ejiang River" longLatPrecision="11" longitude="105.768166" pageId="9" pageNumber="1648" specimenCount="1" stateProvince="Zhejiang">
Nos 471035, 471045–46, male and female, 53.4–
<quantity id="4CB3B990FFEC5B60FC73FF62FBB2FF6F" box="[1015,1115,136,160]" metricMagnitude="-2" metricUnit="m" metricValue="4.61" pageId="9" pageNumber="1648" unit="mm" value="46.1">46.1 mm</quantity>
TL, 46.7–
<quantity id="4CB3B990FFEC5B60FF45FF42FECCFF0F" box="[193,293,168,192]" metricMagnitude="-2" metricUnit="m" metricValue="3.91" pageId="9" pageNumber="1648" unit="mm" value="39.1">39.1 mm</quantity>
ST,
<specimenCount id="9D4DDFFCFFEC5B60FEF7FF42FDDCFF0F" box="[371,565,168,192]" pageId="9" pageNumber="1648" type="generic">three specimens</specimenCount>
from the
<location id="8E9442AEFFEC5B60FD40FF42FC7BFF0F" LSID="urn:lsid:plazi:treatment:03E2A563FFED5B63FE7AFC44FB48FBE0:8E9442AEFFEC5B60FD40FF42FC7BFF0F" box="[708,914,168,192]" county="Chengxian County" latitude="33.749832" longLatPrecision="11" longitude="105.768166" name="Cao'ejiang River" pageId="9" pageNumber="1648" stateProvince="Zhejiang">Cao’ejiang River</location>
,
<collectingCounty id="62956CF9FFEC5B60FC23FF42FB79FF0F" box="[935,1168,168,192]" pageId="9" pageNumber="1648">Chengxian County</collectingCounty>
,
<geoCoordinate id="EE7F72B2FFEC5B60FFF9FF22FF12FF2F" box="[125,251,200,224]" degrees="33" direction="north" minutes="44.99" orientation="latitude" pageId="9" pageNumber="1648" precision="9" value="33.749832">
33
<emphasis id="B93FC867FFEC5B60FF1FFF22FF4CFF2F" bold="true" box="[155,165,200,224]" italics="true" pageId="9" pageNumber="1648">°</emphasis>
44.99N
</geoCoordinate>
,
<geoCoordinate id="EE7F72B2FFEC5B60FE8DFF22FE78FF2F" box="[265,401,200,224]" degrees="105" direction="east" minutes="46.09" orientation="longitude" pageId="9" pageNumber="1648" precision="9" value="105.768166">
105
<emphasis id="B93FC867FFEC5B60FEB1FF22FED6FF2F" bold="true" box="[309,319,200,224]" italics="true" pageId="9" pageNumber="1648">°</emphasis>
46.09E
</geoCoordinate>
,
<collectingRegion id="498FDA97FFEC5B60FE1BFF22FD9CFF2F" box="[415,629,200,224]" country="China" name="Zhejiang" pageId="9" pageNumber="1648">Zhejiang Province</collectingRegion>
</materialsCitation>
, 1947.
</paragraph>
<paragraph id="8BF41475FFEC5B60FF1CFF02FC72FECF" blockId="9.[125,1172,136,256]" box="[152,923,232,256]" pageId="9" pageNumber="1648">
All specimens kept in FFM, IHB, CAS, Wuhan,
<collectingRegion id="498FDA97FFEC5B60FD58FF02FC71FECF" box="[732,920,232,256]" country="China" name="Hubei" pageId="9" pageNumber="1648">Hubei Province</collectingRegion>
.
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FEDAFF07FE87" blockId="9.[125,238,304,328]" box="[125,238,304,328]" pageId="9" pageNumber="1648">
<emphasis id="B93FC867FFEC5B60FFF9FEDAFF07FE87" box="[125,238,304,328]" italics="true" pageId="9" pageNumber="1648">Etymology</emphasis>
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FE8AFE8EFE57" blockId="9.[125,1172,352,408]" pageId="9" pageNumber="1648">
The specific name
<taxonomicName id="4C4B6FF6FFEC5B60FED2FE8AFE2EFEB7" authorityName="Chen &amp; Chen" authorityYear="2005" box="[342,455,352,376]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="1648" phylum="Chordata" rank="species" species="longibarba">
<emphasis id="B93FC867FFEC5B60FED2FE8AFE2EFEB7" box="[342,455,352,376]" italics="true" pageId="9" pageNumber="1648">longibarba</emphasis>
</taxonomicName>
is Latin for ‘‘with long barbel’’, and refers to the long maxillomandibular barbels.
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FE22FB7DFDDF" blockId="9.[125,245,456,476]" box="[125,1172,456,528]" lastBlockId="9.[125,1173,504,1166]" pageId="9" pageNumber="1648">
<emphasis id="B93FC867FFEC5B60FFF9FE22FF1CFE2F" box="[125,245,456,480]" italics="true" pageId="9" pageNumber="1648">Description</emphasis>
D. iii-6; A. iii-5; V. i-6; P. i-7. Vertebral 4 + 41–42. Morphometric characters are given in
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FDF2FF3EFDFF" blockId="9.[125,1173,504,1166]" box="[125,215,536,560]" pageId="9" pageNumber="1648">
<tableCitation id="C6C921CEFFEC5B60FFF9FDF2FF3BFDFF" box="[125,210,536,560]" captionStart="Table I" captionStartId="4.[138,192,979,998]" captionText="Table I. Statistical parameters for the morphometric characters of three Chinese species of Niwaella (mm)." pageId="9" pageNumber="1648">Table I</tableCitation>
.
</paragraph>
<paragraph id="8BF41475FFEC5B60FF1CFDD2FDEAFD00" blockId="9.[125,1173,504,1166]" pageId="9" pageNumber="1648">
Body elongated and laterally compressed. Dorsal surface and abdomen almost parallel. Head small, slightly compressed, snout bluntly rounded, length longer than that of space between eye and nape. Mouth small, inferior, with undeveloped mental lobes. Barbels slender, shorter than or slightly equal to eye diameter (
<figureCitation id="137008F0FFEC5B60FC81FD7DFC94FD60" box="[773,893,663,687]" captionStart="Figure 2" captionStartId="6.[138,198,1634,1653]" captionTargetBox="[267,1055,1234,1598]" captionTargetId="figure-354@6.[267,1056,1234,1598]" captionTargetPageId="6" captionText="Figure 2. Ventral view of the mouth with barbels and mental lobes. (A) N. laterimaculata; (B) N. longibarba sp. n.; (C) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214956" httpUri="https://zenodo.org/record/5214956/files/figure.png" pageId="9" pageNumber="1648">Figure 2B</figureCitation>
). Eye interorbital width equal to or longer than diameter.
</paragraph>
<paragraph id="8BF41475FFEC5B60FF1CFD3DFCD1FC01" blockId="9.[125,1173,504,1166]" pageId="9" pageNumber="1648">Dorsal fin moderately long, situated on front half of the space between nape and caudal fin, length shorter than that of head. Dorsal fin short and with tip rounded, the second branched ray the longest, not reaching one-quarter of the distance between pectoral fin and pelvic fin. Pelvic fin short and small, and approximately on the same line with the dorsal fin, reaching one-third of the distance between pelvic fin and anal fin. Anal fin short and small, situated on posterior half of the space between pelvic fin and caudal fin, tip blunt. Caudal fin long, tip emarginate. Anus close to anal fin. Caudal peduncle long, adipose crests developed. Lateral line short, exceeding length of pectorals.</paragraph>
<paragraph id="8BF41475FFEC5B60FF1CFC3CFE18FB41" blockId="9.[125,1173,504,1166]" pageId="9" pageNumber="1648">
No scales on the head and sides of head. Body scales are small, round, with a large, almost central focal area, 24–25 radial grooves and few supplementary radial grooves. The radial grooves in the front and on the side of scale are broad and sparse, at the base of scale closely spaced. The circular striae relatively scarce (
<figureCitation id="137008F0FFEC5B60FD6DFBDCFC88FB81" box="[745,865,1078,1102]" captionStart="Figure 3" captionStartId="7.[180,240,425,444]" captionTargetBox="[249,1046,134,389]" captionTargetId="figure-219@7.[249,1047,134,389]" captionTargetPageId="7" captionText="Figure 3. Subdorsal scales. (A) N. laterimaculata; (B) N. longibarba sp. n.; (C) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214958" httpUri="https://zenodo.org/record/5214958/files/figure.png" pageId="9" pageNumber="1648">Figure 3B</figureCitation>
). Suborbital spine curved opposite the caudal processes, and the mediorostral process is more developed in this species than others (
<figureCitation id="137008F0FFEC5B60FEE8FB9CFE0DFB41" box="[364,484,1142,1166]" captionStart="Figure 4" captionStartId="7.[180,240,925,944]" captionTargetBox="[254,1041,550,886]" captionTargetId="figure-196@7.[250,1047,549,889]" captionTargetPageId="7" captionText="Figure 4. Suborbital spine. (A) N. laterimaculata; (B) N. longibarba sp. n.; (C) N. xinjiangensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5214960" httpUri="https://zenodo.org/record/5214960/files/figure.png" pageId="9" pageNumber="1648">Figure 4B</figureCitation>
).
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FB54FE88FB1A" blockId="9.[125,353,1214,1238]" box="[125,353,1214,1238]" pageId="9" pageNumber="1648">
<emphasis id="B93FC867FFEC5B60FFF9FB54FE88FB1A" box="[125,353,1214,1238]" italics="true" pageId="9" pageNumber="1648">Pigmentation pattern</emphasis>
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FB04FBE7FA4A" blockId="9.[125,1172,1262,1413]" pageId="9" pageNumber="1648">Body pale yellowish. Head sprinkled with many black spots, a black stripe was visible from nape through eye to first barbel insertion. Dorsal body colour pattern 14–16 angular black stripes, moderately large, long diffuse dark vertical bar on the dorsolateral surface and 16– 19 blotches below the lateral midline. There are two or three striations on the caudal fin and dorsal fin, and a conspicuous dark spot on upper part of the caudal base.</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FA5FFEA7FA02" blockId="9.[125,334,1461,1485]" box="[125,334,1461,1485]" pageId="9" pageNumber="1648">
<emphasis id="B93FC867FFEC5B60FFF9FA5FFEA7FA02" box="[125,334,1461,1485]" italics="true" pageId="9" pageNumber="1648">Sexual dimorphism</emphasis>
</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9FA0FFEF8FA32" blockId="9.[125,273,1509,1533]" box="[125,273,1509,1533]" pageId="9" pageNumber="1648">Not evident.</paragraph>
<paragraph id="8BF41475FFEC5B60FFF9F9C7FF35F98A" blockId="9.[125,220,1581,1605]" box="[125,220,1581,1605]" pageId="9" pageNumber="1648">
<emphasis id="B93FC867FFEC5B60FFF9F9C7FF35F98A" box="[125,220,1581,1605]" italics="true" pageId="9" pageNumber="1648">Remarks</emphasis>
</paragraph>
<paragraph id="8BF41475FFEC5B63FFF9F9B7FC4DFC58" blockId="9.[125,1172,1629,1685]" lastBlockId="10.[138,1186,799,919]" lastPageId="10" lastPageNumber="1649" pageId="9" pageNumber="1648">
This new species is similar to
<taxonomicName id="4C4B6FF6FFEC5B60FE53F9B4FD71F9BA" box="[471,664,1629,1653]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="1648" phylum="Chordata" rank="species" species="laterimaculata">
<emphasis id="B93FC867FFEC5B60FE53F9B4FD71F9BA" box="[471,664,1629,1653]" italics="true" pageId="9" pageNumber="1648">N. laterimaculata</emphasis>
</taxonomicName>
in appearance, but differs from the latter in its colour pattern of a row of moderately large and long scattered dark brown vertical bars on the dorsolateral surface, versus small and densely scattered dark speckles on the dorsolateral surface, and by its two or three striations on the caudal fin, versus four or five. It is also distinguished from
<taxonomicName id="4C4B6FF6FFEF5B63FE5CFCB5FD73FCB8" box="[472,666,863,887]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="1649" phylum="Chordata" rank="species" species="laterimaculata">
<emphasis id="B93FC867FFEF5B63FE5CFCB5FD73FCB8" box="[472,666,863,887]" italics="true" pageId="10" pageNumber="1649">N. laterimaculata</emphasis>
</taxonomicName>
by its long barbels and undeveloped mental lobes, versus shorter barbels and developed mental lobes (Table II).
</paragraph>
<caption id="DF3444FDFFEF5B63FF0EFF7BFC73FF6B" box="[138,922,145,164]" pageId="10" pageNumber="1649" startId="10.[138,192,145,164]">
<paragraph id="8BF41475FFEF5B63FF0EFF7BFC73FF6B" blockId="10.[138,922,145,164]" box="[138,922,145,164]" pageId="10" pageNumber="1649">
Table II. Comparison of diagnostic characters of three Chinese species of
<taxonomicName id="4C4B6FF6FFEF5B63FCC0FF7BFC7FFF6B" authorityName="Nalbant" authorityYear="1963" box="[836,918,145,164]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="1649" phylum="Chordata" rank="genus">Niwaella</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF41475FFEF5B63FF0EFF2FFB9EFD11" pageId="10" pageNumber="1649">
<table id="F94BE6D5FFEFA496FF0EFF2FFB7FFD11" box="[138,1174,197,734]" gridcols="4" gridrows="20" pageId="10" pageNumber="1649">
<tr id="357B1637FFEFA496FF0EFF2FFB7FFF17" box="[138,1174,197,216]" gridrow="0" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFF2FFEA6FF17" box="[138,335,197,216]" gridcol="0" gridrow="0" pageId="10" pageNumber="1649">Characters</th>
<th id="76AA7F4BFFEFA496FEE3FF2FFDB3FF17" box="[359,602,197,216]" gridcol="1" gridrow="0" pageId="10" pageNumber="1649">
<taxonomicName id="4C4B6FF6FFEF5B63FE25FF2FFDD2FF17" box="[417,571,197,216]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="1649" phylum="Chordata" rank="species" species="laterimaculata">
<emphasis id="B93FC867FFEF5B63FE25FF2FFDD2FF17" box="[417,571,197,216]" italics="true" pageId="10" pageNumber="1649">N. laterimaculata</emphasis>
</taxonomicName>
</th>
<th id="76AA7F4BFFEFA496FDF1FF2FFC82FF17" box="[629,875,197,216]" gridcol="2" gridrow="0" pageId="10" pageNumber="1649">
<taxonomicName id="4C4B6FF6FFEF5B63FD2CFF2FFCC8FF17" authority="Chen &amp; Chen, 2005" authorityName="Chen &amp; Chen" authorityYear="2005" box="[680,801,197,216]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="1649" phylum="Chordata" rank="species" species="longibarba" status="sp. nov.">
<emphasis id="B93FC867FFEF5B63FD2CFF2FFCC8FF17" box="[680,801,197,216]" italics="true" pageId="10" pageNumber="1649">N. longibarba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C751CFFEF5B63FCACFF2FFCB4FF17" box="[808,861,197,216]" pageId="10" pageNumber="1649" rank="species">sp. n.</taxonomicNameLabel>
</th>
<th id="76AA7F4BFFEFA496FC0BFF2FFB7FFF17" box="[911,1174,197,216]" gridcol="3" gridrow="0" pageId="10" pageNumber="1649">
<taxonomicName id="4C4B6FF6FFEF5B63FC31FF2FFBA8FF17" authority="Chen &amp; Chen, 2005" authorityName="Chen &amp; Chen" authorityYear="2005" box="[949,1089,197,216]" class="Actinopterygii" family="Cobitidae" genus="Niwaella" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="1649" phylum="Chordata" rank="species" species="xinjiangensis" status="sp. nov.">
<emphasis id="B93FC867FFEF5B63FC31FF2FFBA8FF17" box="[949,1089,197,216]" italics="true" pageId="10" pageNumber="1649">N. xinjiangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C751CFFEF5B63FBCCFF2FFB95FF17" box="[1096,1148,197,216]" pageId="10" pageNumber="1649" rank="species">sp. n.</taxonomicNameLabel>
</th>
</tr>
<tr id="357B1637FFEFA496FF0EFF07FB7FFECF" box="[138,1174,237,256]" gridrow="1" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFF07FEA6FECF" box="[138,335,237,256]" gridcol="0" gridrow="1" pageId="10" pageNumber="1649">Caudal peduncle</th>
<td id="76AA7F4BFFEFA496FEE3FF07FDB3FECF" box="[359,602,237,256]" gridcol="1" gridrow="1" pageId="10" pageNumber="1649">Short, the ratio of depth</td>
<td id="76AA7F4BFFEFA496FDF1FF07FC82FECF" box="[629,875,237,256]" gridcol="2" gridrow="1" pageId="10" pageNumber="1649">Shorter, the ratio of depth</td>
<td id="76AA7F4BFFEFA496FC0BFF07FB7FFECF" box="[911,1174,237,256]" gridcol="3" gridrow="1" pageId="10" pageNumber="1649">Long, the ratio of depth</td>
</tr>
<tr id="357B1637FFEFA496FF0EFEEDFB7FFED5" box="[138,1174,263,282]" gridrow="2" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FEEDFDB3FED5" box="[359,602,263,282]" gridcol="1" gridrow="2" pageId="10" pageNumber="1649">to length is 0.9–1.2 (1.1)</td>
<td id="76AA7F4BFFEFA496FDF1FEEDFC82FED5" box="[629,875,263,282]" gridcol="2" gridrow="2" pageId="10" pageNumber="1649">to length is 1.1–1.8 (1.4)</td>
<td id="76AA7F4BFFEFA496FC0BFEEDFB7FFED5" box="[911,1174,263,282]" gridcol="3" gridrow="2" pageId="10" pageNumber="1649">to length is 1.2–1.7 (1.4)</td>
</tr>
<tr id="357B1637FFEFA496FF0EFEC8FB7FFEFA" box="[138,1174,290,309]" gridrow="3" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFEC8FEA6FEFA" box="[138,335,290,309]" gridcol="0" gridrow="3" pageId="10" pageNumber="1649">Mental lobes</th>
<td id="76AA7F4BFFEFA496FEE3FEC8FDB3FEFA" box="[359,602,290,309]" gridcol="1" gridrow="3" pageId="10" pageNumber="1649">Well developed</td>
<td id="76AA7F4BFFEFA496FDF1FEC8FC82FEFA" box="[629,875,290,309]" gridcol="2" gridrow="3" pageId="10" pageNumber="1649">Undeveloped</td>
<td id="76AA7F4BFFEFA496FC0BFEC8FB7FFEFA" box="[911,1174,290,309]" gridcol="3" gridrow="3" pageId="10" pageNumber="1649">Developed</td>
</tr>
<tr id="357B1637FFEFA496FF0EFED6FB7FFE80" box="[138,1174,316,335]" gridrow="4" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFED6FEA6FE80" box="[138,335,316,335]" gridcol="0" gridrow="4" pageId="10" pageNumber="1649">Maxillo-mandibular</th>
<td id="76AA7F4BFFEFA496FEE3FED6FDB3FE80" box="[359,602,316,335]" gridcol="1" gridrow="4" pageId="10" pageNumber="1649">Shorter than eye diameter</td>
<td id="76AA7F4BFFEFA496FDF1FED6FC82FE80" box="[629,875,316,335]" gridcol="2" gridrow="4" pageId="10" pageNumber="1649">Shorter or almost equal</td>
<td id="76AA7F4BFFEFA496FC0BFED6FB7FFE80" box="[911,1174,316,335]" gridcol="3" gridrow="4" pageId="10" pageNumber="1649">Shorter than eye diameter</td>
</tr>
<tr id="357B1637FFEFA496FF0EFEBDFB7FFEA5" box="[138,1174,343,362]" gridrow="5" pageId="10" pageNumber="1649" rowspan-1="1" rowspan-3="1">
<th id="76AA7F4BFFEFA496FF0EFEBDFEA6FEA5" box="[138,335,343,362]" gridcol="0" gridrow="5" pageId="10" pageNumber="1649">length</th>
<td id="76AA7F4BFFEFA496FDF1FEBDFC82FEA5" box="[629,875,343,362]" gridcol="2" gridrow="5" pageId="10" pageNumber="1649">to eye diameter</td>
</tr>
<tr id="357B1637FFEFA496FF0EFE9BFB7FFE4B" box="[138,1174,369,388]" gridrow="6" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFE9BFEA6FE4B" box="[138,335,369,388]" gridcol="0" gridrow="6" pageId="10" pageNumber="1649">Origin of dorsal fin</th>
<td id="76AA7F4BFFEFA496FEE3FE9BFDB3FE4B" box="[359,602,369,388]" gridcol="1" gridrow="6" pageId="10" pageNumber="1649">Midway between nape</td>
<td id="76AA7F4BFFEFA496FDF1FE9BFC82FE4B" box="[629,875,369,388]" gridcol="2" gridrow="6" pageId="10" pageNumber="1649">Nearer nape than</td>
<td id="76AA7F4BFFEFA496FC0BFE9BFB7FFE4B" box="[911,1174,369,388]" gridcol="3" gridrow="6" pageId="10" pageNumber="1649">Nearer nape than caudal fin</td>
</tr>
<tr id="357B1637FFEFA496FF0EFE66FB7FFE50" box="[138,1174,396,415]" gridrow="7" pageId="10" pageNumber="1649" rowspan-0="1" rowspan-3="1">
<td id="76AA7F4BFFEFA496FEE3FE66FDB3FE50" box="[359,602,396,415]" gridcol="1" gridrow="7" pageId="10" pageNumber="1649">and caudal fin</td>
<td id="76AA7F4BFFEFA496FDF1FE66FC82FE50" box="[629,875,396,415]" gridcol="2" gridrow="7" pageId="10" pageNumber="1649">caudal fin</td>
</tr>
<tr id="357B1637FFEFA496FF0EFE4DFB7FFE75" box="[138,1174,423,442]" gridrow="8" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFE4DFEA6FE75" box="[138,335,423,442]" gridcol="0" gridrow="8" pageId="10" pageNumber="1649">Scales</th>
<td id="76AA7F4BFFEFA496FEE3FE4DFDB3FE75" box="[359,602,423,442]" gridcol="1" gridrow="8" pageId="10" pageNumber="1649">Round, or oval, with a</td>
<td id="76AA7F4BFFEFA496FDF1FE4DFC82FE75" box="[629,875,423,442]" gridcol="2" gridrow="8" pageId="10" pageNumber="1649">Round, with a large,</td>
<td id="76AA7F4BFFEFA496FC0BFE4DFB7FFE75" box="[911,1174,423,442]" gridcol="3" gridrow="8" pageId="10" pageNumber="1649">Round or square, with a</td>
</tr>
<tr id="357B1637FFEFA496FF0EFE2BFB7FFE1B" box="[138,1174,449,468]" gridrow="9" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FE2BFDB3FE1B" box="[359,602,449,468]" gridcol="1" gridrow="9" pageId="10" pageNumber="1649">large, eccentric focal area,</td>
<td id="76AA7F4BFFEFA496FDF1FE2BFC82FE1B" box="[629,875,449,468]" gridcol="2" gridrow="9" pageId="10" pageNumber="1649">almost central focal area,</td>
<td id="76AA7F4BFFEFA496FC0BFE2BFB7FFE1B" box="[911,1174,449,468]" gridcol="3" gridrow="9" pageId="10" pageNumber="1649">large eccentric focal area,</td>
</tr>
<tr id="357B1637FFEFA496FF0EFE36FB7FFE20" box="[138,1174,476,495]" gridrow="10" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FE36FDB3FE20" box="[359,602,476,495]" gridcol="1" gridrow="10" pageId="10" pageNumber="1649">being closer to the base,</td>
<td id="76AA7F4BFFEFA496FDF1FE36FC82FE20" box="[629,875,476,495]" gridcol="2" gridrow="10" pageId="10" pageNumber="1649">24–25 radial grooves</td>
<td id="76AA7F4BFFEFA496FC0BFE36FB7FFE20" box="[911,1174,476,495]" gridcol="3" gridrow="10" pageId="10" pageNumber="1649">being closer to the base,</td>
</tr>
<tr id="357B1637FFEFA496FF0EFE1CFB7FFDC6" box="[138,1174,502,521]" gridrow="11" pageId="10" pageNumber="1649" rowspan-0="1" rowspan-2="1">
<td id="76AA7F4BFFEFA496FEE3FE1CFDB3FDC6" box="[359,602,502,521]" gridcol="1" gridrow="11" pageId="10" pageNumber="1649">23–26 radial grooves</td>
<td id="76AA7F4BFFEFA496FC0BFE1CFB7FFDC6" box="[911,1174,502,521]" gridcol="3" gridrow="11" pageId="10" pageNumber="1649">24–25 radial grooves,</td>
</tr>
<tr id="357B1637FFEFA496FF0EFDFBFB7FFDEB" box="[138,1174,529,548]" gridrow="12" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFDFBFEA6FDEB" box="[138,335,529,548]" gridcol="0" gridrow="12" pageId="10" pageNumber="1649">Suborbital spine</th>
<td id="76AA7F4BFFEFA496FEE3FDFBFDB3FDEB" box="[359,602,529,548]" gridcol="1" gridrow="12" pageId="10" pageNumber="1649">Curved</td>
<td id="76AA7F4BFFEFA496FDF1FDFBFC82FDEB" box="[629,875,529,548]" gridcol="2" gridrow="12" pageId="10" pageNumber="1649">More curved</td>
<td id="76AA7F4BFFEFA496FC0BFDFBFB7FFDEB" box="[911,1174,529,548]" gridcol="3" gridrow="12" pageId="10" pageNumber="1649">Slightly curved</td>
</tr>
<tr id="357B1637FFEFA496FF0EFDC1FB7FFDF1" box="[138,1174,555,574]" gridrow="13" pageId="10" pageNumber="1649">
<th id="76AA7F4BFFEFA496FF0EFDC1FEA6FDF1" box="[138,335,555,574]" gridcol="0" gridrow="13" pageId="10" pageNumber="1649">Pigmentation pattern</th>
<td id="76AA7F4BFFEFA496FEE3FDC1FDB3FDF1" box="[359,602,555,574]" gridcol="1" gridrow="13" pageId="10" pageNumber="1649">Small scattered dark</td>
<td id="76AA7F4BFFEFA496FDF1FDC1FC82FDF1" box="[629,875,555,574]" gridcol="2" gridrow="13" pageId="10" pageNumber="1649">Moderately large and long</td>
<td id="76AA7F4BFFEFA496FC0BFDC1FB7FFDF1" box="[911,1174,555,574]" gridcol="3" gridrow="13" pageId="10" pageNumber="1649">17–20 large and long dark</td>
</tr>
<tr id="357B1637FFEFA496FF0EFDACFB7FFD96" box="[138,1174,582,601]" gridrow="14" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FDACFDB3FD96" box="[359,602,582,601]" gridcol="1" gridrow="14" pageId="10" pageNumber="1649">speckles on the</td>
<td id="76AA7F4BFFEFA496FDF1FDACFC82FD96" box="[629,875,582,601]" gridcol="2" gridrow="14" pageId="10" pageNumber="1649">scattered dark vertical bar</td>
<td id="76AA7F4BFFEFA496FC0BFDACFB7FFD96" box="[911,1174,582,601]" gridcol="3" gridrow="14" pageId="10" pageNumber="1649">brown vertical bars on the</td>
</tr>
<tr id="357B1637FFEFA496FF0EFD8BFB7FFDBB" box="[138,1174,609,628]" gridrow="15" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FD8BFDB3FDBB" box="[359,602,609,628]" gridcol="1" gridrow="15" pageId="10" pageNumber="1649">dorsolateral surface and</td>
<td id="76AA7F4BFFEFA496FDF1FD8BFC82FDBB" box="[629,875,609,628]" gridcol="2" gridrow="15" pageId="10" pageNumber="1649">on the dorsolateral surface</td>
<td id="76AA7F4BFFEFA496FC0BFD8BFB7FFDBB" box="[911,1174,609,628]" gridcol="3" gridrow="15" pageId="10" pageNumber="1649">dorsolateral surface and a</td>
</tr>
<tr id="357B1637FFEFA496FF0EFD91FB7FFD41" box="[138,1174,635,654]" gridrow="16" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FD91FDB3FD41" box="[359,602,635,654]" gridcol="1" gridrow="16" pageId="10" pageNumber="1649">16–20 blotches below</td>
<td id="76AA7F4BFFEFA496FDF1FD91FC82FD41" box="[629,875,635,654]" gridcol="2" gridrow="16" pageId="10" pageNumber="1649">and 16–19 blotches below</td>
<td id="76AA7F4BFFEFA496FC0BFD91FB7FFD41" box="[911,1174,635,654]" gridcol="3" gridrow="16" pageId="10" pageNumber="1649">dark stripe along the lateral</td>
</tr>
<tr id="357B1637FFEFA496FF0EFD7CFB7FFD66" box="[138,1174,662,681]" gridrow="17" pageId="10" pageNumber="1649" rowspan-0="1">
<td id="76AA7F4BFFEFA496FEE3FD7CFDB3FD66" box="[359,602,662,681]" gridcol="1" gridrow="17" pageId="10" pageNumber="1649">the lateral midline</td>
<td id="76AA7F4BFFEFA496FDF1FD7CFC82FD66" box="[629,875,662,681]" gridcol="2" gridrow="17" pageId="10" pageNumber="1649">the lateral midline</td>
<td id="76AA7F4BFFEFA496FC0BFD7CFB7FFD66" box="[911,1174,662,681]" gridcol="3" gridrow="17" pageId="10" pageNumber="1649">midline beyond the dorsal</td>
</tr>
<tr id="357B1637FFEFA496FF0EFD5AFB7FFD0C" box="[138,1174,688,707]" gridrow="18" pageId="10" pageNumber="1649" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="76AA7F4BFFEFA496FC0BFD5AFB7FFD0C" box="[911,1174,688,707]" gridcol="3" gridrow="18" pageId="10" pageNumber="1649">fin, and then gradually</td>
</tr>
<tr id="357B1637FFEFA496FF0EFD21FB7FFD11" box="[138,1174,715,734]" gridrow="19" pageId="10" pageNumber="1649" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="76AA7F4BFFEFA496FC0BFD21FB7FFD11" box="[911,1174,715,734]" gridcol="3" gridrow="19" pageId="10" pageNumber="1649">below the lateral midline</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF41475FFEF5B63FF0EFC2DFEE2FC10" blockId="10.[138,267,967,991]" box="[138,267,967,991]" pageId="10" pageNumber="1649">
<emphasis id="B93FC867FFEF5B63FF0EFC2DFEE2FC10" box="[138,267,967,991]" italics="true" pageId="10" pageNumber="1649">Distribution</emphasis>
</paragraph>
<paragraph id="8BF41475FFEF5B63FF0EFC1DFB48FBE0" blockId="10.[138,1185,1015,1071]" pageId="10" pageNumber="1649">
Previously the species was known only in the Cao’ejiang River.
<bibRefCitation id="EFDA6984FFEF5B63FC2FFC1DFB48FBC0" author="Son YM &amp; He SP" box="[939,1185,1015,1039]" pageId="10" pageNumber="1649" pagination="1 - 5" refId="ref5327" refString="Son YM, He SP. 2001. Transfer of Cobitis laterimaculata to the genus Niwaella (Cobitidae). Korean Journal of Ichthyology 13 (1): 1 - 5." type="journal article" year="2001">Son and He (2001)</bibRefCitation>
misspelled the Cao’ejiang as the Caozejiang, and Chengxian County as Shengxian County.
</paragraph>
</subSubSection>
</treatment>
</document>
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