154 lines
16 KiB
XML
154 lines
16 KiB
XML
<document id="992A06C9F5304A2A8F63CA8E9AEBE0B1" ID-DOI="10.11646/zootaxa.3666.2.2" ID-GBIF-Dataset="930d392c-8720-4820-85eb-501985209cf3" ID-ISSN="1175-5326" ID-Zenodo-Dep="220466" ID-ZooBank="1F8D4E8A-2447-4556-9B98-D8229D858194" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460622323338" checkinUser="plazi" docAuthor="Prokin, Alexander A., Petrov, Pyotr N., Wang, Bo & Ponomarenko, Alexander G." docDate="2013" docId="03AB6D60FF9CD235FF440B8BB7374C81" docLanguage="en" docName="zt03666p159.pdf" docOrigin="Zootaxa 3666 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liadytidae Ponomarenko 1977" docType="treatment" docVersion="8" lastPageNumber="140" masterDocId="FF921518FF9FD236FFD30973B6134A15" masterDocTitle="New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)" masterLastPageNumber="159" masterPageNumber="137" pageNumber="140" updateTime="1698334052497" updateUser="plazi">
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<mods:mods id="CBB0BE232F57A4E93985072A6C10D185" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="09FF36163A96C463CF714E97CF4011B7">
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<mods:title id="C4C5EACF227D4C1C9B931A235B116266">New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="01533C2913A9D61564641AB7E2FE9935" type="personal">
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<mods:roleTerm id="8FA1AAB2C7732E5F89BDF1D888210C36">Author</mods:roleTerm>
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<mods:namePart id="761D426DA9DA3234D2A677A7BBD23415">Prokin, Alexander A.</mods:namePart>
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<mods:name id="7E3A175C2B571B6C9E963D910466DB03" type="personal">
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<mods:roleTerm id="7D1D928F2794B03FA09E684478BC306D">Author</mods:roleTerm>
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<mods:namePart id="CA66AB93437CF6EE111E2FEDF4B1F69D">Petrov, Pyotr N.</mods:namePart>
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<mods:name id="C6348AF0A3432EE09DAC2C34135D77BA" type="personal">
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<mods:namePart id="6CDF85F49245FF6E4782C6264CD93616">Wang, Bo</mods:namePart>
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</mods:name>
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<mods:name id="008DDB78736A808F20510E1036724574" type="personal">
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<mods:roleTerm id="C27C0E72A8428FD35075D2275AA74B10">Author</mods:roleTerm>
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<mods:namePart id="315B657D2BD0EB34AB20CBE9A486EF9D">Ponomarenko, Alexander G.</mods:namePart>
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<mods:typeOfResource id="3BA744A099644465C1E2DD237BCD6B46">text</mods:typeOfResource>
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<mods:title id="FE1B802806DAD33F238E1A27200F6092">Zootaxa</mods:title>
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<mods:part id="F80D8F3D2A0DB37BDD220B281B7816C3">
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<mods:date id="E05475D63EE876BC9360A1570FE6E9E0">2013</mods:date>
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<mods:detail id="A66AA14F5658DE9B99FE23BFA51B52D1" type="volume">
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<mods:number id="AAA96CD066ACC22FFD1A6B1C1FED9EDD">3666</mods:number>
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</mods:detail>
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<mods:detail id="4895185E8031A67B53FCA7CAA6AE1A7D" type="issue">
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<mods:number id="B09E874FDAC9505D2ABFE0BCE60D9AB1">2</mods:number>
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<mods:classification id="9D584FF070FF23701F542497729719E7">journal article</mods:classification>
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<mods:identifier id="16D773DDE332D4164AB327F7AEC78AC2" type="DOI">10.11646/zootaxa.3666.2.2</mods:identifier>
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<mods:identifier id="A3DEABC33010F72B0E69A73F467F2645" type="GBIF-Dataset">930d392c-8720-4820-85eb-501985209cf3</mods:identifier>
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<mods:identifier id="A05A810528BFA60C7136ED67194AC3F5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="170CEA24BD757D4931F0A373A1D44EEC" type="Zenodo-Dep">220466</mods:identifier>
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<treatment id="03AB6D60FF9CD235FF440B8BB7374C81" ID-DOI="http://doi.org/10.5281/zenodo.5680594" ID-GBIF-Taxon="119569011" ID-Zenodo-Dep="5680594" LSID="urn:lsid:plazi:treatment:03AB6D60FF9CD235FF440B8BB7374C81" httpUri="http://treatment.plazi.org/id/03AB6D60FF9CD235FF440B8BB7374C81" lastPageNumber="140" pageId="3" pageNumber="140">
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<subSubSection id="C3188FFDFF9CD235FF440B8BB46E4907" box="[151,637,760,786]" pageId="3" pageNumber="140" type="nomenclature">
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<paragraph id="8BBDDC76FF9CD235FF440B8BB46E4907" blockId="3.[151,637,760,786]" box="[151,637,760,786]" pageId="3" pageNumber="140">
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<heading id="D0F56B1AFF9CD235FF440B8BB46E4907" bold="true" box="[151,637,760,786]" fontSize="11" level="1" pageId="3" pageNumber="140" reason="1">
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<emphasis id="B9760064FF9CD235FF440B8BB46E4907" bold="true" box="[151,637,760,786]" pageId="3" pageNumber="140">
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Family
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<taxonomicName id="4C02A7F5FF9CD235FF250B8BB46E4907" authority="Ponomarenko, 1977" authorityName="Ponomarenko" authorityYear="1977" box="[246,637,760,786]" class="Insecta" family="Liadytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Liadytidae Ponomarenko, 1977</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3188FFDFF9CD235FF440A4CB7374C81" pageId="3" pageNumber="140" type="diagnosis">
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<paragraph id="8BBDDC76FF9CD235FF440A4CB5934E25" blockId="3.[151,1437,831,1684]" pageId="3" pageNumber="140">
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<emphasis id="B9760064FF9CD235FF440A4CB719494D" bold="true" box="[151,266,831,856]" pageId="3" pageNumber="140">Diagnosis</emphasis>
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(modified from Ponomarenko 1992, English translation of Ponomarenko 1977): Medium-sized aquatic beetles with oval, biconvex body. Head noticeably retraced into prothorax, scutellum visible. Mesocoxae located close to each other. Metepisterna reaching mesocoxal cavities. Metaventrite without anteromedial process protruding between mesocoxae, medially with elevated area sharply limited laterally. Anterior margin of metacoxae forming small median emargination bounded in front by extension of transverse metaventral suture. Legs slender and long. Metatibia not shorter than metafemur and metatarsus, very slender, linear. Tarsal segments not broad or paddle-shaped. Tibiae and tarsi with swimming setae.
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</paragraph>
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<paragraph id="8BBDDC76FF9CD235FF140D4FB79E4F5A" blockId="3.[151,1437,831,1684]" pageId="3" pageNumber="140">
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<emphasis id="B9760064FF9CD235FF140D4FB7194E41" bold="true" box="[199,266,1084,1108]" pageId="3" pageNumber="140">Notes</emphasis>
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: A broad lateral part of the metathoracic praeepisternum, a metathoracic anepisternum that reaches the mesocoxal cavities, and a transverse suture of the metaventrite characterize only two known families of Dytiscoidea:
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<taxonomicName id="4C02A7F5FF9CD235FEE30DF7B7BB4E89" box="[304,424,1156,1180]" class="Insecta" family="Liadytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Liadytidae</taxonomicName>
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and
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<taxonomicName id="4C02A7F5FF9CD235FE340DF7B4614E89" box="[487,626,1156,1180]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
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, the only observable difference between which listed by Balke
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<emphasis id="B9760064FF9CD235FAB30DF6B38F4E89" box="[1376,1436,1156,1180]" italics="true" pageId="3" pageNumber="140">et al.</emphasis>
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(2003) is the absence of swimming setae in
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<taxonomicName id="4C02A7F5FF9CD235FD430DD4B5084EAA" box="[656,795,1191,1215]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
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. This character is, in fact, insufficient for distinguishing between families of Dytiscoidea, because the families
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<taxonomicName id="4C02A7F5FF9CD235FCD10DBFB5604EF1" box="[770,883,1228,1252]" class="Insecta" family="Noteridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Noteridae</taxonomicName>
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,
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<taxonomicName id="4C02A7F5FF9CD235FC530DBFB5EB4EF1" box="[896,1016,1228,1252]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
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and
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<taxonomicName id="4C02A7F5FF9CD235FBE20DBFB2C04EF1" box="[1073,1235,1228,1252]" class="Insecta" family="Coptoclavidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Coptoclavidae</taxonomicName>
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include members that vary in the state of this character. The characters most clearly distinguishing
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<taxonomicName id="4C02A7F5FF9CD235FB8C0D9CB2F54F12" box="[1119,1254,1263,1287]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
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from the other Dytiscoidea, configuration of the scapus and pedicellus (Ribera
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<emphasis id="B9760064FF9CD235FC560C66B5A94F39" box="[901,954,1300,1324]" italics="true" pageId="3" pageNumber="140">et al</emphasis>
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. 2002), are not discernible in the known species of
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<taxonomicName id="4C02A7F5FF9CD235FEDE0C44B79A4F5A" box="[269,393,1335,1359]" class="Insecta" family="Liadytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Liadytidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BBDDC76FF9CD235FF1A0C2FB5C34C11" blockId="3.[151,1437,831,1684]" pageId="3" pageNumber="140">
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The new species
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<taxonomicName id="4C02A7F5FF9CD235FE5F0C2FB46D4F61" box="[396,638,1372,1396]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="aspidytoides" status="sp. nov.">
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<emphasis id="B9760064FF9CD235FE5F0C2FB46D4F61" box="[396,638,1372,1396]" italics="true" pageId="3" pageNumber="140">Liadytes aspidytoides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A245BD1FFF9CD235FD550C2FB4D54F61" box="[646,710,1372,1396]" pageId="3" pageNumber="140" rank="species">
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<emphasis id="B9760064FF9CD235FD550C2FB4D54F61" bold="true" box="[646,710,1372,1396]" pageId="3" pageNumber="140">sp. n.</emphasis>
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</taxonomicNameLabel>
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is placed here in
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<taxonomicName id="4C02A7F5FF9CD235FC5C0C2FB2194F61" box="[911,1034,1372,1396]" class="Insecta" family="Liadytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Liadytidae</taxonomicName>
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, based on the age of the fossils and on their mesocoxae located close to each other and the absence of the anteromedial metaventral process. This character distinguishes all
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<taxonomicName id="4C02A7F5FF9CD235FE160CD7B4374FA9" box="[453,548,1444,1468]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9CD235FE160CD7B4374FA9" box="[453,548,1444,1468]" italics="true" pageId="3" pageNumber="140">Liadytes</emphasis>
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</taxonomicName>
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from all
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<taxonomicName id="4C02A7F5FF9CD235FD420CD7B2A34FA9" authority="Ribera, Beutel, Balke & Vogler, 2002" authorityName="Ribera, Beutel, Balke & Vogler" authorityYear="2002" box="[657,1200,1444,1468]" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9CD235FD420CD7B4EE4FA9" box="[657,765,1444,1468]" italics="true" pageId="3" pageNumber="140">Aspidytes</emphasis>
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Ribera, Beutel, Balke & Vogler, 2002
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</taxonomicName>
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and is reported here for the first time. Since
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<taxonomicName id="4C02A7F5FF9CD235FE7D0CBBB41E4FF5" box="[430,525,1480,1504]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9CD235FE7D0CBBB41E4FF5" box="[430,525,1480,1504]" italics="true" pageId="3" pageNumber="140">Liadytes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C02A7F5FF9CD235FD9A0CBBB4A74FF5" box="[585,692,1480,1504]" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9CD235FD9A0CBBB4A74FF5" box="[585,692,1480,1504]" italics="true" pageId="3" pageNumber="140">Aspidytes</emphasis>
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</taxonomicName>
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are the single genera in the respective families, we can use this character also for distinguishing between the
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<taxonomicName id="4C02A7F5FF9CD235FD400C9FB5184C11" box="[659,779,1516,1540]" class="Insecta" family="Liadytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Liadytidae</taxonomicName>
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and
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<taxonomicName id="4C02A7F5FF9CD235FC920C9FB5DF4C11" box="[833,972,1516,1540]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BBDDC76FF9CD235FF140F63B7374C81" blockId="3.[151,1437,831,1684]" pageId="3" pageNumber="140">
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The monotypic genus
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<emphasis id="B9760064FF9CD235FE030F63B4214C3D" box="[464,562,1552,1576]" italics="true" pageId="3" pageNumber="140">Ovidytes</emphasis>
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and species
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<emphasis id="B9760064FF9CD235FD030F63B53A4C3D" box="[720,809,1552,1576]" italics="true" pageId="3" pageNumber="140">
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<collectingCountry id="F3159CE6FF9CD235FD030F63B4F44C32" box="[720,743,1552,1575]" name="Iceland" pageId="3" pageNumber="140">O</collectingCountry>
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. gaoi
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</emphasis>
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Ren, Zhu & Lu, 1995 were described in the family
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<taxonomicName id="4C02A7F5FF9CD235FF440F47B7F34C59" authority="Ren et al. 1995" authorityName="Ren et al." authorityYear="1995" box="[151,480,1588,1612]" class="Insecta" family="Liadytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">
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Liadytidae (Ren
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<emphasis id="B9760064FF9CD235FE8B0F46B7804C59" box="[344,403,1588,1612]" italics="true" pageId="3" pageNumber="140">et al.</emphasis>
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1995)
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</taxonomicName>
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. We could not study the
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<typeStatus id="54B962D4FF9CD235FCD10F46B5214C58" box="[770,818,1589,1613]" pageId="3" pageNumber="140">type</typeStatus>
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material of this species, but judging by the relatively short non-swimming legs, shown in an illustration to the original description,
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<emphasis id="B9760064FF9CD235FBED0F2BB28A4C65" box="[1086,1177,1624,1648]" italics="true" pageId="3" pageNumber="140">
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<collectingCountry id="F3159CE6FF9CD235FBED0F2BB2464C7A" box="[1086,1109,1624,1647]" name="Iceland" pageId="3" pageNumber="140">O</collectingCountry>
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. gaoi
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</emphasis>
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can hardly belong to Dytiscoidea.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |