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<mods:title id="E318AE10E2132F74FED8797A344BA50E">Revision of the Mesobuthus caucasicus complex from Central Asia with descriptions of six new species (Scorpiones Buthidae)</mods:title>
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<heading id="D0FCF6287A436A6BFF5D5CC2FCE34588" box="[222,771,1709,1736]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="29" reason="4">
<taxonomicName id="4C0B3AC77A436A6BFF5D5CC2FD444588" authority="(Birula, 1904)" baseAuthorityName="Birula" baseAuthorityYear="1904" box="[222,676,1709,1736]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="kaznakovi">
<emphasis id="B97F9D567A436A6BFF5D5CC2FE144588" bold="true" box="[222,500,1709,1736]" italics="true" pageId="30" pageNumber="29">Mesobuthus kaznakovi</emphasis>
(Birula, 1904)
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,
<emphasis id="B97F9D567A436A6BFD335CC2FD3D4587" bold="true" box="[688,733,1709,1735]" pageId="30" pageNumber="29">stat</emphasis>
.
<emphasis id="B97F9D567A436A6BFD6F5CC2FD1C4587" bold="true" box="[748,764,1709,1735]" pageId="30" pageNumber="29">n</emphasis>
.
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<paragraph id="8BB441447A436A6BFF615CA0FDD04445" blockId="30.[222,771,1709,1797]" pageId="30" pageNumber="29">
(
<figureCitation id="13305DC17A436A6BFF685CA0FE6645A6" box="[235,390,1743,1766]" captionStart-0="Figures 119122" captionStart-1="Figures 123130" captionStart-2="Figures 131138" captionStart-3="Figures 139151" captionStartId-0="30.[192,278,1470,1494]" captionStartId-1="31.[192,278,1409,1433]" captionStartId-2="32.[192,278,1757,1781]" captionStartId-3="34.[192,278,1252,1276]" captionTargetBox-0="[192,1437,144,1455]" captionTargetBox-1="[191,1438,191,1381]" captionTargetBox-2="[196,1433,208,1720]" captionTargetBox-3="[201,1434,180,1206]" captionTargetId-0="figure-11@30.[192,1437,144,1455]" captionTargetId-1="figure-6@31.[192,1438,192,1380]" captionTargetId-2="figure-11@32.[192,1439,197,1728]" captionTargetId-3="figure-11@34.[192,1439,168,1223]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionTargetPageId-3="34" captionText-0="Figures 119122: Mesobuthus kaznakovi. Figures 119120. Male from Ramit, Tajikistan, dorsal (119) and ventral (120) views. Figures 121122. Female from Ramit, Tajikistan, dorsal (121) and ventral (122) views." captionText-1="Figures 123130: Mesobuthus kaznakovi. Figures 123, 125, 127130. Male from Ramit, Tajikistan, chelicerae, carapace and tergites IIII (123), sternopectinal region and sternites IIIV (125), left legs IIV, retrolateral aspect (127130). Figures 124, 126: Paratype female, chelicerae, carapace and tergites IIII (124), and sternopectinal region and sternites IIIIV (126)." captionText-2="Figures 131138: Mesobuthus kaznakovi. Figures 131, 133135. Male from Ramit, Tajikistan, metasoma V and telson lateral (131), and metasoma and telson, lateral (133), ventral (134), and dorsal (135) views. Figures 132, 136138. Female from Ramit, Tajikistan, metasoma V and telson lateral (132), and metasoma and telson, lateral (136), ventral (137), and dorsal (138) views." captionText-3="Figures 139151: Mesobuthus kaznakovi. Figures 139141. Female from Ramit, Tajikistan, Pedipalp chela, dorsal (139) and external (140), and pedipalp patella dorsal (141). Figures 142151. Male from Ramit, Tajikistan, Pedipalp chela, dorsal (142), external (143), and ventral (144) views. Pedipalp patella, dorsal (145), external (146), and ventral (147) views. Pedipalp femur and trochanter, internal (148) and dorsal (149) views. Right chelicera dorsal (150) and ventral (151) views. The trichobothrial pattern is indicated in Figures 142146, 148149." pageId="30" pageNumber="29">Figs. 119151</figureCitation>
,
<figureCitation id="13305DC17A436A6BFE125CA0FE5945A6" box="[401,441,1743,1766]" captionStart="Figures 265273" captionStartId="56.[192,278,1569,1593]" captionTargetBox="[208,1427,176,1539]" captionTargetId="figure-49@56.[192,1439,168,1540]" captionTargetPageId="56" captionText="Figures 265273: Mesobuthus, metasoma and telson ventral of males. Figure 265. M. brutus sp. n., holotype. Figure 266. M. caucasicus, Turkey. Figure 267. M. gorelovi sp. n., holotype. Figure 268. M. fuscus, Tajikistan. Figure 269. M. kaznakovi, Tajikistan. Figure 270. M. kreuzbergi sp. n., holotype. Figure 271. M. mischi sp. n., holotype. Figure 272. M. nenilini sp. n., holotype. Figure 273. M. parthorum, Afghanistan." pageId="30" pageNumber="29">269</figureCitation>
,
<figureCitation id="13305DC17A436A6BFE455CA0FE1145A6" box="[454,497,1743,1766]" captionStart="Figures 274284" captionStartId="57.[192,278,1468,1492]" captionTargetBox="[196,1428,206,1431]" captionTargetId="figure-6@57.[192,1440,192,1439]" captionTargetPageId="57" captionText="Figures 274284: Mesobuthus, tarsomeres of third leg. Figure 274. M. brutus sp. n., male holotype. Figure 275. M. caucasicus, female from Turkey. Figure 276. M. elenae sp. n., female holotype. Figure 277. M. gorelovi sp. n., female paratype from type locality. Figure 278. M. fuscus, female from Tajikistan. Figure 279. M. kaznakovi, male from Tajikistan. Figure 280. M. kreuzbergi sp. n., female paratype. Figure 281. M. mischi sp. n., male holotype. Figure 282. M. nenilini sp. n., female paratype. Figure 283284. M. parthorum, male from Ghazni, Afghanistan (283) and female from Kholm, Afghanistan (284)." pageId="30" pageNumber="29">279</figureCitation>
,
<figureCitation id="13305DC17A436A6BFE785CA0FDB945A6" box="[507,601,1743,1766]" captionStart="Figures 285303" captionStartId="58.[192,278,1281,1305]" captionTargetBox="[198,1431,202,1246]" captionTargetId="figure-11@58.[192,1440,192,1252]" captionTargetPageId="58" captionText="Figures 285303: Mesobuthus, pedipalp chela dorsal. Figure 285286. M. brutus sp. n., male holotype (285) and female paratype (286). Figure 287288. M. caucasicus, male (287) and female (288) from Turkey. Figure 289. M. elenae sp. n., female holotype. Figure 290291. M. fuscus, differently colored male (290) and female (291) from Tajikistan. Figure 292293. M. gorelovi sp. n., male holotype (292) and female paratype (293). Figure 294295. M. kaznakovi, male (294) and female (295) from Tajikistan. Figure 296297. M. kreuzbergi sp. n., male holotype (296) and female paratype (297). Figure 298299. M. mischi sp. n., male holotype (298) and female paratype (299). Figure 300301. M. nenilini sp. n., male holotype (300) and female paratype (301). Figure 302303. M. parthorum, male (302) and female (303) from Afghanistan." pageId="30" pageNumber="29">294295</figureCitation>
,
<figureCitation id="13305DC17A436A6BFDE55CA0FD7145A6" box="[614,657,1743,1766]" captionStart="Figures 304315" captionStartId="59.[192,278,953,977]" captionTargetBox="[208,1427,193,918]" captionTargetId="figure-6@59.[192,1437,192,924]" captionTargetPageId="59" captionText="Figures 304315: Mesobuthus, pedipalp movable (304313) and fixed (314315) fingers. Figure 304. M. brutus sp. n., male paratype. Figure 305. M. caucasicus, male from Turkey. Figure 306. M. elenae sp. n., female holotype. Figure 307. M. gorelovi sp. n., male holotype. Figure 308. M. fuscus, female from Tajikistan. Figure 309. M. kaznakovi, male from Tajikistan. Figure 310, 314. M. kreuzbergi sp. n., male holotype. Figure 311. M. mischi sp. n., male holotype. Figure 312, 315. M. nenilini sp. n., male holotype. Figure 313. M. parthorum, male from Afghanistan." pageId="30" pageNumber="29">309</figureCitation>
,
<figureCitation id="13305DC17A436A6BFD185CA0FD1945A6" box="[667,761,1743,1766]" captionStart="Figures 316321" captionStartId="60.[192,278,1445,1469]" captionTargetBox="[191,1437,204,1417]" captionTargetId="figure-304@60.[192,1437,205,1416]" captionTargetPageId="60" captionText="Figures 316321: Mesobuthus, live specimens. Figure 316. M. kaznakovi, female from Uzbekistan. Figure 317. M. kaznakovi, male from Tajikistan. Figure 318. M. caucasicus, female from Iran. Figure 319. M. elenae sp. n., female. Figure 320. M. kreuzbergi sp. n., female paratype. Figure 321. M. parthorum, juvenile from Afghanistan." pageId="30" pageNumber="29">316317</figureCitation>
,
<tableCitation id="C68974FF7A436A6BFE325C81FDC74445" box="[433,551,1774,1797]" captionStart-0="Table 4" captionStart-1="Table 5" captionStart-2="Table 6" captionStartId-0="53.[192,257,1057,1081]" captionStartId-1="54.[192,257,1090,1114]" captionStartId-2="62.[192,257,967,991]" captionText-0="Table 4: Comparison among Mesobuthus species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." captionText-1="Table 5: Comparison among Mesobuhus species (specimens), based upon selected morphometric ratios of adult females. Abbreviations: length (L), width (W), depth (D)." captionText-2="Table 6: Comparison among Mesobuhus species, based upon pectinal teeth number. Data from selected specimens studied in this revision; additional data for M. caucasicus (Caucasus) from ZISP collection (Fet, 1989) scored by VF in 1987-88." pageId="30" pageNumber="29">Tables 46</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31112CF7A436A6BFF435D47FDA4447F" box="[192,580,1832,1855]" pageId="30" pageNumber="29" type="reference_group">
<paragraph id="8BB441447A436A6BFF435D47FDA4447F" blockId="30.[192,802,1832,1916]" box="[192,580,1832,1855]" pageId="30" pageNumber="29">
<taxonomicName id="4C0B3AC77A436A6BFF435D47FDDE447F" authority="Birula, 1904 b: 32" authorityName="Birula" authorityPageNumber="32" authorityYear="1904" box="[192,574,1832,1855]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="kaznakovi">
<emphasis id="B97F9D567A436A6BFF435D47FE9B447F" box="[192,379,1832,1855]" italics="true" pageId="30" pageNumber="29">Buthus kaznakovi</emphasis>
Birula, 1904b: 32
</taxonomicName>
.
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<typeStatus id="54B0FFE67A436A6BFE405D25FE18441D" box="[451,504,1866,1885]" pageId="30" pageNumber="29">TYPE</typeStatus>
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<collectingCountry id="F31C01D47A436A6BFD255D29FCFB441E" box="[678,795,1862,1886]" name="Tajikistan" pageId="30" pageNumber="29">Tajikistan</collectingCountry>
</emphasis>
,
<emphasis id="B97F9D567A436A6BFF435D0AFE9D443C" box="[192,381,1893,1916]" italics="true" pageId="30" pageNumber="29">
<collectingRegion id="49CF8FA67A436A6BFF435D0AFE9D443C" box="[192,381,1893,1916]" country="Tajikistan" name="Khatlon" pageId="30" pageNumber="29">Khatlon Province</collectingRegion>
</emphasis>
, road from
<collectingMunicipality id="6BD0DB3E7A436A6BFE785D0AFDB7443C" box="[507,599,1893,1916]" pageId="30" pageNumber="29">Sarypul</collectingMunicipality>
to
<location id="8ED4179F7A436A6BFDFA5D0AFD22443C" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A436A6BFDFA5D0AFD22443C" box="[633,706,1893,1916]" country="Tajikistan" municipality="Sarypul'" name="Talbar" pageId="30" pageNumber="29" stateProvince="Khatlon">Talbar</location>
</materialsCitation>
;
<collectionCode id="ED1AD9817A436A6BFD4E5D0AFCE4443C" box="[717,772,1893,1916]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="30" pageNumber="29">ZISP</collectionCode>
.
</paragraph>
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<subSubSection id="C31112CF7A436A6BFCBD5C7BFBC445C7" pageId="30" pageNumber="29" type="reference_group">
<paragraph id="8BB441447A436A6BFCBD5C7BFB404509" blockId="30.[830,1442,1556,1671]" pageId="30" pageNumber="29">
REFERENCES (selected); see
<bibRefCitation id="EF9A3CB57A436A6BFBF75C7BFB06456B" author="FET" box="[1140,1254,1556,1579]" pageId="30" pageNumber="29" pagination="73 - 171" refId="ref33201" refString="FET, V. 1989. A catalogue of scorpions (Chelicerata: Scorpiones) of the USSR. Rivista del Museo Civico di Scienze Naturali &quot; Enrico Caffi &quot;, 13 (1988): 73 - 171." type="journal article" year="1989">Fet (1989)</bibRefCitation>
and Fet &amp; Lowe (2000) for a full list before 1998:
</paragraph>
<paragraph id="8BB441447A436A6BFCED5C3EFA404528" blockId="30.[830,1442,1556,1671]" box="[878,1440,1617,1640]" pageId="30" pageNumber="29">
<taxonomicName id="4C0B3AC77A436A6BFCED5C3EFA794528" authority=": Birula, 1911: 167" authorityName="Birula" authorityPageNumber="167" authorityYear="1911" box="[878,1433,1617,1640]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subSpecies="kaznakovi">
<emphasis id="B97F9D567A436A6BFCED5C3EFB564528" box="[878,1206,1617,1640]" italics="true" pageId="30" pageNumber="29">Buthus caucasicus kaznakovi</emphasis>
: Birula, 1911: 167
</taxonomicName>
;
</paragraph>
<paragraph id="8BB441447A436A6BFCED5C1FFBC445C7" blockId="30.[830,1442,1556,1671]" box="[878,1060,1648,1671]" pageId="30" pageNumber="29">Birula, 1917: 71.</paragraph>
</subSubSection>
<subSubSection id="C31112CF7A436A6BFCED5CC2FBA645A3" pageId="30" pageNumber="29" type="description">
<paragraph id="8BB441447A436A6BFCED5CC2FA404584" blockId="30.[832,1440,1709,1824]" box="[878,1440,1709,1732]" pageId="30" pageNumber="29">
<taxonomicName id="4C0B3AC77A436A6BFCED5CC2FA404584" authority=": Vachon, 1958: 150" authorityName=": Vachon" authorityYear="1958" box="[878,1440,1709,1732]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subSpecies="kaznakovi">
<emphasis id="B97F9D567A436A6BFCED5CC2FB094584" box="[878,1257,1709,1732]" italics="true" pageId="30" pageNumber="29">Mesobuthus caucasicus kaznakovi</emphasis>
:
<bibRefCitation id="EF9A3CB57A436A6BFB7E5CC2FA404584" author="VACHON" box="[1277,1440,1709,1732]" pageId="30" pageNumber="29" pagination="121 - 187" refId="ref35395" refString="VACHON, M. 1958. Scorpionidea (Chelicerata) de l'Afghanistan. (The 3 d Danish Expedition to Central Asia. Zoological Results 23). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KObehavn, 120: 121 - 187." type="journal article" year="1958">Vachon, 1958:</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BB441447A436A6BFCED5CA3FBA645A3" blockId="30.[832,1440,1709,1824]" box="[878,1094,1740,1763]" pageId="30" pageNumber="29">
150;
<bibRefCitation id="EF9A3CB57A436A6BFC265CA3FBDF45A3" author="FET" box="[933,1087,1740,1763]" pageId="30" pageNumber="29" pagination="73 - 171" refId="ref33201" refString="FET, V. 1989. A catalogue of scorpions (Chelicerata: Scorpiones) of the USSR. Rivista del Museo Civico di Scienze Naturali &quot; Enrico Caffi &quot;, 13 (1988): 73 - 171." type="journal article" year="1989">Fet, 1989: 112</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31112CF7A436A6BFCED5C85FC8E4460" pageId="30" pageNumber="29" type="reference_group">
<paragraph id="8BB441447A436A6BFCED5C85FC8E4460" blockId="30.[832,1440,1709,1824]" pageId="30" pageNumber="29">
<taxonomicName id="4C0B3AC77A436A6BFCED5C85FC8A4460" ID-CoL="498SH" authority=": Fet &amp; Lowe, 2000: 191" authorityName="Fet &amp; Lowe" authorityPageNumber="191" authorityYear="2000" class="Arachnida" family="Buthidae" genus="Olivierus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subSpecies="kaznakovi">
<emphasis id="B97F9D567A436A6BFCED5C85FB214441" box="[878,1217,1770,1793]" italics="true" pageId="30" pageNumber="29">Olivierus caucasicus kaznakovi</emphasis>
: Fet &amp; Lowe, 2000: 191
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31112CF7A436A6AFCBD5D29FBEC45D1" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29" type="materials_examined">
<paragraph id="8BB441447A436A6BFCBD5D29FA76443C" blockId="30.[830,1440,1862,1916]" pageId="30" pageNumber="29">
<materialsCitation id="3B634B197A436A6BFCBD5D29FA76443C" collectionCode="TYPE, MATERIAL" country="Tajikistan" latitude="38.42" location="Talbar" longLatPrecision="705" longitude="70.13" municipality="Sarypul'" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Khatlon" typeStatus="syntype">
<collectionCode id="ED1AD9817A436A6BFCBD5D29FC11441D" box="[830,1009,1862,1885]" pageId="30" pageNumber="29">TYPE MATERIAL</collectionCode>
.
<emphasis id="B97F9D567A436A6BFB865D29FB8B441D" box="[1029,1131,1862,1885]" italics="true" pageId="30" pageNumber="29">
<typeStatus id="54B0FFE67A436A6BFB865D29FB82441D" box="[1029,1122,1862,1885]" pageId="30" pageNumber="29" type="syntype">Syntypes</typeStatus>
:
</emphasis>
<emphasis id="B97F9D567A436A6BFBFB5D29FB0D441E" bold="true" box="[1144,1261,1862,1886]" pageId="30" pageNumber="29">
<collectingCountry id="F31C01D47A436A6BFBFB5D29FB0D441E" box="[1144,1261,1862,1886]" name="Tajikistan" pageId="30" pageNumber="29">Tajikistan</collectingCountry>
</emphasis>
,
<emphasis id="B97F9D567A436A6BFA825D29FC8A443C" italics="true" pageId="30" pageNumber="29">
<collectingRegion id="49CF8FA67A436A6BFA825D29FC8A443C" country="Tajikistan" name="Khatlon" pageId="30" pageNumber="29">Khatlon Province</collectingRegion>
</emphasis>
, road from
<collectingMunicipality id="6BD0DB3E7A436A6BFC6A5D0AFBA5443C" box="[1001,1093,1893,1916]" pageId="30" pageNumber="29">Sarypul</collectingMunicipality>
to
<location id="8ED4179F7A436A6BFBE45D0AFB4F443C" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A436A6BFBE45D0AFB4F443C" box="[1127,1199,1893,1916]" country="Tajikistan" latitude="38.42" longLatPrecision="705" longitude="70.13" municipality="Sarypul'" name="Talbar" pageId="30" pageNumber="29" stateProvince="Khatlon">Talbar</location>
, [
<geoCoordinate id="EE3F27837A436A6BFB415D0AFAFC443C" box="[1218,1308,1893,1916]" degrees="38. 42" direction="north" orientation="latitude" pageId="30" pageNumber="29" precision="555" value="38.42">38.42°N</geoCoordinate>
<geoCoordinate id="EE3F27837A436A6BFAA05D0AFA9A443C" box="[1315,1402,1893,1916]" degrees="70.13" direction="east" orientation="longitude" pageId="30" pageNumber="29" precision="555" value="70.13">70.13°E</geoCoordinate>
to
</materialsCitation>
</paragraph>
<caption id="DF7411CC7A426A6AFF435FEEFADC468B" pageId="31" pageNumber="30" startId="31.[192,278,1409,1433]" targetBox="[191,1438,191,1381]" targetPageId="31">
<paragraph id="8BB441447A426A6AFF435FEEFADC468B" blockId="31.[192,1441,1409,1483]" pageId="31" pageNumber="30">
<emphasis id="B97F9D567A426A6AFF435FEEFE6446DA" bold="true" box="[192,388,1409,1434]" pageId="31" pageNumber="30">Figures 123130:</emphasis>
<taxonomicName id="4C0B3AC77A426A6AFE0F5FEBFD8746D9" baseAuthorityName="Birula" baseAuthorityYear="1904" box="[396,615,1412,1433]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="kaznakovi">
<emphasis id="B97F9D567A426A6AFE0F5FEBFD8746D9" box="[396,615,1412,1433]" italics="true" pageId="31" pageNumber="30">Mesobuthus kaznakovi</emphasis>
</taxonomicName>
.
<emphasis id="B97F9D567A426A6AFDF65FECFD0E46D8" bold="true" box="[629,750,1411,1433]" pageId="31" pageNumber="30">Figures 123</emphasis>
,
<emphasis id="B97F9D567A426A6AFD785FECFCFF46D8" bold="true" box="[763,799,1411,1432]" pageId="31" pageNumber="30">125</emphasis>
,
<emphasis id="B97F9D567A426A6AFCAE5FECFC6146D8" bold="true" box="[813,897,1411,1433]" pageId="31" pageNumber="30">127130</emphasis>
. Male from Ramit, Tajikistan, chelicerae, carapace and tergites IIII (123), sternopectinal region and sternites IIIV (125), left legs IIV, retrolateral aspect (127130).
<emphasis id="B97F9D567A426A6AFAA35FF3FA7A46F1" bold="true" box="[1312,1434,1436,1458]" pageId="31" pageNumber="30">Figures 124</emphasis>
,
<emphasis id="B97F9D567A426A6AFF435FDAFF04468A" bold="true" box="[192,228,1461,1482]" pageId="31" pageNumber="30">126</emphasis>
: Paratype female, chelicerae, carapace and tergites IIII (124), and sternopectinal region and sternites IIIIV (126).
</paragraph>
</caption>
<paragraph id="8BB441447A426A6AFF435C6FFD6D444C" blockId="31.[192,802,1536,1804]" pageId="31" pageNumber="30">
<materialsCitation id="3B634B197A426A6AFF435C6FFDD64575" collectingDate="1897-05-28" collectingDateMax="1897-05-29" collectingDateMin="1897-05-28" collectionCode="ZISP" collectorName="A. N. Kaznakov" latitude="38.6" location="Old Style" longLatPrecision="705" longitude="70.35" pageId="31" pageNumber="30" specimenCode="ZISP 1252" specimenCount="1" specimenCount-female="1">
<geoCoordinate id="EE3F27837A426A6AFF435C6FFEFA4557" box="[192,282,1536,1559]" degrees="38.60" direction="north" orientation="latitude" pageId="31" pageNumber="30" precision="555" value="38.6">38.60°N</geoCoordinate>
<geoCoordinate id="EE3F27837A426A6AFEA65C6FFE9C4557" box="[293,380,1536,1559]" degrees="70.35" direction="east" orientation="longitude" pageId="31" pageNumber="30" precision="555" value="70.35">70.35°E</geoCoordinate>
]
<date id="FFB567847A426A6AFE085C6FFDA54557" box="[395,581,1536,1559]" pageId="31" pageNumber="30" value="1897-05-28" valueMax="1897-05-29" valueMin="1897-05-28">
<collectingDate id="EFF19E6C7A426A6AFE085C6FFDA54557" box="[395,581,1536,1559]" pageId="31" pageNumber="30" value="1897-05-28" valueMax="1897-05-29" valueMin="1897-05-28">2829 May 1897</collectingDate>
</date>
(
<location id="8ED4179F7A426A6AFDDB5C6FFD224557" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFDDB5C6FFD224557" box="[600,706,1536,1559]" latitude="38.6" longLatPrecision="705" longitude="70.35" name="Old Style" pageId="31" pageNumber="30">Old Style</location>
), leg.
<collectorName id="26FE24927A426A6AFC8B5C6FFEB84575" pageId="31" pageNumber="30">A. N. Kaznakov</collectorName>
,
<specimenCount id="9D0D8ACD7A426A6AFEF25C71FE724575" box="[369,402,1566,1589]" pageId="31" pageNumber="30" type="female">1♀</specimenCount>
(
<specimenCode id="DBADE93F7A426A6AFE285C71FDCB4575" box="[427,555,1566,1589]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="31" pageNumber="30">ZISP 1252</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B634B197A426A6AFDCE5C71FDA74514" location="Tavildara District" municipality="Obi-Khingou River" pageId="31" pageNumber="30" specimenCount="1">
<emphasis id="B97F9D567A426A6AFDCE5C71FCFB4575" box="[589,795,1566,1589]" italics="true" pageId="31" pageNumber="30">
<location id="8ED4179F7A426A6AFDCE5C71FCFB4575" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFDCE5C71FCFB4575" box="[589,795,1566,1589]" municipality="Obi-Khingou River" name="Tavildara District" pageId="31" pageNumber="30">Tavildara District</location>
</emphasis>
: Childara (on
<collectingMunicipality id="6BD0DB3E7A426A6AFEE65C52FDDD4514" box="[357,573,1597,1620]" pageId="31" pageNumber="30">Obi-Khingou River</collectingMunicipality>
)
</materialsCitation>
,
<materialsCitation id="3B634B197A426A6AFDDF5C52FE3745D1" collectingDate="1897-05-30" collectingDateMax="1897-06-17" collectingDateMin="1897-05-30" collectionCode="ZISP" collectorName="A. N. Kaznakov" latitude="38.78" location="Old Style" longLatPrecision="704" longitude="70.3" pageId="31" pageNumber="30" specimenCode="ZISP 1254" specimenCount="1" specimenCount-female="1">
<geoCoordinate id="EE3F27837A426A6AFDDF5C52FD554514" box="[604,693,1597,1620]" degrees="38.78" direction="north" orientation="latitude" pageId="31" pageNumber="30" precision="555" value="38.78">38.78°N</geoCoordinate>
<geoCoordinate id="EE3F27837A426A6AFD475C52FCFE4514" box="[708,798,1597,1620]" degrees="70.30" direction="east" orientation="longitude" pageId="31" pageNumber="30" precision="555" value="70.3">70.30°E</geoCoordinate>
, leg.
<collectorName id="26FE24927A426A6AFF705C33FE494533" box="[243,425,1628,1651]" pageId="31" pageNumber="30">A. N. Kaznakov</collectorName>
,
<date id="FFB567847A426A6AFE385C33FD054533" box="[443,741,1628,1651]" pageId="31" pageNumber="30" value="1897-05-30" valueMax="1897-06-17" valueMin="1897-05-30">
<collectingDate id="EFF19E6C7A426A6AFE385C33FD054533" box="[443,741,1628,1651]" pageId="31" pageNumber="30" value="1897-05-30" valueMax="1897-06-17" valueMin="1897-05-30">30 May and 17 June 1897</collectingDate>
</date>
(
<location id="8ED4179F7A426A6AFD795C33FF1B45D1" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFD795C33FF1B45D1" latitude="38.78" longLatPrecision="704" longitude="70.3" name="Old Style" pageId="31" pageNumber="30">Old Style</location>
),
<specimenCount id="9D0D8ACD7A426A6AFE945C15FED845D1" box="[279,312,1658,1681]" pageId="31" pageNumber="30" type="female">1♀</specimenCount>
(
<specimenCode id="DBADE93F7A426A6AFED35C15FE2E45D1" box="[336,462,1658,1681]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="31" pageNumber="30">ZISP 1254</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B634B197A426A6AFE6C5C15FDCE458F" collectingDate="1897-05-01" collectingDateMax="1897-05-03" collectingDateMin="1897-05-01" collectionCode="ZISP" collectorName="Amudarya &amp; A. N. Kaznakov" location="Karasu River" municipality="Old Style" pageId="31" pageNumber="30" specimenCount="2" specimenCount-female="2">
<location id="8ED4179F7A426A6AFE6C5C15FD6645D1" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFE6C5C15FD6645D1" box="[495,646,1658,1681]" municipality="Old Style" name="Karasu River" pageId="31" pageNumber="30">Karasu River</location>
(tributary of
<collectorName id="26FE24927A426A6AFF435CF6FED545F0" box="[192,309,1689,1712]" pageId="31" pageNumber="30">Amudarya</collectorName>
), leg.
<collectorName id="26FE24927A426A6AFEFE5CF6FDD245F0" box="[381,562,1689,1712]" pageId="31" pageNumber="30">A. N. Kaznakov</collectorName>
,
<date id="FFB567847A426A6AFDC05CF6FD0645F0" box="[579,742,1689,1712]" pageId="31" pageNumber="30" value="1897-05-01" valueMax="1897-05-03" valueMin="1897-05-01">
<collectingDate id="EFF19E6C7A426A6AFDC05CF6FD0645F0" box="[579,742,1689,1712]" pageId="31" pageNumber="30" value="1897-05-01" valueMax="1897-05-03" valueMin="1897-05-01">13 May 1897</collectingDate>
</date>
(
<collectingMunicipality id="6BD0DB3E7A426A6AFD7B5CF6FF1B458F" pageId="31" pageNumber="30">Old Style</collectingMunicipality>
),
<specimenCount id="9D0D8ACD7A426A6AFE905CD7FEA0458F" box="[275,320,1720,1743]" pageId="31" pageNumber="30" type="female">2 ♀</specimenCount>
(not found in
<collectionCode id="ED1AD9817A426A6AFE6D5CD7FDC6458F" box="[494,550,1720,1743]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="31" pageNumber="30">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B634B197A426A6AFDC25CD7FD69444C" collectingDate="1897-08" collectionCode="ZISP" collectorName="Shugnan &amp; A. N. Kaznakov" location="Shakhdara River valley" pageId="31" pageNumber="30" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis id="B97F9D567A426A6AFDC25CD7FD7E458F" box="[577,670,1720,1743]" italics="true" pageId="31" pageNumber="30">
<collectorName id="26FE24927A426A6AFDC25CD7FD7E458F" box="[577,670,1720,1743]" pageId="31" pageNumber="30">Shugnan</collectorName>
</emphasis>
,
<location id="8ED4179F7A426A6AFD325CD7FEA745AD" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFD325CD7FEA745AD" name="Shakhdara River valley" pageId="31" pageNumber="30">Shakhdara River valley</location>
,
<date id="FFB567847A426A6AFED75CB9FE0145AD" box="[340,481,1750,1773]" pageId="31" pageNumber="30" value="1897-08">
<collectingDate id="EFF19E6C7A426A6AFED75CB9FE0145AD" box="[340,481,1750,1773]" pageId="31" pageNumber="30" value="1897-08">August 1897</collectingDate>
</date>
, leg.
<collectorName id="26FE24927A426A6AFD9C5CB9FD2E45AD" box="[543,718,1750,1773]" pageId="31" pageNumber="30">A. N. Kaznakov</collectorName>
, “numerous
<specimenCount id="9D0D8ACD7A426A6AFE825C9AFEF5444C" box="[257,277,1781,1804]" pageId="31" pageNumber="30" type="female"></specimenCount>
,
<specimenCount id="9D0D8ACD7A426A6AFEA15C9AFED6444C" box="[290,310,1781,1804]" pageId="31" pageNumber="30" type="male"></specimenCount>
, and pulli” (not found in
<collectionCode id="ED1AD9817A426A6AFDC55C9AFD61444C" box="[582,641,1781,1804]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="31" pageNumber="30">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB441447A426A6AFF435D5DFBEC45D1" blockId="31.[192,802,1841,1896]" lastBlockId="31.[830,1441,1536,1681]" pageId="31" pageNumber="30">
<collectionCode id="ED1AD9817A426A6AFF435D5DFE424409" box="[192,418,1842,1865]" pageId="31" pageNumber="30">MATERIAL STUDIED</collectionCode>
.
<materialsCitation id="3B634B197A426A6AFE3D5D5EFA794575" collectingDate="1962-06-27" collectionCode="FKCP" country="Tajikistan" latitude="38.78" location="Sardai-Miena River valley" longLatPrecision="704" longitude="69.35" municipality="Ramit Nature Reserve" pageId="31" pageNumber="30" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Khatlon">
<emphasis id="B97F9D567A426A6AFE3D5D5EFDD34409" bold="true" box="[446,563,1841,1865]" pageId="31" pageNumber="30">
<collectingCountry id="F31C01D47A426A6AFE3D5D5EFDD34409" box="[446,563,1841,1865]" name="Tajikistan" pageId="31" pageNumber="30">Tajikistan</collectingCountry>
</emphasis>
,
<emphasis id="B97F9D567A426A6AFDD35D5DFCFB4409" box="[592,795,1842,1865]" italics="true" pageId="31" pageNumber="30">
<collectingRegion id="49CF8FA67A426A6AFDD35D5DFCFB4409" box="[592,795,1842,1865]" country="Tajikistan" name="Khatlon" pageId="31" pageNumber="30">Khatlon Province</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BD0DB3E7A426A6AFF435D3EFE4D4428" box="[192,429,1873,1896]" pageId="31" pageNumber="30">Ramit Nature Reserve</collectingMunicipality>
(zapovednik),
<location id="8ED4179F7A426A6AFDCD5D3EFC624557" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFDCD5D3EFC624557" box="[590,898,1536,1896]" country="Tajikistan" latitude="38.78" longLatPrecision="704" longitude="69.35" municipality="Ramit Nature Reserve" name="Sardai-Miena River valley" pageId="31" pageNumber="30" stateProvince="Khatlon">Sardai-Miena River valley</location>
,
<geoCoordinate id="EE3F27837A426A6AFC105C6FFC0D4557" box="[915,1005,1536,1559]" degrees="38.78" direction="north" orientation="latitude" pageId="31" pageNumber="30" precision="555" value="38.78">38.78°N</geoCoordinate>
<geoCoordinate id="EE3F27837A426A6AFC795C6FFBB44557" box="[1018,1108,1536,1559]" degrees="69.35" direction="east" orientation="longitude" pageId="31" pageNumber="30" precision="555" value="69.35">69.35°E</geoCoordinate>
,
<date id="FFB567847A426A6AFBE75C6FFAE24557" box="[1124,1282,1536,1559]" pageId="31" pageNumber="30" value="1962-06-27">
<collectingDate id="EFF19E6C7A426A6AFBE75C6FFAE24557" box="[1124,1282,1536,1559]" pageId="31" pageNumber="30" value="1962-06-27">27 June 1962</collectingDate>
</date>
,
<specimenCount id="9D0D8ACD7A426A6AFA905C6FFAD44557" box="[1299,1332,1536,1559]" pageId="31" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="9D0D8ACD7A426A6AFAB75C6FFAB64557" box="[1332,1366,1536,1559]" pageId="31" pageNumber="30" type="female">1♀</specimenCount>
(
<figureCitation id="13305DC17A426A6AFAE95C6FFC7C4575" captionStart-0="Figures 119122" captionStart-1="Figures 123130" captionStart-2="Figures 131138" captionStart-3="Figures 139151" captionStartId-0="30.[192,278,1470,1494]" captionStartId-1="31.[192,278,1409,1433]" captionStartId-2="32.[192,278,1757,1781]" captionStartId-3="34.[192,278,1252,1276]" captionTargetBox-0="[192,1437,144,1455]" captionTargetBox-1="[191,1438,191,1381]" captionTargetBox-2="[196,1433,208,1720]" captionTargetBox-3="[201,1434,180,1206]" captionTargetId-0="figure-11@30.[192,1437,144,1455]" captionTargetId-1="figure-6@31.[192,1438,192,1380]" captionTargetId-2="figure-11@32.[192,1439,197,1728]" captionTargetId-3="figure-11@34.[192,1439,168,1223]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionTargetPageId-3="34" captionText-0="Figures 119122: Mesobuthus kaznakovi. Figures 119120. Male from Ramit, Tajikistan, dorsal (119) and ventral (120) views. Figures 121122. Female from Ramit, Tajikistan, dorsal (121) and ventral (122) views." captionText-1="Figures 123130: Mesobuthus kaznakovi. Figures 123, 125, 127130. Male from Ramit, Tajikistan, chelicerae, carapace and tergites IIII (123), sternopectinal region and sternites IIIV (125), left legs IIV, retrolateral aspect (127130). Figures 124, 126: Paratype female, chelicerae, carapace and tergites IIII (124), and sternopectinal region and sternites IIIIV (126)." captionText-2="Figures 131138: Mesobuthus kaznakovi. Figures 131, 133135. Male from Ramit, Tajikistan, metasoma V and telson lateral (131), and metasoma and telson, lateral (133), ventral (134), and dorsal (135) views. Figures 132, 136138. Female from Ramit, Tajikistan, metasoma V and telson lateral (132), and metasoma and telson, lateral (136), ventral (137), and dorsal (138) views." captionText-3="Figures 139151: Mesobuthus kaznakovi. Figures 139141. Female from Ramit, Tajikistan, Pedipalp chela, dorsal (139) and external (140), and pedipalp patella dorsal (141). Figures 142151. Male from Ramit, Tajikistan, Pedipalp chela, dorsal (142), external (143), and ventral (144) views. Pedipalp patella, dorsal (145), external (146), and ventral (147) views. Pedipalp femur and trochanter, internal (148) and dorsal (149) views. Right chelicera dorsal (150) and ventral (151) views. The trichobothrial pattern is indicated in Figures 142146, 148149." pageId="31" pageNumber="30">Figs. 119151</figureCitation>
,
<figureCitation id="13305DC17A426A6AFC3B5C71FC034575" box="[952,995,1566,1589]" captionStart="Figures 265273" captionStartId="56.[192,278,1569,1593]" captionTargetBox="[208,1427,176,1539]" captionTargetId="figure-49@56.[192,1439,168,1540]" captionTargetPageId="56" captionText="Figures 265273: Mesobuthus, metasoma and telson ventral of males. Figure 265. M. brutus sp. n., holotype. Figure 266. M. caucasicus, Turkey. Figure 267. M. gorelovi sp. n., holotype. Figure 268. M. fuscus, Tajikistan. Figure 269. M. kaznakovi, Tajikistan. Figure 270. M. kreuzbergi sp. n., holotype. Figure 271. M. mischi sp. n., holotype. Figure 272. M. nenilini sp. n., holotype. Figure 273. M. parthorum, Afghanistan." pageId="31" pageNumber="30">269</figureCitation>
,
<figureCitation id="13305DC17A426A6AFC7F5C71FBC74575" box="[1020,1063,1566,1589]" captionStart="Figures 274284" captionStartId="57.[192,278,1468,1492]" captionTargetBox="[196,1428,206,1431]" captionTargetId="figure-6@57.[192,1440,192,1439]" captionTargetPageId="57" captionText="Figures 274284: Mesobuthus, tarsomeres of third leg. Figure 274. M. brutus sp. n., male holotype. Figure 275. M. caucasicus, female from Turkey. Figure 276. M. elenae sp. n., female holotype. Figure 277. M. gorelovi sp. n., female paratype from type locality. Figure 278. M. fuscus, female from Tajikistan. Figure 279. M. kaznakovi, male from Tajikistan. Figure 280. M. kreuzbergi sp. n., female paratype. Figure 281. M. mischi sp. n., male holotype. Figure 282. M. nenilini sp. n., female paratype. Figure 283284. M. parthorum, male from Ghazni, Afghanistan (283) and female from Kholm, Afghanistan (284)." pageId="31" pageNumber="30">279</figureCitation>
,
<figureCitation id="13305DC17A426A6AFBC35C71FB404575" box="[1088,1184,1566,1589]" captionStart="Figures 285303" captionStartId="58.[192,278,1281,1305]" captionTargetBox="[198,1431,202,1246]" captionTargetId="figure-11@58.[192,1440,192,1252]" captionTargetPageId="58" captionText="Figures 285303: Mesobuthus, pedipalp chela dorsal. Figure 285286. M. brutus sp. n., male holotype (285) and female paratype (286). Figure 287288. M. caucasicus, male (287) and female (288) from Turkey. Figure 289. M. elenae sp. n., female holotype. Figure 290291. M. fuscus, differently colored male (290) and female (291) from Tajikistan. Figure 292293. M. gorelovi sp. n., male holotype (292) and female paratype (293). Figure 294295. M. kaznakovi, male (294) and female (295) from Tajikistan. Figure 296297. M. kreuzbergi sp. n., male holotype (296) and female paratype (297). Figure 298299. M. mischi sp. n., male holotype (298) and female paratype (299). Figure 300301. M. nenilini sp. n., male holotype (300) and female paratype (301). Figure 302303. M. parthorum, male (302) and female (303) from Afghanistan." pageId="31" pageNumber="30">294295</figureCitation>
,
<figureCitation id="13305DC17A426A6AFB395C71FB024575" box="[1210,1250,1566,1589]" captionStart="Figures 304315" captionStartId="59.[192,278,953,977]" captionTargetBox="[208,1427,193,918]" captionTargetId="figure-6@59.[192,1437,192,924]" captionTargetPageId="59" captionText="Figures 304315: Mesobuthus, pedipalp movable (304313) and fixed (314315) fingers. Figure 304. M. brutus sp. n., male paratype. Figure 305. M. caucasicus, male from Turkey. Figure 306. M. elenae sp. n., female holotype. Figure 307. M. gorelovi sp. n., male holotype. Figure 308. M. fuscus, female from Tajikistan. Figure 309. M. kaznakovi, male from Tajikistan. Figure 310, 314. M. kreuzbergi sp. n., male holotype. Figure 311. M. mischi sp. n., male holotype. Figure 312, 315. M. nenilini sp. n., male holotype. Figure 313. M. parthorum, male from Afghanistan." pageId="31" pageNumber="30">309</figureCitation>
,
<figureCitation id="13305DC17A426A6AFB7D5C71FAC74575" box="[1278,1319,1566,1589]" captionStart="Figures 316321" captionStartId="60.[192,278,1445,1469]" captionTargetBox="[191,1437,204,1417]" captionTargetId="figure-304@60.[192,1437,205,1416]" captionTargetPageId="60" captionText="Figures 316321: Mesobuthus, live specimens. Figure 316. M. kaznakovi, female from Uzbekistan. Figure 317. M. kaznakovi, male from Tajikistan. Figure 318. M. caucasicus, female from Iran. Figure 319. M. elenae sp. n., female. Figure 320. M. kreuzbergi sp. n., female paratype. Figure 321. M. parthorum, juvenile from Afghanistan." pageId="31" pageNumber="30">317</figureCitation>
) (
<collectionCode id="ED1AD9817A426A6AFACD5C71FA724575" box="[1358,1426,1566,1589]" pageId="31" pageNumber="30">FKCP</collectionCode>
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</materialsCitation>
.
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<collectingCountry id="F31C01D47A426A6AFCBD5C53FC5D4514" box="[830,957,1596,1620]" name="Uzbekistan" pageId="31" pageNumber="30">Uzbekistan</collectingCountry>
</emphasis>
,
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<collectingRegion id="49CF8FA67A426A6AFC535C52FB694514" box="[976,1161,1597,1620]" inferredBySuffix="true" pageId="31" pageNumber="30">Jizzakh Province</collectingRegion>
</emphasis>
, a
<collectingMunicipality id="6BD0DB3E7A426A6AFB315C52FA884514" box="[1202,1384,1597,1620]" pageId="31" pageNumber="30">Turkmen village</collectingMunicipality>
near
<location id="8ED4179F7A426A6AFCBD5C33FC6F4533" LSID="urn:lsid:plazi:treatment:03A2F0527A436A54FF5D5CC2FAA647B1:8ED4179F7A426A6AFCBD5C33FC6F4533" box="[830,911,1628,1651]" country="Uzbekistan" latitude="39.61" longLatPrecision="701" longitude="68.5" municipality="Turkmen village" name="Zaamin" pageId="31" pageNumber="30" stateProvince="Jizzakh Province">Zaamin</location>
,
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,
<date id="FFB567847A426A6AFBE05C33FAEF4533" box="[1123,1295,1628,1651]" pageId="31" pageNumber="30" value="2010-11">
<collectingDate id="EFF19E6C7A426A6AFBE05C33FAEF4533" box="[1123,1295,1628,1651]" pageId="31" pageNumber="30" value="2010-11">November 2010</collectingDate>
</date>
,
<specimenCount id="9D0D8ACD7A426A6AFA9E5C33FADE4533" box="[1309,1342,1628,1651]" pageId="31" pageNumber="30" type="male">2♂</specimenCount>
<specimenCount id="9D0D8ACD7A426A6AFABD5C33FA804533" box="[1342,1376,1628,1651]" pageId="31" pageNumber="30" type="female">3♀</specimenCount>
2juvs. (
<figureCitation id="13305DC17A426A6AFCC55C15FC4245D1" box="[838,930,1658,1681]" captionStart="Figures 316321" captionStartId="60.[192,278,1445,1469]" captionTargetBox="[191,1437,204,1417]" captionTargetId="figure-304@60.[192,1437,205,1416]" captionTargetPageId="60" captionText="Figures 316321: Mesobuthus, live specimens. Figure 316. M. kaznakovi, female from Uzbekistan. Figure 317. M. kaznakovi, male from Tajikistan. Figure 318. M. caucasicus, female from Iran. Figure 319. M. elenae sp. n., female. Figure 320. M. kreuzbergi sp. n., female paratype. Figure 321. M. parthorum, juvenile from Afghanistan." pageId="31" pageNumber="30">Fig. 316</figureCitation>
) (
<collectionCode id="ED1AD9817A426A6AFC3A5C15FBE145D1" box="[953,1025,1658,1681]" pageId="31" pageNumber="30">FKCP</collectionCode>
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31112CF7A426A6AFCBD5CD7FA9F458F" box="[830,1407,1720,1743]" pageId="31" pageNumber="30" type="distribution">
<paragraph id="8BB441447A426A6AFCBD5CD7FA9F458F" blockId="31.[830,1407,1720,1743]" box="[830,1407,1720,1743]" pageId="31" pageNumber="30">
DISTRIBUTION.
<collectingCountry id="F31C01D47A426A6AFC595CD7FBAB458F" box="[986,1099,1720,1743]" name="Tajikistan" pageId="31" pageNumber="30">Tajikistan</collectingCountry>
,
<collectingCountry id="F31C01D47A426A6AFBD55CD7FB2E458F" box="[1110,1230,1720,1743]" name="Uzbekistan" pageId="31" pageNumber="30">Uzbekistan</collectingCountry>
(
<figureCitation id="13305DC17A426A6AFB5D5CD7FAA4458F" box="[1246,1348,1720,1743]" captionStart="Figure 328" captionStartId="63.[192,268,1348,1372]" captionTargetBox="[187,1924,187,1319]" captionTargetId="figure-24@63.[192,1920,192,1314]" captionTargetPageId="63" captionText="Figure 328: Distribution map showing all species examined in this study. See Appendix A for distribution maps of each species." pageId="31" pageNumber="30">Figs. 328</figureCitation>
, A
<figureCitation id="13305DC17A426A6AFAE15CD7FA8F458F" box="[1378,1391,1720,1743]" captionStart="Figures 514" captionStartId="6.[192,278,1635,1659]" captionTargetBox="[191,1440,191,1606]" captionTargetId="figure-11@6.[192,1439,192,1605]" captionTargetPageId="6" captionText="Figures 514: Mesobuthus brutus sp. n. Figures 5, 7, 911. Holotype male, chelicerae, carapace and tergites IIII (5), sternopectinal region and sternite III (7), pedipalp chela, dorsal (9) and external (10), pedipalp patella dorsal (11). The trichobothrial pattern is indicated in Figures 910. Figures 6, 8, 1214. Paratype female, chelicerae, carapace and tergites IIII (6), sternopectinal region and sternite III (8), pedipalp chela, dorsal (12) and external (13), pedipalp patella dorsal (14)." pageId="31" pageNumber="30">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31112CF7A426A54FCBD5C9AFD144022" lastPageId="33" lastPageNumber="32" pageId="31" pageNumber="30" type="diagnosis">
<paragraph id="8BB441447A426A54FCBD5C9AFD144022" blockId="31.[830,1441,1781,1896]" lastBlockId="33.[192,802,199,866]" lastPageId="33" lastPageNumber="32" pageId="31" pageNumber="30">
DIAGNOSIS. (
<emphasis id="B97F9D567A426A6AFC4D5C9AFB0A444C" box="[974,1258,1781,1804]" italics="true" pageId="31" pageNumber="30">based on studied material</emphasis>
) Total length of adult males
<quantity id="4CF3ECA17A426A6AFC475D7BFBD9446B" box="[964,1081,1812,1835]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="31" pageNumber="30" unit="mm" value="69.0" valueMax="70.0" valueMin="68.0">6870 mm</quantity>
, 68
<specimenCount id="9D0D8ACD7A426A6AFBF25D7BFB0A446B" box="[1137,1258,1812,1835]" pageId="31" pageNumber="30" type="female">75 females</specimenCount>
. Trichobothrium
<emphasis id="B97F9D567A426A6AFCBD5D5DFCB94409" box="[830,857,1842,1865]" italics="true" pageId="31" pageNumber="30">db</emphasis>
on fixed finger of pedipalp situated between trichobothria
<emphasis id="B97F9D567A426A6AFC3D5D3EFC3C4428" box="[958,988,1873,1896]" italics="true" pageId="31" pageNumber="30">est</emphasis>
and
<emphasis id="B97F9D567A426A6AFB9E5D3EFBA04428" box="[1053,1088,1873,1896]" italics="true" pageId="31" pageNumber="30">esb</emphasis>
, near to
<emphasis id="B97F9D567A426A6AFB275D3EFB224428" box="[1188,1218,1873,1896]" italics="true" pageId="31" pageNumber="30">est</emphasis>
. Male with fingers proximally little more twisted than female. Female has longer and narrower chela of pedipalps. Pedipalp chela length/ width ratio
<quantity id="4CF3ECA17A7C6A54FE115B6BFDFB425B" box="[402,539,260,283]" metricMagnitude="-2" metricUnit="m" metricValue="9.0805" metricValueMax="9.270999999999999" metricValueMin="8.89" pageId="33" pageNumber="32" unit="in" value="3.575" valueMax="3.65" valueMin="3.5">3.503.65 in</quantity>
males and
<quantity id="4CF3ECA17A7C6A54FD195B6BFCC1425B" box="[666,801,260,283]" metricMagnitude="-2" metricUnit="m" metricValue="9.8298" metricValueMax="9.9822" metricValueMin="9.6774" pageId="33" pageNumber="32" unit="in" value="3.87" valueMax="3.93" valueMin="3.81">3.813.93 in</quantity>
females. Pectinal teeth number
<quantity id="4CF3ECA17A7C6A54FD925B4CFD90427A" box="[529,624,291,314]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" metricValueMax="6.858" metricValueMin="6.35" pageId="33" pageNumber="32" unit="in" value="26.0" valueMax="27.0" valueMin="25.0">2527 in</quantity>
males,
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females. Chelicerae yellow, without reticulation. Pedipalps and metasoma sparsely hirsute. Carapace and tergites yellowish brown, black pigmented; metasoma, telson, pedipalps and legs yellowish brown, only anterior part of metasomal segment V black. Femur of pedipalp with 45 granulated carinae. Patella with 8 granulated or smooth carinae. Chela lacks carinae. Movable fingers of pedipalps with 13 cutting rows of denticles and 5 terminal denticles. Seventh sternite bears 4 well marked granulate carinae. First metasomal segment with 10 carinae; second to third with 8 carinae, other two carinae indicated by incomplete row of denticles; fourth with 8 carinae; fifth with 5 carinae. All carinae granulated, dorsal carinae bear slightly larger terminal denticles. Length to width ratio of fourth metasomal segment
<quantity id="4CF3ECA17A7C6A54FD195880FCC24046" box="[666,802,751,774]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" metricValueMax="4.5466" metricValueMin="4.3434" pageId="33" pageNumber="32" unit="in" value="1.75" valueMax="1.79" valueMin="1.71">1.711.79 in</quantity>
males,
<quantity id="4CF3ECA17A7C6A54FE935961FE7C4065" box="[272,412,782,805]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" metricValueMax="4.3434" metricValueMin="4.2926" pageId="33" pageNumber="32" unit="in" value="1.7" valueMax="1.71" valueMin="1.69">1.691.71 in</quantity>
females. Telotarsus III ventral setation represented by short and strong spiniform setae. Pedal spur of legs obviously with solitary setae only.
</paragraph>
</subSubSection>
<caption id="DF7411CC7A7D6A55FF435CB2FA9F4466" pageId="32" pageNumber="31" startId="32.[192,278,1757,1781]" targetBox="[196,1433,208,1720]" targetPageId="32">
<paragraph id="8BB441447A7D6A55FF435CB2FA9F4466" blockId="32.[192,1440,1757,1830]" pageId="32" pageNumber="31">
<emphasis id="B97F9D567A7D6A55FF435CB2FE6345B5" bold="true" box="[192,387,1757,1781]" pageId="32" pageNumber="31">Figures 131138:</emphasis>
<taxonomicName id="4C0B3AC77A7D6A55FE085CB0FD8645B4" baseAuthorityName="Birula" baseAuthorityYear="1904" box="[395,614,1759,1780]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="kaznakovi">
<emphasis id="B97F9D567A7D6A55FE085CB0FD8645B4" box="[395,614,1759,1780]" italics="true" pageId="32" pageNumber="31">Mesobuthus kaznakovi</emphasis>
</taxonomicName>
.
<emphasis id="B97F9D567A7D6A55FDF15CB0FD0B45B4" bold="true" box="[626,747,1759,1781]" pageId="32" pageNumber="31">Figures 131</emphasis>
,
<emphasis id="B97F9D567A7D6A55FD7B5CB0FCAC45B4" bold="true" box="[760,844,1759,1780]" pageId="32" pageNumber="31">133135</emphasis>
. Male from Ramit, Tajikistan, metasoma V and telson lateral (131), and metasoma and telson, lateral (133), ventral (134), and dorsal (135) views.
<emphasis id="B97F9D567A7D6A55FC7B5C97FB90444D" bold="true" box="[1016,1136,1784,1806]" pageId="32" pageNumber="31">Figures 132</emphasis>
,
<emphasis id="B97F9D567A7D6A55FBFF5C97FB30444D" bold="true" box="[1148,1232,1784,1805]" pageId="32" pageNumber="31">136138</emphasis>
. Female from Ramit, Tajikistan, metasoma V and telson lateral (132), and metasoma and telson, lateral (136), ventral (137), and dorsal (138) views.
</paragraph>
</caption>
<subSubSection id="C31112CF7A7C6A54FF4359E5FAA647B1" pageId="33" pageNumber="32" type="description">
<paragraph id="8BB441447A7C6A54FF4359E5FED447F5" blockId="33.[192,802,906,1910]" pageId="33" pageNumber="32">
REDESCRIPTION. The total length of adult males
<quantity id="4CF3ECA17A7C6A54FD5C59E5FF0C40FF" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="33" pageNumber="32" unit="mm" value="69.0" valueMax="70.0" valueMin="68.0">6870 mm</quantity>
, 68
<specimenCount id="9D0D8ACD7A7C6A54FEA159C7FE7B40FF" box="[290,411,936,959]" pageId="33" pageNumber="32" type="female">75 females</specimenCount>
. Trichobothrium
<emphasis id="B97F9D567A7C6A54FDD959C7FD9540FF" box="[602,629,936,959]" italics="true" pageId="33" pageNumber="32">db</emphasis>
on fixed finger of pedipalp is situated between trichobothria
<emphasis id="B97F9D567A7C6A54FD2659A8FD23409E" box="[677,707,967,990]" italics="true" pageId="33" pageNumber="32">est</emphasis>
and
<emphasis id="B97F9D567A7C6A54FD7B59A8FCFC409E" box="[760,796,967,990]" italics="true" pageId="33" pageNumber="32">esb</emphasis>
, near to
<emphasis id="B97F9D567A7C6A54FE915989FED040BD" box="[274,304,998,1021]" italics="true" pageId="33" pageNumber="32">est</emphasis>
. Male has the fingers proximally a little more twisted than female. Female has longer and slightly narrower chela of pedipalps. Chelicerae yellow, without reticulation, the tips of teeth on cheliceral fingers are black. For the position and distribution of trichobothria see
<figureCitation id="13305DC17A7C6A54FF715E10FDED47D6" box="[242,525,1151,1174]" captionStart="Figures 139151" captionStartId="34.[192,278,1252,1276]" captionTargetBox="[201,1434,180,1206]" captionTargetId="figure-11@34.[192,1439,168,1223]" captionTargetPageId="34" captionText="Figures 139151: Mesobuthus kaznakovi. Figures 139141. Female from Ramit, Tajikistan, Pedipalp chela, dorsal (139) and external (140), and pedipalp patella dorsal (141). Figures 142151. Male from Ramit, Tajikistan, Pedipalp chela, dorsal (142), external (143), and ventral (144) views. Pedipalp patella, dorsal (145), external (146), and ventral (147) views. Pedipalp femur and trochanter, internal (148) and dorsal (149) views. Right chelicera dorsal (150) and ventral (151) views. The trichobothrial pattern is indicated in Figures 142146, 148149." pageId="33" pageNumber="32">Figs. 142146, 148149</figureCitation>
. For measurements see tables 45.
</paragraph>
<paragraph id="8BB441447A7C6A54FF435ED3FF11460E" blockId="33.[192,802,906,1910]" pageId="33" pageNumber="32">
COLORATION (
<figureCitation id="13305DC17A7C6A54FEDF5ED3FE1C4793" box="[348,508,1212,1235]" captionStart="Figures 119122" captionStartId="30.[192,278,1470,1494]" captionTargetBox="[192,1437,144,1455]" captionTargetId="figure-11@30.[192,1437,144,1455]" captionTargetPageId="30" captionText="Figures 119122: Mesobuthus kaznakovi. Figures 119120. Male from Ramit, Tajikistan, dorsal (119) and ventral (120) views. Figures 121122. Female from Ramit, Tajikistan, dorsal (121) and ventral (122) views." pageId="33" pageNumber="32">Figs. 119122</figureCitation>
). The carapace and tergites are yellowish brown strongly black pigmented; the metasoma, telson, pedipalps and legs yellowish brown to black. Metasomal segment V is black mainly in anterior part.
</paragraph>
<paragraph id="8BB441447A7C6A54FF435F39FEFB4479" blockId="33.[192,802,906,1910]" pageId="33" pageNumber="32">
MESOSOMA AND CARAPACE (
<figureCitation id="13305DC17A7C6A54FD825F39FD43462D" box="[513,675,1366,1389]" captionStart="Figures 123130" captionStartId="31.[192,278,1409,1433]" captionTargetBox="[191,1438,191,1381]" captionTargetId="figure-6@31.[192,1438,192,1380]" captionTargetPageId="31" captionText="Figures 123130: Mesobuthus kaznakovi. Figures 123, 125, 127130. Male from Ramit, Tajikistan, chelicerae, carapace and tergites IIII (123), sternopectinal region and sternites IIIV (125), left legs IIV, retrolateral aspect (127130). Figures 124, 126: Paratype female, chelicerae, carapace and tergites IIII (124), and sternopectinal region and sternites IIIIV (126)." pageId="33" pageNumber="32">Figs. 123126</figureCitation>
). The carapace is carinated and unevenly covered by granules of varying size; much of the granulation is fine, but some granules are larger and distinctly rounded. Tergites IVI bear three carinae and are granulated, with some intercarinal granules small and others larger and rounded. Tergite VII is pentacarinate. The pectinal tooth count is
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males,
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females. The pectinal marginal tips extend to about third of the fifth sternite in males and third of the fourth sternite in females. The pectines have three marginal lamellae and seven to nine middle lamellae. The lamellae bear numerous long setae, each fulcrum with three to five setae. All sternites are smooth and sparsely hirsute. The seventh sternite bears four well marked granulate carinae. The other sternites bear two furrows.
</paragraph>
<paragraph id="8BB441447A7C6A54FF435D2FFAC242F5" blockId="33.[192,802,906,1910]" lastBlockId="33.[830,1441,199,1111]" pageId="33" pageNumber="32">
PEDIPALPS (
<figureCitation id="13305DC17A7C6A54FEC05D2FFE054417" box="[323,485,1856,1879]" captionStart="Figures 139151" captionStartId="34.[192,278,1252,1276]" captionTargetBox="[201,1434,180,1206]" captionTargetId="figure-11@34.[192,1439,168,1223]" captionTargetPageId="34" captionText="Figures 139151: Mesobuthus kaznakovi. Figures 139141. Female from Ramit, Tajikistan, Pedipalp chela, dorsal (139) and external (140), and pedipalp patella dorsal (141). Figures 142151. Male from Ramit, Tajikistan, Pedipalp chela, dorsal (142), external (143), and ventral (144) views. Pedipalp patella, dorsal (145), external (146), and ventral (147) views. Pedipalp femur and trochanter, internal (148) and dorsal (149) views. Right chelicera dorsal (150) and ventral (151) views. The trichobothrial pattern is indicated in Figures 142146, 148149." pageId="33" pageNumber="32">Figs. 139149</figureCitation>
). The pedipalps are sparsely hirsute and smooth. The femur bears four to five granulated carinae, the middle carina on internal surface could be incomplete indicated by several strong granules. The patella bears eight carinae from which three internal are granulated and other are smooth in males but could by granulated in females. The chela is without carinae. The movable fingers of pedipalps bear 13 cutting rows of denticles, every with external and internal denticles present, and five terminal denticles.
</paragraph>
<paragraph id="8BB441447A7C6A54FCBD5BD3FBDC4189" blockId="33.[830,1441,199,1111]" pageId="33" pageNumber="32">
LEGS (
<figureCitation id="13305DC17A7C6A54FC055BD3FBC64293" box="[902,1062,444,467]" captionStart="Figures 123130" captionStartId="31.[192,278,1409,1433]" captionTargetBox="[191,1438,191,1381]" captionTargetId="figure-6@31.[192,1438,192,1380]" captionTargetPageId="31" captionText="Figures 123130: Mesobuthus kaznakovi. Figures 123, 125, 127130. Male from Ramit, Tajikistan, chelicerae, carapace and tergites IIII (123), sternopectinal region and sternites IIIV (125), left legs IIV, retrolateral aspect (127130). Figures 124, 126: Paratype female, chelicerae, carapace and tergites IIII (124), and sternopectinal region and sternites IIIIV (126)." pageId="33" pageNumber="32">Figs. 127130</figureCitation>
). The tarsomeres bear two rows of spiniform setae on the ventral surface and numerous macrosetae on the other surfaces. Ventral setation of tarsomere II (telotarsus) represented by short and strong spiniform setae. Pedal spur of legs obviously with solitary setae only. Femur bears only several macrosetae. Femur and patella with carinae well developed. Tibial spurs present and long on third and fourth legs and absent in the other legs.
</paragraph>
<paragraph id="8BB441447A7C6A54FCBD58BFFAD14046" blockId="33.[830,1441,199,1111]" pageId="33" pageNumber="32">
METASOMA AND TELSON (
<figureCitation id="13305DC17A7C6A54FBF358BFFAF741A7" box="[1136,1303,720,743]" captionStart="Figures 131138" captionStartId="32.[192,278,1757,1781]" captionTargetBox="[196,1433,208,1720]" captionTargetId="figure-11@32.[192,1439,197,1728]" captionTargetPageId="32" captionText="Figures 131138: Mesobuthus kaznakovi. Figures 131, 133135. Male from Ramit, Tajikistan, metasoma V and telson lateral (131), and metasoma and telson, lateral (133), ventral (134), and dorsal (135) views. Figures 132, 136138. Female from Ramit, Tajikistan, metasoma V and telson lateral (132), and metasoma and telson, lateral (136), ventral (137), and dorsal (138) views." pageId="33" pageNumber="32">Figs. 131138</figureCitation>
). All metasomal segments are only very sparsely hirsute.
</paragraph>
<paragraph id="8BB441447A7C6A54FCBD5961FB174717" blockId="33.[830,1441,199,1111]" pageId="33" pageNumber="32">The metasomal segment I with 10 carinae, IIIII with 8 carinae but other two lateromedian carinae are indicated by incomplete row of denticles, IV with 8 carinae, and V with 5 carinae. All carinae with denticles, dorsal carinae on metasomal segment IIII bear slightly larger terminal denticles. The dorsal surface of all segments is smooth in the middle and bumpy on margins. Other surfaces are bumpy with several solitary granules. Ventrolateral carinae of metasomal segment V posteriorly with several large lobate denticles. The telson is only sparsely hirsute, rather elongate, bumpy and smooth.</paragraph>
<paragraph id="8BB441447A7C6A54FCBD5E11FC6E47D5" blockId="33.[830,910,1150,1173]" box="[830,910,1150,1173]" pageId="33" pageNumber="32">NOTES.</paragraph>
<paragraph id="8BB441447A7C6A54FCD85ED4FAA647B1" blockId="33.[859,1439,1211,1265]" pageId="33" pageNumber="32">
<bibRefCitation id="EF9A3CB57A7C6A54FCD85ED4FC334792" author="FET" box="[859,979,1211,1234]" pageId="33" pageNumber="32" pagination="73 - 171" refId="ref33201" refString="FET, V. 1989. A catalogue of scorpions (Chelicerata: Scorpiones) of the USSR. Rivista del Museo Civico di Scienze Naturali &quot; Enrico Caffi &quot;, 13 (1988): 73 - 171." type="journal article" year="1989">Fet (1989)</bibRefCitation>
did not list
<typeStatus id="54B0FFE67A7C6A54FBF25ED4FB2F4792" box="[1137,1231,1211,1234]" pageId="33" pageNumber="32" type="syntype">syntypes</typeStatus>
from Karasu and Shugnan, which could not be located in ZISP.
</paragraph>
</subSubSection>
</treatment>
</document>