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<document id="FA7441599DCB2030B910D331EFEF9D21" ID-DOI="10.5281/zenodo.174070" ID-GBIF-Dataset="0af0afa8-e726-4eff-8444-7d419d5ee9da" ID-ISSN="1175­5326" ID-Zenodo-Dep="174070" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459789414158" checkinUser="plazi" docAuthor="Tourinho, Ana Lúcia &amp; González, Abel Pérez" docDate="2006" docId="03A2CA197659FFDFFEF4237E2CD0FD1F" docLanguage="en" docName="zt01325p254.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Fissiphallius chicoi Tourinho &amp; González, 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="242" masterDocId="FF9BB261765DFFD8FFFC22622F0FFF81" masterDocTitle="On the family Fissiphalliidae Martens, 1988, with descriptions of two new Amazonian species (Arachnida: Opiliones: Laniatores)" masterLastPageNumber="254" masterPageNumber="235" pageNumber="239" updateTime="1698219041982" updateUser="plazi">
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<mods:title id="6C272390B489B4A1F08E2FABBD8EEAF3">On the family Fissiphalliidae Martens, 1988, with descriptions of two new Amazonian species (Arachnida: Opiliones: Laniatores)</mods:title>
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<mods:namePart id="D2A971738828C06394499B4F6DA1C46C">Tourinho, Ana Lúcia</mods:namePart>
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<paragraph id="8BB47B0F7659FFDCFEF4237E2D30FEB4" blockId="4.[264,672,283,345]" box="[264,575,283,310]" pageId="4" pageNumber="239">
<heading id="D0FCCC637659FFDCFEF4237E2D30FEB4" bold="true" box="[264,575,283,310]" fontSize="11" level="1" pageId="4" pageNumber="239" reason="1">
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<taxonomicName id="4C0B008C7659FFDCFEF4237E2EFBFEB7" box="[264,500,284,310]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="chicoi" status="sp. nov.">
<emphasis id="B97FA71D7659FFDCFEF4237E2EFBFEB7" bold="true" box="[264,500,284,310]" italics="true" pageId="4" pageNumber="239">Fissiphallius chicoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C1A667659FFDCFE0023792D30FEB4" box="[508,575,283,309]" pageId="4" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BB47B0F7659FFDCFEF4235D2DAFFED8" blockId="4.[264,672,283,345]" box="[264,672,319,345]" pageId="4" pageNumber="239">
<figureCitation id="1330678A7659FFDCFEF4235D2EA4FED8" box="[264,427,319,345]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Figs 1, 3, 5, 7</figureCitation>
,
<figureCitation id="1330678A7659FFDCFE4B235D2D10FED8" box="[439,543,319,345]" captionStart="FIGURES 9 14" captionStartId="6.[264,382,1645,1669]" captionTargetBox="[295,1298,332,1619]" captionTargetId="figure@6.[276,1314,296,1623]" captionTargetPageId="6" captionText="FIGURES 9 14. Fissiphallius chicoi n. sp., male holotype (9, 11 12); F. t u c u p i n. sp., male holotype (10, 13 14). 9 10 right leg IV, lateral view; 11 12 right pedipalp (11 ectal view, 12 mesal view); 13 14 right pedipalp (13 ectal view, 14 mesal view). Scale bars = 1.0 mm (9, 11 12), 0.5 mm (10, 13 14)." httpUri="https://zenodo.org/record/174072/files/figure.png" pageId="4" pageNumber="239">9, 1112</figureCitation>
,
<figureCitation id="1330678A7659FFDCFDD7235D2D78FED8" box="[555,631,319,345]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="4" pageNumber="239">1618</figureCitation>
,
<figureCitation id="1330678A7659FFDCFD7F235D2DAFFED8" box="[643,672,319,345]" captionStart="FIGURE 22" captionStartId="14.[264,367,1755,1779]" captionTargetBox="[359,1230,953,1740]" captionTargetId="figure@14.[359,1231,953,1741]" captionTargetPageId="14" captionText="FIGURE 22. Northern South America showing the localities of the six nominal species of Fissiphallius. F. s t u r m i Martens, F. spinulatus Martens, F. sympatricus Martens (pentagon), F. martensi Pinto­da­Rocha (triangle), F. t u c u p i n. sp. (star), F. c h i c o i n. sp. (circle). Dotted lines = country boundaries, thin lines = rivers." httpUri="https://zenodo.org/record/174075/files/figure.png" pageId="4" pageNumber="239">22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31128847659FFDCFEF423EC2EF6FDA0" pageId="4" pageNumber="239" type="materials_examined">
<paragraph id="8BB47B0F7659FFDCFEF423EC2EF6FDA0" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">
<emphasis id="B97FA71D7659FFDCFEF423EC2EB9FE29" bold="true" box="[264,438,398,424]" pageId="4" pageNumber="239">
<typeStatus id="54B0C5AD7659FFDCFEF423EC2E4CFE29" box="[264,323,398,424]" pageId="4" pageNumber="239">Type</typeStatus>
material
</emphasis>
. Male
<typeStatus id="54B0C5AD7659FFDCFDF323ED2D79FE28" box="[527,630,399,425]" pageId="4" pageNumber="239" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED1AE3CA7659FFDCFD7A23ED2DC5FE28" box="[646,714,399,425]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="4" pageNumber="239">INPA</collectionCode>
159),
<collectingCountry id="F31C3B9F7659FFDCFCE623ED2C6AFE28" box="[794,869,399,425]" name="Brazil" pageId="4" pageNumber="239">Brazil</collectingCountry>
, Pará State, Gurupá municipality, São José community,
<geoCoordinate id="EE3F1DC87659FFDCFE1D23D52D30FE50" box="[481,575,439,465]" direction="south" orientation="latitude" pageId="4" pageNumber="239" precision="55" value="-1.197">1.197ºS</geoCoordinate>
,
<geoCoordinate id="EE3F1DC87659FFDCFDAD23D52DC8FE50" box="[593,711,439,465]" direction="west" orientation="longitude" pageId="4" pageNumber="239" precision="55" value="-51.783">51.783ºW</geoCoordinate>
, Próvárzea,
<date id="FFB55DCF7659FFDCFC9F23D52CEDFE50" box="[867,994,439,465]" pageId="4" pageNumber="239" value="2003-10-18">18.X.2003</date>
,
<taxonomicName id="4C0B008C7659FFDCFC0823D52E97FE78" authority="Anne Rheims &amp; Felipe Rego" authorityName="Anne Rheims &amp; Felipe Rego" class="Arachnida" family="Phalangiidae" genus="Cristina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="239" phylum="Arthropoda" rank="genus">Cristina Anne Rheims &amp; Felipe Rego</taxonomicName>
leg.
<typeStatus id="54B0C5AD7659FFDCFE2F23BD2D46FE78" box="[467,585,479,505]" pageId="4" pageNumber="239" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0DB0867659FFDCFDA923BD2DB3FE78" box="[597,700,479,505]" pageId="4" pageNumber="239" type="female">1 female</specimenCount>
(
<collectionCode id="ED1AE3CA7659FFDCFD3023BD2C1FFE78" box="[716,784,479,505]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="4" pageNumber="239">INPA</collectionCode>
89),
<specimenCount id="9D0DB0867659FFDCFCB323BD2CCEFE78" box="[847,961,479,505]" pageId="4" pageNumber="239" type="female">3 females</specimenCount>
(
<collectionCode id="ED1AE3CA7659FFDCFC2D23BD2B2EFE78" box="[977,1057,479,505]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="4" pageNumber="239">MNRJ</collectionCode>
17814) with the same data as the
<typeStatus id="54B0C5AD7659FFDCFE7720652EFAFDA0" box="[395,501,519,545]" pageId="4" pageNumber="239" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C31128847659FFDCFEC4204C2BB8FD18" pageId="4" pageNumber="239" type="etymology">
<paragraph id="8BB47B0F7659FFDCFEC4204C2BB8FD18" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">
<emphasis id="B97FA71D7659FFDCFEC4204C2ECBFDC9" bold="true" box="[312,452,558,584]" pageId="4" pageNumber="239">Etymology.</emphasis>
The species name is a patronym in honor of Chico Mendes, a Brazilian national hero, symbol of the defense of the Amazonian environment and of rubber workers rights. Chico was cowardly assassinated in
<quantity id="4CF3D6EA7659FFDCFC89201D2CC1FD18" box="[885,974,639,665]" metricMagnitude="1" metricUnit="m" metricValue="5.04952" pageId="4" pageNumber="239" unit="in" value="1988.0">1988 in</quantity>
Xapuri, Acre State.
</paragraph>
</subSubSection>
<subSubSection id="C31128847659FFDCFEC420C42CD1FB80" pageId="4" pageNumber="239" type="diagnosis">
<paragraph id="8BB47B0F7659FFDCFEC420C42CD1FB80" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">
<emphasis id="B97FA71D7659FFDCFEC420C42EBFFD41" bold="true" box="[312,432,678,704]" pageId="4" pageNumber="239">Diagnosis</emphasis>
. Eye mound armed with one pair of small sharp spiniform aphophysis (
<figureCitation id="1330678A7659FFDCFB0420C52E3AFD68" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Figs 1, 3</figureCitation>
). Male genitalia clearly different from those of all other congeners. Rutrum with large number of setae irregularly arranged (paired in other species), pergula narrow, without setae. Portion of truncus below pergula very thin and narrower than stragulum (
<figureCitation id="1330678A7659FFDCFB0B217D2E5AFCE0" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="4" pageNumber="239">Figs 1618</figureCitation>
). Ventral side of truncus, near base of stragulum, with a chitinous structure in the shape of a balcony and armed with three sharp­pointed setae (
<figureCitation id="1330678A7659FFDCFC03210D2B85FC08" box="[1023,1162,879,905]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="4" pageNumber="239">Figs 1718</figureCitation>
), two ventral rows of conic protuberances (similar laterodorsal rows of protuberances present in
<taxonomicName id="4C0B008C7659FFDCFAE921F52E8FFC59" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="spinulatus">
<emphasis id="B97FA71D7659FFDCFAE921F52E8FFC59" italics="true" pageId="4" pageNumber="239">F. spinulatus</emphasis>
</taxonomicName>
) (see
<bibRefCitation id="EF9A06FE7659FFDCFE3221DD2D78FC58" author="Martens" box="[462,631,959,985]" pageId="4" pageNumber="253" refString="Martens, J. (1988) Fissiphalliidae, a new family of South American laniatorean harvestmen (Arachnida: Opiliones). Zeitschrift fur Zoologische Systematik und Evolutionsforschung, 26 (2), 114 - 127." type="journal article" year="1988">Martens 1988</bibRefCitation>
: figs 1718). Truncus basis ventrally with other balconyshaped structure and dorsally with a folded area (
<figureCitation id="1330678A7659FFDCFCB421852CDEFB80" box="[840,977,999,1025]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="4" pageNumber="239">Figs 1618</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31128847659FFDDFEC4266C2C8AF8EE" lastPageId="5" lastPageNumber="240" pageId="4" pageNumber="239" type="description">
<paragraph id="8BB47B0F7659FFDCFEC4266C2EBCFBF8" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">
<emphasis id="B97FA71D7659FFDCFEC4266C2DA1FBA9" bold="true" box="[312,686,1038,1064]" pageId="4" pageNumber="239">
Description of male
<typeStatus id="54B0C5AD7659FFDCFDC0266C2DA6FBA9" box="[572,681,1038,1064]" pageId="4" pageNumber="239" type="holotype">holotype</typeStatus>
.
</emphasis>
Measurements. Total length: 3.2 Carapace region of scutum: 1.0 long, 1.2 wide. Scutum: 2.4 long, 2.1 maximum width. Pedipalp: 2.7. Legs: 7.3/5.1/6.2/4.0
</paragraph>
<paragraph id="8BB47B0F7659FFDCFEC426E52BD4FA38" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">
Dorsal view. Anterior border of prosoma with one median tubercle and a pair of tubercles on each side (
<figureCitation id="1330678A7659FFDCFDE126CD2D6DFB48" box="[541,610,1199,1225]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Fig. 1</figureCitation>
). Spiniform aphophyses of eye mound short (
<figureCitation id="1330678A7659FFDCFB7E26CD2BC6FB48" box="[1154,1225,1199,1225]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Fig. 3</figureCitation>
). Scutal areas IIV with a row of very small tubercles each (almost indiscernible), lateral margin of posterior half of dorsal scutum armed on each side with six sharp­pointed tubercles increasing in length posteriorly (
<figureCitation id="1330678A7659FFDCFD7B27452DC3FAC0" box="[647,716,1319,1345]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Fig. 1</figureCitation>
); posterior margin armed with very small tubercles (almost indiscernible). Free tergites IIII armed with sharp­pointed tubercles, the three median and the lateral ones slightly larger than others; free tergite I with 15 tubercles, III with 13, II with 11; the median one always larger than other tubercles (
<figureCitation id="1330678A7659FFDCFBBA27FD2BC2FA38" box="[1094,1229,1439,1465]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Figs 1, 3, 7</figureCitation>
).
</paragraph>
<paragraph id="8BB47B0F7659FFDCFEC427A52B68F9D8" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">
Ventral view. Posterior margin and sternites with tubercles increasing in length laterally, three to five lateral ones visibly larger (
<figureCitation id="1330678A7659FFDCFCC3278D2C88F988" box="[831,903,1519,1545]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Fig. 5</figureCitation>
). Sternite IV and V with one pair of larger median tubercles (
<figureCitation id="1330678A7659FFDCFDCF24752D72F9B0" box="[563,637,1559,1585]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Fig. 5</figureCitation>
), anal operculum armed with an anterior and a posterior row of tubercles (
<figureCitation id="1330678A7659FFDCFE2A245D2D12F9D8" box="[470,541,1599,1625]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Fig. 7</figureCitation>
), median spine of posterior row larger (
<figureCitation id="1330678A7659FFDCFC13245D2B57F9D8" box="[1007,1112,1599,1625]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="4" pageNumber="239">Figs 5, 7</figureCitation>
).
</paragraph>
<paragraph id="8BB47B0F7659FFDCFEC424052C12F928" blockId="4.[264,1325,398,1905]" pageId="4" pageNumber="239">Chelicerae. Without remarkable armature. Two tubercles on the basichelicerite, bulla short and well marked. Fingers without teeth.</paragraph>
<paragraph id="8BB47B0F7659FFDDFEC424D52EFAFEB7" blockId="4.[264,1325,398,1905]" lastBlockId="5.[264,1323,284,550]" lastPageId="5" lastPageNumber="240" pageId="4" pageNumber="239">
Pedipalps. Coxa with one ventral tubercle carrying setae and one small dorsal protuberance. Femur ventrally with two basal and one distal setiferous tubercle, one distal setiferous tubercle in the mesal region. Patella mesally with one setiferous tubercle. Tibia mesally and ectally with one row of three setiferous tubercles. Tarsus mesally and ectally with two setiferous tubercles on each side; small setae scattered. Tarsal claw long and strong (
<figureCitation id="1330678A7658FFDDFE9D237E2EE7FEB7" box="[353,488,284,310]" captionStart="FIGURES 9 14" captionStartId="6.[264,382,1645,1669]" captionTargetBox="[295,1298,332,1619]" captionTargetId="figure@6.[276,1314,296,1623]" captionTargetPageId="6" captionText="FIGURES 9 14. Fissiphallius chicoi n. sp., male holotype (9, 11 12); F. t u c u p i n. sp., male holotype (10, 13 14). 9 10 right leg IV, lateral view; 11 12 right pedipalp (11 ectal view, 12 mesal view); 13 14 right pedipalp (13 ectal view, 14 mesal view). Scale bars = 1.0 mm (9, 11 12), 0.5 mm (10, 13 14)." httpUri="https://zenodo.org/record/174072/files/figure.png" pageId="5" pageNumber="240">Figs 1112</figureCitation>
).
</paragraph>
<paragraph id="8BB47B0F7658FFDDFEC423262EB4FDA7" blockId="5.[264,1323,284,550]" pageId="5" pageNumber="240">
Legs. IIV tuberculate. Coxae IV with sharp­pointed dorsal tubercles, posterior margin of coxae IIIII armed with larger tubercles (
<figureCitation id="1330678A7658FFDDFC90230E2CD8FE07" box="[876,983,364,390]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="5" pageNumber="240">Figs 1, 5</figureCitation>
). Trochanters IIV with one sharp­pointed retrolateral tubercle (
<figureCitation id="1330678A7658FFDDFD5123F62DF8FE2F" box="[685,759,404,430]" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="5" pageNumber="240">Fig. 5</figureCitation>
). Femur IV with two rows of 1011 large and thin tubercles. Patella IV with sharp­pointed distal tubercles larger than others. Tibiae with one ventroapical tubercle larger than others. Tarsi tuberculate (
<figureCitation id="1330678A7658FFDDFBF723862B57FE7F" box="[1035,1112,484,510]" captionStart="FIGURES 9 14" captionStartId="6.[264,382,1645,1669]" captionTargetBox="[295,1298,332,1619]" captionTargetId="figure@6.[276,1314,296,1623]" captionTargetPageId="6" captionText="FIGURES 9 14. Fissiphallius chicoi n. sp., male holotype (9, 11 12); F. t u c u p i n. sp., male holotype (10, 13 14). 9 10 right leg IV, lateral view; 11 12 right pedipalp (11 ectal view, 12 mesal view); 13 14 right pedipalp (13 ectal view, 14 mesal view). Scale bars = 1.0 mm (9, 11 12), 0.5 mm (10, 13 14)." httpUri="https://zenodo.org/record/174072/files/figure.png" pageId="5" pageNumber="240">Fig. 9</figureCitation>
). Tarsal formula: 3(2), 5(3), 5, 5.
</paragraph>
<caption id="DF742B877658FFDDFEF4273E2C72FA38" httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="5" pageNumber="240" targetBox="[265,1319,621,1329]" targetPageId="5">
<paragraph id="8BB47B0F7658FFDDFEF4273E2C72FA38" blockId="5.[264,1323,1372,1465]" pageId="5" pageNumber="240">
<emphasis id="B97FA71D7658FFDDFEF4273E2EA1FAF5" bold="true" box="[264,430,1372,1396]" pageId="5" pageNumber="240">FIGURES 18</emphasis>
.
<taxonomicName id="4C0B008C7658FFDDFE41273F2D9FFAF5" box="[445,656,1373,1396]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="chicoi" status="sp. nov.">
<emphasis id="B97FA71D7658FFDDFE41273F2D9FFAF5" box="[445,656,1373,1396]" italics="true" pageId="5" pageNumber="240">Fissiphallius chicoi</emphasis>
</taxonomicName>
<emphasis id="B97FA71D7658FFDDFD65273E2DD9FAF5" bold="true" box="[665,726,1372,1396]" pageId="5" pageNumber="240">
<taxonomicNameLabel id="A24C1A667658FFDDFD65273E2DD9FAF5" box="[665,726,1372,1396]" pageId="5" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (1, 3, 5, 7);
<taxonomicName id="4C0B008C7658FFDDFBF6273F2BD1FAF5" box="[1034,1246,1373,1396]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="tucupi" status="sp. nov.">
<emphasis id="B97FA71D7658FFDDFBF6273F2BD1FAF5" box="[1034,1246,1373,1396]" italics="true" pageId="5" pageNumber="240">Fissiphallius tucupi</emphasis>
</taxonomicName>
<emphasis id="B97FA71D7658FFDDFB1A273E2A2BFAF5" bold="true" box="[1254,1316,1372,1396]" pageId="5" pageNumber="240">
<taxonomicNameLabel id="A24C1A667658FFDDFB1A273E2A2BFAF5" box="[1254,1316,1372,1396]" pageId="5" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (2, 4, 6, 8). Body (12 dorsal view, 34 lateral view, 56 ventral view, 78 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8).
</paragraph>
</caption>
<paragraph id="8BB47B0F7658FFDDFEC4278F2C8AF8EE" blockId="5.[264,1323,1517,1903]" pageId="5" pageNumber="240">
Penis. Stragulum entire, with a long central furrow only visible distally. Apical portion of stragulum in the shape of a “parrot bill” (
<figureCitation id="1330678A7658FFDDFCF124772C6BF9AE" box="[781,868,1557,1583]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="5" pageNumber="240">Fig. 17</figureCitation>
). Stragulum as long as in
<emphasis id="B97FA71D7658FFDDFB6924772BF6F9AF" box="[1173,1273,1557,1582]" italics="true" pageId="5" pageNumber="240">F. s t u r m i</emphasis>
but with shorter rutrum (
<figureCitation id="1330678A7658FFDDFDF3245F2D90F9D6" box="[527,671,1597,1623]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="5" pageNumber="240">Figs 1718</figureCitation>
). Rutrum medially with large number of irregularly arranged setae (paired in other species) (
<figureCitation id="1330678A7658FFDDFCF524072C98F9FE" box="[777,919,1637,1663]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="5" pageNumber="240">Figs 1718</figureCitation>
), pergula narrow, without setae. Rutrum with truncus portion of pergula very thin and narrower than stragulum (
<figureCitation id="1330678A7658FFDDFB0924EF2E5AF94E" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="5" pageNumber="240">Figs 1618</figureCitation>
). Ventral side of truncus near level of stragulum basis with sclerotised structure in shape of a balcony, armed with three sharp­pointed setae and with two ventral rows of conical protuberances (similar laterodorsal rows of protuberances present in
<taxonomicName id="4C0B008C7658FFDDFB7725672A2FF89F" box="[1163,1312,1797,1822]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="spinulatus">
<emphasis id="B97FA71D7658FFDDFB7725672A2FF89F" box="[1163,1312,1797,1822]" italics="true" pageId="5" pageNumber="240">F. spinulatus</emphasis>
</taxonomicName>
) (see
<bibRefCitation id="EF9A06FE7658FFDDFEB8254F2EFEF8C6" author="Martens" box="[324,497,1837,1863]" pageId="5" pageNumber="253" refString="Martens, J. (1988) Fissiphalliidae, a new family of South American laniatorean harvestmen (Arachnida: Opiliones). Zeitschrift fur Zoologische Systematik und Evolutionsforschung, 26 (2), 114 - 127." type="journal article" year="1988">Martens 1988</bibRefCitation>
: figs 1718). Truncus basis ventrally with another balcony­shaped structure and dorsally with a folded area (
<figureCitation id="1330678A7658FFDDFD1325372C76F8EE" box="[751,889,1877,1903]" captionStart="FIGURES 16 21" captionStartId="9.[264,383,1595,1619]" captionTargetBox="[310,1267,880,1587]" captionTargetId="figure@9.[265,1321,865,1588]" captionTargetPageId="9" captionText="FIGURES 16 21. Fissiphallius chicoi n. sp., male holotype (16 18); F. t u c u p i n. sp., male holotype (19 21). Penis (16, 19 dorsal view, 17, 20 lateral view, 18, 21 ventral view)." httpUri="https://zenodo.org/record/174074/files/figure.png" pageId="5" pageNumber="240">Figs 1618</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF742B87765BFFDEFEF4240F2EB4F96C" httpUri="https://zenodo.org/record/174072/files/figure.png" pageId="6" pageNumber="241" targetBox="[295,1298,332,1619]" targetPageId="6">
<paragraph id="8BB47B0F765BFFDEFEF4240F2EB4F96C" blockId="6.[264,1323,1645,1773]" pageId="6" pageNumber="241">
<emphasis id="B97FA71D765BFFDEFEF4240F2EC9F904" bold="true" box="[264,454,1645,1669]" pageId="6" pageNumber="241">FIGURES 914.</emphasis>
<taxonomicName id="4C0B008C765BFFDEFE2E240C2DA7F904" box="[466,680,1646,1669]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="chicoi" status="sp. nov.">
<emphasis id="B97FA71D765BFFDEFE2E240C2DA7F904" box="[466,680,1646,1669]" italics="true" pageId="6" pageNumber="241">Fissiphallius chicoi</emphasis>
</taxonomicName>
<emphasis id="B97FA71D765BFFDEFD48240F2DFAF904" bold="true" box="[692,757,1645,1669]" pageId="6" pageNumber="241">
<taxonomicNameLabel id="A24C1A66765BFFDEFD48240F2DFAF904" box="[692,757,1645,1669]" pageId="6" pageNumber="241" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (9, 1112);
<emphasis id="B97FA71D765BFFDEFBCA240C2B9BF904" box="[1078,1172,1646,1669]" italics="true" pageId="6" pageNumber="241">F. t u c u p i</emphasis>
<emphasis id="B97FA71D765BFFDEFB5F240F2BEBF904" bold="true" box="[1187,1252,1645,1669]" pageId="6" pageNumber="241">n. sp.</emphasis>
, male holotype (10, 1314). 910 right leg IV, lateral view; 1112 right pedipalp (11 ectal view, 12 mesal view); 1314 right pedipalp (13 ectal view, 14 mesal view). Scale bars = 1.0 mm (9, 1112), 0.5 mm (10, 1314).
</paragraph>
</caption>
<subSubSection id="C3112884765BFFDFFEC425422CD1FE2F" lastPageId="7" lastPageNumber="242" pageId="6" pageNumber="241" type="key">
<paragraph id="8BB47B0F765BFFDEFEC425422A27F8E3" blockId="6.[264,1321,1824,1890]" pageId="6" pageNumber="241">
<keyLead id="30FAD83A765BFFDEFEC425422A27F8E3" pageId="6" pageNumber="241">
Color (in 75% ethanol). Body yellowish, prosoma with reticulated brown stains (
<figureCitation id="1330678A765BFFDEFB0625422E19F8E3" captionStart="FIGURES 1 8" captionStartId="5.[264,383,1372,1396]" captionTargetBox="[265,1319,621,1329]" captionTargetId="figure@5.[239,1345,586,1347]" captionTargetPageId="5" captionText="FIGURES 1 8. Fissiphallius chicoi n. sp., male holotype (1, 3, 5, 7); Fissiphallius tucupi n. sp., male holotype (2, 4, 6, 8). Body (1 2 dorsal view, 3 4 lateral view, 5 6 ventral view, 7 8 posterior view). Scale bars = 0.1 mm (1, 3, 5, 7), 0.5 mm (2, 4, 6, 8)." httpUri="https://zenodo.org/record/174071/files/figure.png" pageId="6" pageNumber="241">Fig. 1</figureCitation>
). Lateral borders and median region of scutal areas IIV with brown stains, reticulated
</keyLead>
</paragraph>
<paragraph id="8BB47B0F765AFFDFFEF4237E2CD1FE2F" blockId="7.[264,1323,284,670]" pageId="7" pageNumber="242">brown stains in between them. Posterior margin of dorsal scutum and free tergites and sternites with brown lateral stains. Anal operculum brown, with yellow stain in the middle. Pedipalps, coxae and throchanters pale yellow, all other leg segments brown. Free sternites brown. Coxae IIV yellow, coxae IV with brown dorsal stain.</paragraph>
</subSubSection>
<subSubSection id="C3112884765AFFDFFEC423DE2CD0FD1F" pageId="7" pageNumber="242" type="description">
<paragraph id="8BB47B0F765AFFDFFEC423DE2CFAFE7F" blockId="7.[264,1323,284,670]" pageId="7" pageNumber="242">
Description of the female
<typeStatus id="54B0C5AD765AFFDFFD6D23DE2DF5FE57" box="[657,762,444,470]" pageId="7" pageNumber="242" type="paratype">paratype</typeStatus>
. Very similar to the male, different in the following: legs considerably shorter, tubercles of leg IV shorter.
</paragraph>
<paragraph id="8BB47B0F765AFFDFFEC4206E2D28FDF7" blockId="7.[264,1323,284,670]" pageId="7" pageNumber="242">Measurements. Total length: 3.0. Carapace region of scutum: 1.0 long, 1.2 wide. Scutum: 2.0 long, 2.0 maximum width. Pedipalp: 2.7. Legs: 5.9/4.1/5.2/3.1. Tarsal formula: 3(3), 5(3), 5, 5.</paragraph>
<paragraph id="8BB47B0F765AFFDFFEC420E62CD0FD1F" blockId="7.[264,1323,284,670]" box="[312,991,644,670]" pageId="7" pageNumber="242">
Distribution. Known only from the
<typeStatus id="54B0C5AD765AFFDFFD2520E62C03FD1F" box="[729,780,644,670]" pageId="7" pageNumber="242">type</typeStatus>
locality (
<figureCitation id="1330678A765AFFDFFC8620E62CDCFD1F" box="[890,979,644,670]" captionStart="FIGURE 22" captionStartId="14.[264,367,1755,1779]" captionTargetBox="[359,1230,953,1740]" captionTargetId="figure@14.[359,1231,953,1741]" captionTargetPageId="14" captionText="FIGURE 22. Northern South America showing the localities of the six nominal species of Fissiphallius. F. s t u r m i Martens, F. spinulatus Martens, F. sympatricus Martens (pentagon), F. martensi Pinto­da­Rocha (triangle), F. t u c u p i n. sp. (star), F. c h i c o i n. sp. (circle). Dotted lines = country boundaries, thin lines = rivers." httpUri="https://zenodo.org/record/174075/files/figure.png" pageId="7" pageNumber="242">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>