150 lines
15 KiB
XML
150 lines
15 KiB
XML
<document id="5824FD252086D94D435713792019B424" ID-DOI="10.11646/zootaxa.5403.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561687" ID-ZooBank="0DEB9EDB-4BC9-43F1-A025-DFF59726B738" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706103756776" checkinUser="plazi" docAuthor="Lukashevich, Elena Dmitrievna & Ansorge, Jörg" docDate="2024" docId="03A2AD6EE44E22274B9FF9D0EB2DBD79" docLanguage="en" docName="zootaxa.5403.1.8.pdf" docOrigin="Zootaxa 5403 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boholdoyidae Kovalev" docType="treatment" docVersion="2" lastPageNumber="122" masterDocId="FF9BD516E449222F4B08FFA5E801BC64" masterDocTitle="First Upper Triassic Diptera (Insecta) from Germany" masterLastPageNumber="129" masterPageNumber="115" pageNumber="122" updateTime="1706104313811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="08AD24791E3AE36B2E75E0E4D456218A">
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<mods:title id="0BE5B95EB2454C872BAD22B6F45771EC">First Upper Triassic Diptera (Insecta) from Germany</mods:title>
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</mods:titleInfo>
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<mods:name id="CC518FF01D161A46FD3F8C7B8DDC7D5C" type="personal">
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<mods:role id="CA0998D78DFDB2106FF50528483909DC">
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<mods:roleTerm id="8F43C23378951B40C3E0EAC43CA7F5CB">Author</mods:roleTerm>
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<mods:namePart id="2A39708627E1BA336776A781BABD9316">Lukashevich, Elena Dmitrievna</mods:namePart>
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<mods:affiliation id="BA255DAD21565C4904512FD989BF489A">Elena D. Lukashevich, Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="5369058BF72A7D9D10B621E89A38F40F" type="personal">
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<mods:roleTerm id="9333F5B4D8B9033A531B944F1066FB14">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1E2D9AF7CBDD68C258961E73AF8D5DAA">Ansorge, Jörg</mods:namePart>
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<mods:affiliation id="2E825A5763768872A3D35B93BC2F56DC">Jörg Ansorge, Institute of Geography and Geology, University of Greifswald, Friedrich-Ludwig-Jahn-Straβe 17 a, D- 17487 Greifswald, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="E801B9D4BB85696430784233B1CDF7DA">text</mods:typeOfResource>
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<mods:title id="3ADEE4D544FF42DEEFC09DDFB4CB8333">Zootaxa</mods:title>
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<mods:part id="52668B408A2E1ACB288CAAF6B57F9145">
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<mods:date id="F222BC0357E774C35B739A21B55610CF">2024</mods:date>
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<mods:number id="3192DBB15546D1AF90DB125BEE790CB2">2024-01-18</mods:number>
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<mods:start id="035F6B60F5D1D264A2FC483828CC5BEC">115</mods:start>
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<mods:url id="D1B73A5495F67215A065D3040F1AE2B8">http://dx.doi.org/10.11646/zootaxa.5403.1.8</mods:url>
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<mods:classification id="A2A4EFE4699028D6C376CDD3D38D2C6A">journal article</mods:classification>
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<mods:identifier id="F0451075B2D3004F44ABC6CB621ACAC6" type="DOI">10.11646/zootaxa.5403.1.8</mods:identifier>
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<mods:identifier id="9AB39C3A8914E2F7F605645DD928A27C" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C3114FF3E44E22284B9FF9D0EB81BAEB" box="[151,896,1653,1680]" pageId="7" pageNumber="122" type="nomenclature">
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<paragraph id="8BB41C78E44E22284B9FF9D0EB81BAEB" blockId="7.[151,896,1653,1680]" box="[151,896,1653,1680]" pageId="7" pageNumber="122">
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<heading id="D0FCAB14E44E22284B9FF9D0EB81BAEB" bold="true" box="[151,896,1653,1680]" fontSize="11" level="1" pageId="7" pageNumber="122" reason="1">
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<emphasis id="B97FC06AE44E22284B9FF9D0EB81BAEB" bold="true" box="[151,896,1653,1680]" pageId="7" pageNumber="122">
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Family
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<taxonomicName id="4C0B67FBE44E22284BFFF9D0EA11BAF4" ID-CoL="89NSJ" authority="Kovalev" authorityName="Kovalev" box="[247,528,1653,1680]" class="Insecta" family="Boholdoyidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="122" phylum="Arthropoda" rank="family">Boholdoyidae Kovalev</taxonomicName>
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in
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<bibRefCitation id="EF9A6189E44E2228493FF9D0EB81BAEB" author="Kalugina, N. S. & Kovalev, V. G." box="[567,896,1653,1680]" pageId="7" pageNumber="122" refId="ref7985" refString="Kalugina, N. S. & Kovalev, V. G. (1985) Dvukrylye nasekomye yury Sibiri [Jurassic Diptera of Siberia]. Nauka, Moscow, 198 pp." type="book" year="1985">Kalugina & Kovalev, 1985</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3114FF3E44E22284B9FF91BEB46BB44" pageId="7" pageNumber="122" type="type_taxon">
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<paragraph id="8BB41C78E44E22284B9FF91BEC85BABC" blockId="7.[151,1437,1725,2040]" box="[151,1156,1725,1758]" pageId="7" pageNumber="122">
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<materialsCitation id="3B631625E44E22284B9FF91BEC85BABC" box="[151,1156,1725,1758]" country="Russia" county="Transbaikalia" location="Boholdoyia Kovalev" municipality="Novospasskoye" pageId="7" pageNumber="122" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97FC06AE44E22284B9FF91BE925BABC" bold="true" box="[151,292,1726,1752]" pageId="7" pageNumber="122">
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<typeStatus id="54B0A2DAE44E22284B9FF91BE8D3BABC" box="[151,210,1726,1752]" pageId="7" pageNumber="122">Type</typeStatus>
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genus.
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</emphasis>
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<location id="8ED44AA3E44E22284A23F91BEA12BABC" LSID="urn:lsid:plazi:treatment:03A2AD6EE44E22274B9FF9D0EB2DBD79:8ED44AA3E44E22284A23F91BEA12BABC" box="[299,531,1726,1752]" country="Russia" county="Transbaikalia" municipality="Novospasskoye" name="Boholdoyia Kovalev" pageId="7" pageNumber="122">
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<emphasis id="B97FC06AE44E22284A23F91BE9ADBABC" box="[299,428,1726,1752]" italics="true" pageId="7" pageNumber="122">Boholdoyia</emphasis>
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Kovalev
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</location>
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, 1985 (
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<collectingMunicipality id="6BD08602E44E2228496DF91BEB18BABC" box="[613,793,1726,1752]" pageId="7" pageNumber="122">Novospasskoye</collectingMunicipality>
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,
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<collectingCounty id="62D564F4E44E2228482CF91BEBBFBABC" box="[804,958,1726,1752]" pageId="7" pageNumber="122">Transbaikalia</collectingCounty>
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,
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<collectingCountry id="F31C5CE8E44E222848C1F91BEC16BABC" box="[969,1047,1726,1752]" name="Russia" pageId="7" pageNumber="122">Russia</collectingCountry>
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, J
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<subScript id="178F1E3DE44E22284F25F975EC47BABA" attach="left" box="[1069,1094,1744,1758]" fontSize="6" pageId="7" pageNumber="122">1–2</subScript>
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– K
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<subScript id="178F1E3DE44E22284F7BF975EC7ABABA" attach="right" box="[1139,1147,1744,1758]" fontSize="6" pageId="7" pageNumber="122">1</subScript>
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)
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB41C78E44E22284BCFF944EB46BB44" blockId="7.[151,1437,1725,2040]" pageId="7" pageNumber="122">
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<emphasis id="B97FC06AE44E22284BCFF944E966BA9F" bold="true" box="[199,359,1761,1787]" pageId="7" pageNumber="122">Composition.</emphasis>
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<materialsCitation id="3B631625E44E22284A78F947EB42BB44" country="Russia" county="Transbaikalia" location="Daiamyi" municipality="Lower Cretaceous of Daya" pageId="7" pageNumber="122" specimenCount="1">
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Besides the
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<typeStatus id="54B0A2DAE44E22284AF3F947EA2ABA98" box="[507,555,1762,1788]" pageId="7" pageNumber="122">type</typeStatus>
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and the new genus, the monotypic
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<emphasis id="B97FC06AE44E222848CFF947EC26BA9F" box="[967,1063,1762,1787]" italics="true" pageId="7" pageNumber="122">
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<location id="8ED44AA3E44E222848CFF947EC26BA9F" LSID="urn:lsid:plazi:treatment:03A2AD6EE44E22274B9FF9D0EB2DBD79:8ED44AA3E44E222848CFF947EC26BA9F" box="[967,1063,1762,1787]" country="Russia" county="Transbaikalia" municipality="Lower Cretaceous of Daya" name="Daiamyi" pageId="7" pageNumber="122">Daiamyi</location>
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</emphasis>
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a
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<bibRefCitation id="EF9A6189E44E22284F35F947ECE3BA98" author="Kovalev, V. G." box="[1085,1250,1762,1788]" pageId="7" pageNumber="122" pagination="123 - 177" refId="ref8019" refString="Kovalev, V. G. (1990) Dvukrylye. Muscida [Flies. Muscida]. In: Rasnitsyn, A. P. (Ed.), Pozdnemezozoiskie nasekomye Vostochnogo Zabaikal'ya [Late Mesozoic insects of Eastern Transbaikalia]. Transactions of the Paleontological Institute of the USSR Academy of Sciences, 239, pp. 123 - 177." type="journal article" year="1990">Kovalev, 1990</bibRefCitation>
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from the
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<location id="8ED44AA3E44E22284E5EF947E8F1BB44" LSID="urn:lsid:plazi:treatment:03A2AD6EE44E22274B9FF9D0EB2DBD79:8ED44AA3E44E22284E5EF947E8F1BB44" country="Russia" county="Transbaikalia" municipality="Lower Cretaceous of Daya" name="Upper Jurassic" pageId="7" pageNumber="122">Upper Jurassic</location>
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or
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<collectingMunicipality id="6BD08602E44E22284A1DF8A3EA44BB44" box="[277,581,1798,1824]" pageId="7" pageNumber="122">Lower Cretaceous of Daya</collectingMunicipality>
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,
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<collectingCounty id="62D564F4E44E22284958F8A3EAEBBB44" box="[592,746,1798,1824]" pageId="7" pageNumber="122">Transbaikalia</collectingCounty>
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,
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<collectingCountry id="F31C5CE8E44E222849FDF8A3EB42BB44" box="[757,835,1798,1824]" name="Russia" pageId="7" pageNumber="122">Russia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3114FF3E44E22274BCFF88FEB2DBD79" lastPageId="8" lastPageNumber="123" pageId="7" pageNumber="122" type="discussion">
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<paragraph id="8BB41C78E44E22274BCFF88FEB2DBD79" blockId="7.[151,1437,1725,2040]" lastBlockId="8.[151,1437,151,291]" lastPageId="8" lastPageNumber="123" pageId="7" pageNumber="122">
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<emphasis id="B97FC06AE44E22284BCFF88FE93ABB20" bold="true" box="[199,315,1834,1860]" pageId="7" pageNumber="122">Remarks.</emphasis>
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V. Kovalev described the family based on
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<specimenCount id="9D0DD7F1E44E2228481FF88FEB7DBB20" box="[791,892,1834,1860]" count="1" pageId="7" pageNumber="122" type="male">one male</specimenCount>
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with badly preserved wings and the diagnosis of the family included a lot of body characters (Kovalev in
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<bibRefCitation id="EF9A6189E44E2228480CF8EBEC2DBB0C" author="Kalugina, N. S. & Kovalev, V. G." box="[772,1068,1869,1896]" pageId="7" pageNumber="122" refId="ref7985" refString="Kalugina, N. S. & Kovalev, V. G. (1985) Dvukrylye nasekomye yury Sibiri [Jurassic Diptera of Siberia]. Nauka, Moscow, 198 pp." type="book" year="1985">Kalugina & Kovalev 1985</bibRefCitation>
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). Later
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<specimenCount id="9D0DD7F1E44E22284F76F8EBECEEBB0C" box="[1150,1263,1870,1896]" count="2" pageId="7" pageNumber="122" type="male">two males</specimenCount>
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were described from other Mesozoic deposits and the venation of the family became clear (
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<bibRefCitation id="EF9A6189E44E222848D5F8D7EC81BBE8" author="Kovalev, V. G." box="[989,1152,1906,1932]" pageId="7" pageNumber="122" pagination="123 - 177" refId="ref8019" refString="Kovalev, V. G. (1990) Dvukrylye. Muscida [Flies. Muscida]. In: Rasnitsyn, A. P. (Ed.), Pozdnemezozoiskie nasekomye Vostochnogo Zabaikal'ya [Late Mesozoic insects of Eastern Transbaikalia]. Transactions of the Paleontological Institute of the USSR Academy of Sciences, 239, pp. 123 - 177." type="journal article" year="1990">Kovalev 1990</bibRefCitation>
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). The new Upper Triassic isolated wings from
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<collectingCountry id="F31C5CE8E44E22284A77F833E9E6BBD4" box="[383,487,1942,1968]" name="Germany" pageId="7" pageNumber="122">Germany</collectingCountry>
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share the same pattern of venation. The following characters, visible in the Triassic wings, are diagnostic for the family (so they are not mentioned in the diagnosis of the new genus below): C extends along entire wing, strongly thickened at the anterior and distal margin, weak but distinct along the posterior margin, Sc weak, shorter than half of the wing, R
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<subScript id="178F1E3DE44122274988FF0CEA89BCD3" attach="left" box="[640,648,169,183]" fontSize="6" pageId="8" pageNumber="123">1</subScript>
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ends about wing midlength, Rs stem long, Rs with three long free branches, R
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<subScript id="178F1E3DE44122274A14FF68E934BCBF" attach="left" box="[284,309,205,219]" fontSize="6" pageId="8" pageNumber="123">2+3</subScript>
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begins proximal to
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<emphasis id="B97FC06AE4412227491DFF1EEA3CBCB0" box="[533,573,187,212]" italics="true" pageId="8" pageNumber="123">r-m</emphasis>
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and ends in the wing margin far from R
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<subScript id="178F1E3DE44122274F0CFF68EC0DBCBF" attach="left" box="[1028,1036,205,219]" fontSize="6" pageId="8" pageNumber="123">1</subScript>
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, R
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<subScript id="178F1E3DE44122274F25FF68EC46BCBF" attach="left" box="[1069,1095,205,219]" fontSize="6" pageId="8" pageNumber="123">4+5</subScript>
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fork several times longer than distal portion of R
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<subScript id="178F1E3DE44122274A6DFF54E97EBC9B" attach="left" box="[357,383,241,255]" fontSize="6" pageId="8" pageNumber="123">4+5</subScript>
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stem, R
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<subScript id="178F1E3DE44122274AD4FF54E9E5BC9B" attach="left" box="[476,484,241,255]" fontSize="6" pageId="8" pageNumber="123">5</subScript>
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ends in the wing apex, basal cells closed;
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<emphasis id="B97FC06AE441222748C9FF7AEBDCBC9D" box="[961,989,223,249]" italics="true" pageId="8" pageNumber="123">ba</emphasis>
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longer and narrower than
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<emphasis id="B97FC06AE44122274E0FFF7AED22BC9D" box="[1287,1315,223,249]" italics="true" pageId="8" pageNumber="123">bp</emphasis>
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, ends near wing midlength, three medial veins, M
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<subScript id="178F1E3DE44122274942FEB0EA62BD47" attach="left" box="[586,611,277,291]" fontSize="6" pageId="8" pageNumber="123">1+2</subScript>
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fork distal to
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<emphasis id="B97FC06AE441222749F6FEA6EB27BD78" box="[766,806,259,284]" italics="true" pageId="8" pageNumber="123">r-m</emphasis>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |