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<document id="887B1B760B2E4E43B33FA6FED8300E69" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CA22FFABEEE4FD11426AFC90" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Ceriagrion banditum Kipping &amp; Dijkstra, sp. nov." docType="treatment" docVersion="21" lastPageNumber="510" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="504" updateTime="1695001031402" updateUser="admin">
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<mods:title id="3BDB3971F3711FCD25A1904FEE686B19">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
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<mods:namePart id="FDC38164D2549B1CE023CB41E3E71136">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="D0AFFCED0338BDA74111C1DD17EDF0DD">Kipping, Jens</mods:namePart>
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<mods:namePart id="AFB4E717533260C464E50F9D271F4CF9">Mézière, Nicolas</mods:namePart>
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<mods:title id="89D72EEE88A3DED747774F7286294779">Odonatologica</mods:title>
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<treatment id="03A25264CA22FFABEEE4FD11426AFC90" ID-DOI="http://doi.org/10.5281/zenodo.5640220" ID-GBIF-Taxon="114923421" ID-Zenodo-Dep="5640220" LSID="urn:lsid:zoobank.org:act:03A25264-CA22-FFAB-EEE4-FD11426AFC90" httpUri="http://treatment.plazi.org/id/03A25264CA22FFABEEE4FD11426AFC90" lastPageId="65" lastPageNumber="510" pageId="59" pageNumber="504">
<subSubSection id="C311B0F9CA22FFD1EEE4FD114231FCF0" pageId="59" pageNumber="504" type="nomenclature">
<paragraph id="8BB4E372CA22FFD1EEE4FD114231FCF0" pageId="59" pageNumber="504">
<heading id="D0FC541ECA22FFD1EEE4FD114231FCF0" bold="true" centered="true" fontSize="11" level="2" pageId="59" pageNumber="504" reason="4">
<emphasis id="B97F3F60CA22FFD1EEE4FD1142E1FCA5" bold="true" box="[198,996,764,792]" pageId="59" pageNumber="504">
<taxonomicName id="4C0B98F1CA22FFD1EEE4FD1143A6FCAA" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CA22-FFAB-EEE4-FD11426AFC90" authority="Kipping &amp; Dijkstra" authorityName="Kipping &amp; Dijkstra" box="[198,675,764,792]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="banditum" status="sp. nov.">
<emphasis id="B97F3F60CA22FFD1EEE4FD1140C7FCA5" bold="true" box="[198,450,764,792]" italics="true" pageId="59" pageNumber="504">Ceriagrion banditum</emphasis>
Kipping &amp; Dijkstra
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA22FFD1EC8BFD114203FCA5" box="[681,774,764,791]" pageId="59" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
Band-eyed Citril
</emphasis>
(
<typeStatus id="54B05DD0CA22FFD1EF5FFCCA40BDFCF0" box="[381,440,807,834]" pageId="59" pageNumber="504">Type</typeStatus>
<figureCitation id="1330FFF7CA22FFD1EF9CFCCB4328FCF3" box="[446,557,806,833]" captionStart="Photo 15" captionStartId="43.[782,848,1384,1408]" captionText="Photo 15. Sandy and clear stream on the Batéké Plateau in Gabon. Habitat of Elattoneura aurifex. Photo: A. Günther (27 - i- 2012)" pageId="59" pageNumber="504">Photo 15</figureCitation>
,
<figureCitation id="1330FFF7CA22FFD1EC1AFCCB43D6FCF3" box="[568,723,806,833]" captionStart-0="Photo 21" captionStart-1="Photo 22" captionStartId-0="64.[159,225,1030,1055]" captionStartId-1="64.[159,225,1290,1314]" captionTargetBox-0="[158,1035,461,989]" captionTargetId-0="figure@64.[156,1038,458,991]" captionTargetPageId-0="64" captionText-0="Photo 21. Ceriagrion banditum, holotype male; Lake Chila, Mbala, Zambia. Photo: JK (08 - xii- 2014)" captionText-1="Photo 22. Shallow temporary pools close to the Mansha River at Shiwa Ngandu, Zambia. Habitat of Ceriagrion banditum, C. junceum and some other Ceriagrion. Photo: JK (05 - - xii- 2014)" pageId="59" pageNumber="504">Photo 2122</figureCitation>
,
<figureCitation id="1330FFF7CA22FFD1ECFFFCCA4229FCF0" box="[733,812,807,834]" captionStart="Figure 10" captionStartId="61.[159,227,1468,1493]" captionTargetBox="[159,1035,733,1427]" captionTargetId="figure@61.[156,1038,730,1430]" captionTargetPageId="61" captionText="Figure 10. Male appendages in dorsal (left) and lateral view (right) of suave - complex of Ceriagrion, including the holotype of C. moorei currently treated as a synonym of C. suave, as well as penis in lateral view (far left) of two related species." pageId="59" pageNumber="504">Fig. 10</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA22FFD1EEBDFCB8401AFCDD" box="[159,287,853,879]" pageId="59" pageNumber="504" type="description">
<paragraph id="8BB4E372CA22FFD1EEBDFCB8401AFCDD" box="[159,287,853,879]" pageId="59" pageNumber="504">
<heading id="D0FC541ECA22FFD1EEBDFCB8401AFCDD" bold="true" box="[159,287,853,879]" fontSize="11" level="3" pageId="59" pageNumber="504" reason="6">
<emphasis id="B97F3F60CA22FFD1EEBDFCB8401AFCDD" bold="true" box="[159,287,853,879]" pageId="59" pageNumber="504">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA22FFD5EEBDFC97421EFBB0" lastPageId="63" lastPageNumber="508" pageId="59" pageNumber="504" type="materials_examined">
<paragraph id="8BB4E372CA22FFD4EEBDFC9742F7FE4A" lastPageId="62" lastPageNumber="507" pageId="59" pageNumber="504">
<bibRefCitation id="EFFC8073219A4EBA17D53A61ADCB65C4" author="Dijkstra K. - D. B." journalOrPublisher="Journal of Afrotropical Zoology" pagination="3 - 14" part="2" refId="ref66743" refString="Dijkstra K. - D. B. 2005 a. A review of continental Afrotropical Ceriagrion (Odonata: Coenagrionidae). Journal of Afrotropical Zoology 2: 3 - 14" title="A review of continental Afrotropical Ceriagrion (Odonata: Coenagrionidae)" type="journal article" year="2005">Dijkstra (2005 a)</bibRefCitation>
reviewed the
<taxonomicName id="4C0B98F1CA22FFD1EC3EFC964359FC26" box="[540,604,891,916]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA22FFD1EC3EFC964359FC26" box="[540,604,891,916]" italics="true" pageId="59" pageNumber="504">suave</emphasis>
</taxonomicName>
-complex of
<taxonomicName id="4C0B98F1CA22FFD1ECD0FC97426BFC26" box="[754,878,890,916]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA22FFD1ECD0FC97426BFC26" box="[754,878,890,916]" italics="true" pageId="59" pageNumber="504">Ceriagrion</emphasis>
</taxonomicName>
and suggest- ed that various taxa may be synonymous. However, up to four superficially similar taxa that differ clearly genetically and subtly in ecology, coloration and morphology overlap in southern central Africa (
<figureCitation id="1330FFF7CA22FFD1ECDBFC074241FBB6" box="[761,836,1002,1028]" captionStart="Tree 2. 50" captionStartId="60.[159,207,1440,1464]" captionTargetBox="[159,1033,406,1398]" captionTargetId="figure@60.[156,1038,406,1400]" captionTargetPageId="60" captionText="Tree 2. 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis for the suave - complex of Ceriagrion, related species and two outgroup taxa. For relevant branches posterior probabilities from Bayesian inference analysis are shown first, as is bootstrap support (both as percentages)." pageId="59" pageNumber="504">Tree 2</figureCitation>
). Additional taxa may occur elsewhere: western African specimens appear similar to
<taxonomicName id="4C0B98F1CA22FFD1ED8EFBE2400FFBFD" authority="Ris, 1921" authorityName="Ris" authorityYear="1921" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA22FFD1ED8EFBE2450EFB98" box="[940,1035,1039,1066]" italics="true" pageId="59" pageNumber="504">C.suave</emphasis>
Ris, 1921
</taxonomicName>
but are slightly distinct morphologically (
<figureCitation id="1330FFF7CA22FFD1ECDBFBD9424FFBFD" box="[761,842,1076,1103]" captionStart="Figure 10" captionStartId="61.[159,227,1468,1493]" captionTargetBox="[159,1035,733,1427]" captionTargetId="figure@61.[156,1038,730,1430]" captionTargetPageId="61" captionText="Figure 10. Male appendages in dorsal (left) and lateral view (right) of suave - complex of Ceriagrion, including the holotype of C. moorei currently treated as a synonym of C. suave, as well as penis in lateral view (far left) of two related species." pageId="59" pageNumber="504">Fig. 10</figureCitation>
) and genetically (
<figureCitation id="1330FFF7CA22FFD1EE85FBB741F0FBC6" box="[167,245,1114,1140]" captionStart="Tree 2. 50" captionStartId="60.[159,207,1440,1464]" captionTargetBox="[159,1033,406,1398]" captionTargetId="figure@60.[156,1038,406,1400]" captionTargetPageId="60" captionText="Tree 2. 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis for the suave - complex of Ceriagrion, related species and two outgroup taxa. For relevant branches posterior probabilities from Bayesian inference analysis are shown first, as is bootstrap support (both as percentages)." pageId="59" pageNumber="504">Tree 2</figureCitation>
). These could represent
<taxonomicName id="4C0B98F1CA22FFD1EC3AFBB7424FFBC6" authority="Longfield, 1952" authorityName="Longfield" authorityYear="1952" box="[536,842,1114,1140]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B97F3F60CA22FFD1EC3AFBB7438FFBC6" box="[536,650,1114,1140]" italics="true" pageId="59" pageNumber="504">C. moorei</emphasis>
Longfield, 1952
</taxonomicName>
of which
<taxonomicName id="4C0B98F1CA22FFD1EDE0FBB74070FB28" authority="Fraser, 1955" authorityName="Fraser" authorityYear="1955" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="hamoni">
<emphasis id="B97F3F60CA22FFD1EDE0FBB741DFFB28" italics="true" pageId="59" pageNumber="504">C. hamoni</emphasis>
Fraser, 1955
</taxonomicName>
is possibly a synonym. Marshall &amp; Gambles (1977) separated
<taxonomicName id="4C0B98F1CA22FFD1EF37FB484070FB0D" box="[277,373,1189,1215]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA22FFD1EF37FB484070FB0D" box="[277,373,1189,1215]" italics="true" pageId="59" pageNumber="504">C. suave</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA22FFD1EF8DFB484325FB0D" box="[431,544,1189,1215]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B97F3F60CA22FFD1EF8DFB484325FB0D" box="[431,544,1189,1215]" italics="true" pageId="59" pageNumber="504">C. moorei</emphasis>
</taxonomicName>
from northern Ghana by the penis shape, but while this suggests that similar taxa overlap there too, whether their interpretation agrees with
<typeStatus id="54B05DD0CA22FFD1EFFAFB1D430EFAB8" box="[472,523,1264,1290]" pageId="59" pageNumber="504">type</typeStatus>
material must be investigated.
<bibRefCitation id="FBE1E0EE0CEEC15FD54357AD1130630D" author="Dijkstra K. - D. B. &amp; Clausnitzer V." journalOrPublisher="Royal Museum for Central Africa, Tervuren" refId="ref66897" refString="Dijkstra K. - D. B. &amp; Clausnitzer V. 2014. The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe. Royal Museum for Central Africa, Tervuren" title="The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe" type="book" year="2014">Dijkstra &amp; Clausnitzer (2014)</bibRefCitation>
also provided insufficient information to distinguish the various taxa in the complex, which they separated from
<taxonomicName id="4C0B98F1CA22FFD1EDAEFAD74053FAC8" authority="Campion, 1914" authorityName="Campion" authorityYear="1914" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="ignitum">
<emphasis id="B97F3F60CA22FFD1EDAEFAD74509FAE6" box="[908,1036,1338,1364]" italics="true" pageId="59" pageNumber="504">C. ignitum</emphasis>
Campion, 1914
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA22FFD1EFB4FA8D43D6FAC8" authority="Ris, 1924" authorityName="Ris" authorityYear="1924" box="[406,723,1375,1402]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="59" pageNumber="504" phylum="Arthropoda" rank="species" species="kordofanicum">
<emphasis id="B97F3F60CA22FFD1EFB4FA8D435FFAC8" box="[406,602,1375,1402]" italics="true" pageId="59" pageNumber="504">C. kordofanicum</emphasis>
Ris, 1924
</taxonomicName>
only by (a) the moderate size, Hw 17.023.0 mm rather than 15.017.0 mm; although (b) the greenish rather than deep reddish eyes with maturity are also diagnostic. All species further have (c) rather dull orange to brown dorsa of the head and thorax; (d) uniformly orange to red abdomen; (e) apically pointed penis with acute or finger-like lateral lobes; (f) no toothed processes on the apical border of S 10; and (g) paraprocts that do not or barely reach beyond the cerci and that have a fairly distinct angular heel ventrally. True
<taxonomicName id="4C0B98F1CA25FFD6ED2DFEA5426AFED0" box="[783,879,328,354]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="60" pageNumber="505" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA25FFD6ED2DFEA5426AFED0" box="[783,879,328,354]" italics="true" pageId="60" pageNumber="505">C. suave</emphasis>
</taxonomicName>
favours temporary pools such as rain puddles and, as confirmed with the
<typeStatus id="54B05DD0CA25FFD6ED53FE8042DDFE3A" box="[881,984,365,392]" pageId="60" pageNumber="505">holotype</typeStatus>
and a
<typeStatus id="54B05DD0CA24FFD7EE91FF13401CFEAA" box="[179,281,254,280]" pageId="61" pageNumber="506">paratype</typeStatus>
in
<collectionCode id="ED1A7BB7CA24FFD7EF1DFF134090FEAA" box="[319,405,254,280]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" pageId="61" pageNumber="506">MRAC</collectionCode>
, is (1) notably uniform in colour, brown to orange on the head and thorax grading to yellow on the face and flanks, and typically without a darker greyish or greenish tinge to the head dorsum, nor with distinctly whitish thoracic flanks and/or blackish dots on the humeral and metapleural sutures; (2) often has largely yellow-stained wings; (3) the penis does not have drawn-out finger-like lateral lobes; (4) the dorsal excision of the apical border of S 10 is up to half as deep as the segment is dorsally long, and its borders are pale; (5) the cerci have the apical black teeth turned downward and thus usually not visible in dorsal view; and (6) the tips of the paraprocts reach at most as far as those (or even the apical teeth) of the cerci (
<figureCitation id="1330FFF7CA24FFD7EEC4FD9F403CFD3F" box="[230,313,626,653]" captionStart="Figure 10" captionStartId="61.[159,227,1468,1493]" captionTargetBox="[159,1035,733,1427]" captionTargetId="figure@61.[156,1038,730,1430]" captionTargetPageId="61" captionText="Figure 10. Male appendages in dorsal (left) and lateral view (right) of suave - complex of Ceriagrion, including the holotype of C. moorei currently treated as a synonym of C. suave, as well as penis in lateral view (far left) of two related species." pageId="61" pageNumber="506">Fig. 10</figureCitation>
). The
<taxonomicName id="4C0B98F1CA24FFD7EFA2FD9E40BAFD3F" box="[384,447,627,653]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="61" pageNumber="506" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA24FFD7EFA2FD9E40BAFD3F" box="[384,447,627,653]" italics="true" pageId="61" pageNumber="506">suave</emphasis>
</taxonomicName>
-complex species similarity stems from the shared lack of conspicuous characters rather than from common ancestry (
<figureCitation id="1330FFF7CA24FFD7ED93FD7542F8FD00" box="[945,1021,664,690]" captionStart="Tree 2. 50" captionStartId="60.[159,207,1440,1464]" captionTargetBox="[159,1033,406,1398]" captionTargetId="figure@60.[156,1038,406,1400]" captionTargetPageId="60" captionText="Tree 2. 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis for the suave - complex of Ceriagrion, related species and two outgroup taxa. For relevant branches posterior probabilities from Bayesian inference analysis are shown first, as is bootstrap support (both as percentages)." pageId="61" pageNumber="506">Tree 2</figureCitation>
): ecology, genetics, coloration and appendages and penis shape indicate that
<taxonomicName id="4C0B98F1CA27FFD4EEBDFECE41F8FE8F" box="[159,253,291,317]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="507" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA27FFD4EEBDFECE41F8FE8F" box="[159,253,291,317]" italics="true" pageId="62" pageNumber="507">C. suave</emphasis>
</taxonomicName>
is very close to the ubiquitous
<taxonomicName id="4C0B98F1CA27FFD4EC42FECE42C4FE8F" authority="Burmeister, 1839" authorityName="Burmeister" authorityYear="1839" box="[608,961,290,317]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="507" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="B97F3F60CA27FFD4EC42FECE43E5FE8F" box="[608,736,290,317]" italics="true" pageId="62" pageNumber="507">C. glabrum</emphasis>
(Burmeister, 1839)
</taxonomicName>
, while two new species described here are nearer
<taxonomicName id="4C0B98F1CA27FFD4EC82FEA542A1FED0" authority="Fraser, 1941" authorityName="Fraser" authorityYear="1941" box="[672,932,328,354]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="507" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B97F3F60CA27FFD4EC82FEA5420EFED0" box="[672,779,328,354]" italics="true" pageId="62" pageNumber="507">C. bakeri</emphasis>
Fraser, 1941
</taxonomicName>
but lack its bright blue head and thorax. Known only from the coastal plain of Mozambique and Tanzania, the similarly dull
<taxonomicName id="4C0B98F1CA27FFD4ECABFE7E429EFE1F" authority="Pinhey, 1969" authorityName="Pinhey" authorityYear="1969" box="[649,923,402,429]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="507" phylum="Arthropoda" rank="species" species="mourae">
<emphasis id="B97F3F60CA27FFD4ECABFE7E4205FE1F" box="[649,768,403,429]" italics="true" pageId="62" pageNumber="507">C. mourae</emphasis>
<bibRefCitation id="40084D30F8276CE5361CAFBBA698F61C" author="Pinhey E. C. G." journalOrPublisher="Arnoldia, Rhodesia" pagination="1 - 11" part="4" refId="ref68752" refString="Pinhey E. C. G. 1969. On the genus Umma Kirby (Odonata). Arnoldia, Rhodesia, 4: 1 - 11" title="On the genus Umma Kirby (Odonata)" type="journal article" year="1969">Pinhey, 1969</bibRefCitation>
</taxonomicName>
differs by having 10 rather than 1214 Fw postnodal cross-veins, the rectangular rath- er than trapezoidal Pt, and in the shape of the male appendages (
<figureCitation id="1330FFF7CA27FFD4EDB3FE3042E1FE4A" box="[913,996,477,504]" captionStart="Figure 10" captionStartId="61.[159,227,1468,1493]" captionTargetBox="[159,1035,733,1427]" captionTargetId="figure@61.[156,1038,730,1430]" captionTargetPageId="61" captionText="Figure 10. Male appendages in dorsal (left) and lateral view (right) of suave - complex of Ceriagrion, including the holotype of C. moorei currently treated as a synonym of C. suave, as well as penis in lateral view (far left) of two related species." pageId="62" pageNumber="507">Fig. 10</figureCitation>
).
</paragraph>
<caption id="DF74B3FACA25FFD6EEBDFA4D425BF9A3" pageId="60" pageNumber="505" targetBox="[159,1033,406,1398]" targetPageId="60">
<paragraph id="8BB4E372CA25FFD6EEBDFA4D425BF9A3" pageId="60" pageNumber="505">
<emphasis id="B97F3F60CA25FFD6EEBDFA4D41E9FA0A" bold="true" box="[159,236,1439,1464]" pageId="60" pageNumber="505">Tree 2.</emphasis>
50% majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis for the
<taxonomicName id="4C0B98F1CA25FFD6EF72FA50408BFA67" box="[336,398,1469,1493]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="60" pageNumber="505" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA25FFD6EF72FA50408BFA67" box="[336,398,1469,1493]" italics="true" pageId="60" pageNumber="505">suave</emphasis>
</taxonomicName>
-complex of
<taxonomicName id="4C0B98F1CA25FFD6EC3AFA50438DFA67" box="[536,648,1469,1493]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="60" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA25FFD6EC3AFA50438DFA67" box="[536,648,1469,1493]" italics="true" pageId="60" pageNumber="505">Ceriagrion</emphasis>
</taxonomicName>
, related species and two outgroup taxa. For relevant branches posterior probabilities from Bayesian inference analysis are shown first, as is bootstrap support (both as percentages).
</paragraph>
</caption>
<caption id="DF74B3FACA24FFD7EEBDFA5142F6F9BD" pageId="61" pageNumber="506" targetBox="[159,1035,733,1427]" targetPageId="61">
<paragraph id="8BB4E372CA24FFD7EEBDFA5142F6F9BD" pageId="61" pageNumber="506">
<emphasis id="B97F3F60CA24FFD7EEBDFA514009FA67" bold="true" box="[159,268,1468,1493]" pageId="61" pageNumber="506">Figure 10.</emphasis>
Male appendages in dorsal (left) and lateral view (right) of
<taxonomicName id="4C0B98F1CA24FFD7EDACFA5142C8FA67" box="[910,973,1468,1493]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="61" pageNumber="506" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA24FFD7EDACFA5142C8FA67" box="[910,973,1468,1493]" italics="true" pageId="61" pageNumber="506">suave</emphasis>
</taxonomicName>
-complex of
<taxonomicName id="4C0B98F1CA24FFD7EECEFA374059FA40" box="[236,348,1498,1522]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="61" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA24FFD7EECEFA374059FA40" box="[236,348,1498,1522]" italics="true" pageId="61" pageNumber="506">Ceriagrion</emphasis>
</taxonomicName>
, including the holotype of
<taxonomicName id="4C0B98F1CA24FFD7EC59FA3743E4FA40" box="[635,737,1498,1522]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="61" pageNumber="506" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B97F3F60CA24FFD7EC59FA3743E4FA40" box="[635,737,1498,1522]" italics="true" pageId="61" pageNumber="506">C. moorei</emphasis>
</taxonomicName>
currently treated as a synonym of
<taxonomicName id="4C0B98F1CA24FFD7EED2FA1A404CF9BD" box="[240,329,1527,1551]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="61" pageNumber="506" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA24FFD7EED2FA1A404CF9BD" box="[240,329,1527,1551]" italics="true" pageId="61" pageNumber="506">C. suave</emphasis>
</taxonomicName>
, as well as penis in lateral view (far left) of two related species.
</paragraph>
</caption>
<paragraph id="8BB4E372CA27FFD4EEBDFDC5406DFDF1" box="[159,360,552,579]" pageId="62" pageNumber="507">
<heading id="D0FC541ECA27FFD4EEBDFDC5406DFDF1" bold="true" box="[159,360,552,579]" fontSize="11" level="3" pageId="62" pageNumber="507" reason="6">
<emphasis id="B97F3F60CA27FFD4EEBDFDC5406DFDF1" bold="true" box="[159,360,552,579]" pageId="62" pageNumber="507">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA27FFD5EEBDFDA3421EFBB0" lastPageId="63" lastPageNumber="508" pageId="62" pageNumber="507">
<materialsCitation id="3B63E92FCA27FFD4EEBDFDA342C3FD00" ID-GBIF-Occurrence="1224547715" collectingDate="2014-12-08" collectionCode="RMNH" collectorName="J. Kipping" country="Zambia" elevation="1619" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF422-15" latitude="-8.8341" location="Lake Chila near Mbala" longLatPrecision="7" longitude="31.3948" pageId="62" pageNumber="507" specimenCode="RMNH.INS.508068" specimenCount="1" specimenCount-male="1" stateProvince="Northern Province" typeStatus="holotype">
<typeStatus id="54B05DD0CA27FFD4EEBDFDA34015FDDA" box="[159,272,590,616]" pageId="62" pageNumber="507">
<emphasis id="B97F3F60CA27FFD4EEBDFDA34015FDDA" bold="true" box="[159,272,590,616]" pageId="62" pageNumber="507">Holotype</emphasis>
</typeStatus>
<emphasis id="B97F3F60CA27FFD4EF3BFDA04028FDD5" bold="true" box="[281,301,589,615]" pageId="62" pageNumber="507">
<specimenCount id="52978F8F43F7EDE3826BE10D9E8AB886" type="male"></specimenCount>
</emphasis>
<emphasis id="B97F3F60CA27FFD4EF0FFDA34031FDDA" bold="true" box="[301,308,590,616]" pageId="62" pageNumber="507">.</emphasis>
<specimenCode id="26D62584EE41F57175F9EF525F97C1D8">RMNH.INS.508068</specimenCode>
,
<collectingCountry id="F31CA3E2CA27FFD4EC16FDA04395FDDA" box="[564,656,589,616]" name="Zambia" pageId="62" pageNumber="507" value="Zambia">Zambia</collectingCountry>
,
<collectingRegion id="49CF2D90CA27FFD4ECBFFDA0427BFDDA" box="[669,894,589,616]" pageId="62" pageNumber="507">Northern Province</collectingRegion>
,
<location id="C5B1923F3C1F1003BA4AFC7F77C3F033" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:C5B1923F3C1F1003BA4AFC7F77C3F033" country="Zambia" latitude="-8.8341" longLatPrecision="7" longitude="31.3948" name="Lake Chila near Mbala" stateProvince="Northern Province">Lake Chila near Mbala</location>
, boggy eastern lake shore, at shallow grassy pools,
<elevation id="A9AC3FE564EE3A73B1BC62FE0EFBB2A1" box="[894,978,627,653]" metricMagnitude="3" metricUnit="m" metricValue="1.619" pageId="62" pageNumber="507" unit="m" value="1619.0">
<quantity id="42469A29FC7A00477D8ADC90B0CDE19D" box="[894,978,627,653]" metricMagnitude="3" metricUnit="m" metricValue="1.619" pageId="62" pageNumber="507" unit="m" value="1619.0">1619 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA27FFD4EE85FD75400BFD00" box="[167,270,664,690]" direction="south" orientation="latitude" pageId="62" pageNumber="507" precision="5" value="-8.8341">8.8341 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA27FFD4EF37FD75408AFD00" box="[277,399,664,690]" direction="east" orientation="longitude" pageId="62" pageNumber="507" precision="5" value="31.3948">31.3948 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA27FFD4EF80FD75432EFD00" box="[418,555,664,690]" pageId="62" pageNumber="507" value="2014-12-08">
<collectingDate id="6BD18F18995FD154BA6F955C89695579" box="[418,555,664,690]" pageId="62" pageNumber="507" value="2014-12-08">08-xii- 2014</collectingDate>
</date>
, leg.
<collectorName id="E9A912890F97A3B13DDA2E003CCEC1E7">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA27FFD4ECC5FD744245FD00" box="[743,832,665,690]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="62" pageNumber="507">RMNH</collectionCode>
(
<figureCitation id="1330FFF7CA27FFD4ED6DFD7542B8FD00" box="[847,957,664,690]" captionStart="Photo 21" captionStartId="64.[159,225,1030,1055]" captionTargetBox="[158,1035,461,989]" captionTargetId="figure@64.[156,1038,458,991]" captionTargetPageId="64" captionText="Photo 21. Ceriagrion banditum, holotype male; Lake Chila, Mbala, Zambia. Photo: JK (08 - xii- 2014)" pageId="62" pageNumber="507">Photo 21</figureCitation>
)
</materialsCitation>
.
<emphasis id="B97F3F60CA27FFD4EEBDFD534077FD6A" bold="true" box="[159,370,702,728]" pageId="62" pageNumber="507">
<location id="17A30E198EFF6E9A074BA0FF68076CC8" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:17A30E198EFF6E9A074BA0FF68076CC8" country="Zambia" name="Further" stateProvince="Northern Province">Further</location>
material.
</emphasis>
<materialsCitation id="3B63E92FCA27FFD5EF5BFD5342DBFEAA" ID-GBIF-Occurrence="1224547722" collectingDate="2001-12-27" collectingDateMax="2001-12-29" collectingDateMin="2001-12-27" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="Malawi" elevation="1250" lastPageId="63" lastPageNumber="508" latitude="-14.26" location="SW of Lilongwe" longLatPrecision="558" longitude="33.455" pageId="62" pageNumber="507" specimenCount="2" specimenCount-male="2" stateProvince="Central Region">
<collectingCountry id="F31CA3E2CA27FFD4EF5BFD5340EAFD6A" box="[377,495,702,728]" name="Malawi" pageId="62" pageNumber="507" value="Malawi">MALAWI</collectingCountry>
(
<collectingRegion id="49CF2D90CA27FFD4EFDCFD5043B2FD6A" box="[510,695,701,728]" pageId="62" pageNumber="507">Central Region</collectingRegion>
):
<specimenCount id="9D0D28FBCA27FFD4ECEBFD5343E9FD65" box="[713,748,701,728]" pageId="62" pageNumber="507" type="generic">2 ♂</specimenCount>
,
<quantity id="4CF34E97CA27FFD4ECDBFD534246FD6A" box="[761,835,701,728]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="62" pageNumber="507" unit="km" value="45.0">
<locationDeviation id="84F74691D0C69E8E86623CF761CFF1D5" box="[761,835,701,728]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="62" pageNumber="507" unit="km" value="45.0">45 km</locationDeviation>
</quantity>
<location id="D5F86EF14A1D73ADE283680C33B31827" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:D5F86EF14A1D73ADE283680C33B31827" country="Malawi" latitude="-14.26" longLatPrecision="558" longitude="33.455" name="SW of Lilongwe" stateProvince="Central Region">SW of Lilongwe</location>
,
<collectingRegion id="49CF2D90CA27FFD4EEBDFD0F4077FD4F" box="[159,370,738,765]" pageId="62" pageNumber="507">Lilongwe District</collectingRegion>
,
<location id="B9DAC04C3520E5CC043045EB16343D11" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:B9DAC04C3520E5CC043045EB16343D11" country="Malawi" latitude="-14.26" longLatPrecision="558" longitude="33.455" name="Dzalanyama Forest Reserve, SE of Dzalanyama Forest Lodge" stateProvince="Central Region">Dzalanyama Forest Reserve, SE of Dzalanyama Forest Lodge</location>
, miombo woodland with rocky areas and marshy grassland,
<elevation id="20F1893FED77E552F96333ED83236247" box="[951,1036,776,802]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="62" pageNumber="507" unit="m" value="1250.0">
<quantity id="E5559CE53A5103A31DDA54A7F3431D5F" box="[951,1036,776,802]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="62" pageNumber="507" unit="m" value="1250.0">1250 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA26FFD5EEC0FF10403EFEAA" box="[226,315,253,280]" direction="south" orientation="latitude" pageId="63" pageNumber="508" precision="555" value="-14.26">14.26 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA26FFD5EF61FF1040AAFEAA" box="[323,431,253,280]" direction="east" orientation="longitude" pageId="63" pageNumber="508" precision="55" value="33.455">33.455 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA26FFD5EFE6FF10437DFEAA" box="[452,632,253,280]" pageId="63" pageNumber="508" value="2001-12-27" valueMax="2001-12-29" valueMin="2001-12-27">
<collectingDate id="B85CC1F5F5ADD5ED59D6C844D3305C5E" box="[452,632,253,280]" pageId="63" pageNumber="508" value="2001-12-27" valueMax="2001-12-29" valueMin="2001-12-27">2729 -xii- 2001</collectingDate>
</date>
, leg. K.-
<collectorName id="F02CF1C83A34601A904141479BABDEA1">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EDA0FF1342DBFEAA" box="[898,990,254,280]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EDCBFF1342CDFE3A" ID-GBIF-Occurrence="1224547719" collectingDate="2001-11-06" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="Malawi" elevation="1000" latitude="-15.7922" location="Blantyre" longLatPrecision="7" longitude="34.9943" pageId="63" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Southern Region">
<specimenCount id="9D0D28FBCA26FFD5EDCBFF134509FEA5" box="[1001,1036,253,280]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
(identification suspected),
<collectingCountry id="F31CA3E2CA26FFD5EFFAFECE434AFE8F" box="[472,591,291,317]" name="Malawi" pageId="63" pageNumber="508" value="Malawi">MALAWI</collectingCountry>
(
<collectingRegion id="49CF2D90CA26FFD5EC7DFECF422BFE8F" box="[607,814,290,317]" pageId="63" pageNumber="508">Southern Region</collectingRegion>
):
<collectingRegion id="49CF2D90CA26FFD5ED63FECF4502FE8F" box="[833,1031,290,317]" pageId="63" pageNumber="508">Blantyre District</collectingRegion>
,
<location id="10F6F36B6724F6852D8A91C5417E6134" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:10F6F36B6724F6852D8A91C5417E6134" country="Malawi" latitude="-15.7922" longLatPrecision="7" longitude="34.9943" name="Blantyre" stateProvince="Southern Region">Blantyre</location>
,
<location id="25E664B365ADAC35AAEC777876521E91" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:25E664B365ADAC35AAEC777876521E91" country="Malawi" latitude="-15.7922" longLatPrecision="7" longitude="34.9943" name="Sunnyside" stateProvince="Southern Region">Sunnyside</location>
,
<location id="C039CEEA56363898400D488494F0A557" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:C039CEEA56363898400D488494F0A557" country="Malawi" latitude="-15.7922" longLatPrecision="7" longitude="34.9943" name="Smythe Road" stateProvince="Southern Region">Smythe Road</location>
, rough grass in suburban garden,
<elevation id="D930D4C19C3B4653B211651011D3BB6B" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<quantity id="74BCE732AC16C9F5DE32FB9A4C679E64" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA26FFD5EEC2FE804050FE3A" box="[224,341,365,392]" direction="south" orientation="latitude" pageId="63" pageNumber="508" precision="5" value="-15.7922">15.7922 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA26FFD5EF79FE8040D0FE3A" box="[347,469,365,392]" direction="east" orientation="longitude" pageId="63" pageNumber="508" precision="5" value="34.9943">34.9943 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA26FFD5EFCAFE80436CFE3A" box="[488,617,365,392]" pageId="63" pageNumber="508" value="2001-11-06">
<collectingDate id="577BF2C9D0D569F5409579EA74F08F72" box="[488,617,365,392]" pageId="63" pageNumber="508" value="2001-11-06">06-xi- 2001</collectingDate>
</date>
, leg. K.-
<collectorName id="FD109F8BB0FD5D7715AA014B91AE759A">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5ED4EFE8342CDFE3A" box="[876,968,366,392]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EDF3FE83403DFDAF" ID-GBIF-Occurrence="1224547708" collectingDate="2001-12-05" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="MO- ZAMBIQUE" elevation="1875" latitude="-15.3883" location="NE of Gurue, Namuli Massif, Muretha Plateau" longLatPrecision="7" longitude="37.0467" pageId="63" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Zambezia Province">
<collectingCountry id="C27DD82E5C7F5D8EDDB8AEA5F214ACB6">MO- ZAMBIQUE</collectingCountry>
(
<collectingRegion id="49CF2D90CA26FFD5EF66FE7F4329FE1F" box="[324,556,402,429]" pageId="63" pageNumber="508">Zambezia Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA26FFD5EC1FFE7E435AFE1E" box="[573,607,402,429]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
,
<quantity id="4CF34E97CA26FFD5EC4EFE7E43B1FE1F" box="[620,692,402,429]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="63" pageNumber="508" unit="km" value="15.0">
<locationDeviation id="E4AD66B67AA3C5D76C64B28AAA2CE8B7" box="[620,692,402,429]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="63" pageNumber="508" unit="km" value="15.0">15 km</locationDeviation>
</quantity>
<location id="2A1A6ECE198948D2935703A7089D9F6D" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:2A1A6ECE198948D2935703A7089D9F6D" country="MO- ZAMBIQUE" latitude="-15.3883" longLatPrecision="7" longitude="37.0467" name="NE of Gurue, Namuli Massif, Muretha Plateau" stateProvince="Zambezia Province">NE of Gurue, Namuli Massif, Muretha Plateau</location>
, grassy plateau with streams, boggy pits and forest patches,
<elevation id="247E0E335D3A60B3900C19CFD96F54A6" box="[259,334,478,504]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="1.89" metricValueMin="1.86" pageId="63" pageNumber="508" unit="m" value="1875.0" valueMax="1890.0" valueMin="1860.0">
<quantity id="AAB6BE513927901892272CFDE1CEFE8A" box="[259,334,478,504]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="1.89" metricValueMin="1.86" pageId="63" pageNumber="508" unit="m" value="1875.0" valueMax="1890.0" valueMin="1860.0">1860 1890 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA26FFD5EFB8FE30430AFE4A" box="[410,527,477,504]" direction="south" orientation="latitude" pageId="63" pageNumber="508" precision="5" value="-15.3883">15.3883 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA26FFD5EC3AFE304397FE4A" box="[536,658,477,504]" direction="east" orientation="longitude" pageId="63" pageNumber="508" precision="5" value="37.0467">37.0467 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA26FFD5EC85FE304235FE4A" box="[679,816,477,504]" pageId="63" pageNumber="508" value="2001-12-05">
<collectingDate id="73ED1DA4536AD608B585C820300E6298" box="[679,816,477,504]" pageId="63" pageNumber="508" value="2001-12-05">05-xii- 2001</collectingDate>
</date>
, leg. K.-
<collectorName id="506C5F7480F3793036F82335ED78FBAB">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EEFEFDEE403DFDAF" box="[220,312,515,541]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EF64FDEE4060FD3F" ID-GBIF-Occurrence="1224547702" collectingDate="2014-11-20" collectionCode="RMNH" collectorName="J. Kipping" country="Zambia" elevation="1296" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF421-15" latitude="-12.3676" location="Chimfunshi Wildlife Reserve near Chingola" longLatPrecision="7" longitude="27.4831" pageId="63" pageNumber="508" specimenCode="RMNH.INS.508067" specimenCount="1" stateProvince="Copperbelt Province">
<collectingCountry id="F31CA3E2CA26FFD5EF64FDEE40B1FDAF" box="[326,436,515,541]" name="Zambia" pageId="63" pageNumber="508" value="Zambia">ZAMBIA</collectingCountry>
(
<collectingRegion id="49CF2D90CA26FFD5EFEAFDEF43C0FDAF" box="[456,709,514,541]" pageId="63" pageNumber="508">Copperbelt Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA26FFD5ECFEFDEE43FBFDAE" box="[732,766,514,541]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
(
<specimenCode id="DEFAA8A9FE006BA93872A07577A86288">RMNH.INS.508067</specimenCode>
),
<location id="1705D1AC35EF96308A6EDB6B572F958A" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:1705D1AC35EF96308A6EDB6B572F958A" country="Zambia" latitude="-12.3676" longLatPrecision="7" longitude="27.4831" name="Chimfunshi Wildlife Reserve near Chingola" stateProvince="Copperbelt Province">Chimfunshi Wildlife Reserve near Chingola</location>
, shallow pool in grassy dambo near gallery forest,
<elevation id="3D5E7B939FE228B9B559AE3DB000EA78" box="[410,485,590,616]" metricMagnitude="3" metricUnit="m" metricValue="1.2959999999999998" pageId="63" pageNumber="508" unit="m" value="1296.0">
<quantity id="9C0F940E56F6CC2AFBC915E95764F1A2" box="[410,485,590,616]" metricMagnitude="3" metricUnit="m" metricValue="1.2959999999999998" pageId="63" pageNumber="508" unit="m" value="1296.0">1296 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA26FFD5EC13FDA043A3FDDA" box="[561,678,589,616]" direction="south" orientation="latitude" pageId="63" pageNumber="508" precision="5" value="-12.3676">12.3676 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA26FFD5EC8CFDA0422DFDDA" box="[686,808,589,616]" direction="east" orientation="longitude" pageId="63" pageNumber="508" precision="5" value="27.4831">27.4831 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA26FFD5ED1CFDA042BAFDDA" box="[830,959,589,616]" pageId="63" pageNumber="508" value="2014-11-20">
<collectingDate id="399CF700050941303A158A2ECA74579D" box="[830,959,589,616]" pageId="63" pageNumber="508" value="2014-11-20">20 -xi- 2014</collectingDate>
</date>
, leg.
<collectorName id="4AF2D98C29A116212C643D3A21826728">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EF2BFD9E4060FD3F" box="[265,357,627,653]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EF4CFD9E4236FD3F" ID-GBIF-Occurrence="1224547712" box="[366,819,626,653]" collectingDate="2014-11-20" collectionCode="CJKL" collectorName="J. Kipping" country="Zambia" elevation="1296 m" latitude="-12.3676" location="Chimfunshi Wildlife Reserve near Chingola" longLatPrecision="7" longitude="27.4831" pageId="63" pageNumber="508" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Copperbelt Province">
<specimenCount id="9D0D28FBCA26FFD5EF4CFD9E4095FD3E" box="[366,400,626,653]" pageId="63" pageNumber="508" type="generic">3 ♂</specimenCount>
<specimenCount id="E17C0ECCED9F421E8C5691C2AB2411EE" type="female">3 ♀</specimenCount>
, same locality, same dates,
<collectionCode id="ED1A7BB7CA26FFD5ECCCFD9E4236FD3F" box="[750,819,627,653]" collectionName="CJKL" pageId="63" pageNumber="508">CJKL</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5ED19FD9E4261FD4F" ID-GBIF-Occurrence="1224547703" collectingDate="2014-12-04" collectionCode="RMNH" collectorName="J. Kipping" country="Zambia" elevation="1434" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF422-15" latitude="-11.1703" location="Shiwa Ngandu, Kapishya Hot Springs" longLatPrecision="7" longitude="31.6057" pageId="63" pageNumber="508" specimenCode="RMNH.INS.508066" specimenCount="1" specimenCount-male="1" stateProvince="Northern Province">
<collectingCountry id="F31CA3E2CA26FFD5ED19FD9E42AFFD3F" box="[827,938,627,653]" name="Zambia" pageId="63" pageNumber="508" value="Zambia">ZAMBIA</collectingCountry>
(
<collectingRegion id="49CF2D90CA26FFD5ED9AFD9F4044FD00" pageId="63" pageNumber="508">Northern Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA26FFD5EF79FD754078FD00" box="[347,381,664,690]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
(
<specimenCode id="A0FB71EA3A190D0B039A75514FFCFE74">RMNH.INS.508066</specimenCode>
),
<location id="D20C3CD2505BD034EBCF936FC722E154" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:D20C3CD2505BD034EBCF936FC722E154" country="Zambia" latitude="-11.1703" longLatPrecision="7" longitude="31.6057" name="Shiwa Ngandu, Kapishya Hot Springs" stateProvince="Northern Province">Shiwa Ngandu, Kapishya Hot Springs</location>
, shallow grassy pools near
<location id="C6CFD9EF4FB8D23AC2C591894B2BFBF2" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:C6CFD9EF4FB8D23AC2C591894B2BFBF2" country="Zambia" latitude="-11.1703" longLatPrecision="7" longitude="31.6057" name="Mansha River" stateProvince="Northern Province">Mansha River</location>
(
<figureCitation id="1330FFF7CA26FFD5ECCFFD504259FD6A" box="[749,860,701,728]" captionStart="Photo 22" captionStartId="64.[159,225,1290,1314]" captionText="Photo 22. Shallow temporary pools close to the Mansha River at Shiwa Ngandu, Zambia. Habitat of Ceriagrion banditum, C. junceum and some other Ceriagrion. Photo: JK (05 - - xii- 2014)" pageId="63" pageNumber="508">
<location id="F7FEE57E070237D571B558F5DD65F689" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:F7FEE57E070237D571B558F5DD65F689" country="Zambia" latitude="-11.1703" longLatPrecision="7" longitude="31.6057" name="Photo" stateProvince="Northern Province">Photo</location>
22
</figureCitation>
),
<elevation id="DF816B7C66C8E6650C641BFE0C439289" box="[891,977,702,728]" metricMagnitude="3" metricUnit="m" metricValue="1.434" pageId="63" pageNumber="508" unit="m" value="1434.0">
<quantity id="D63E9D0F005A21FE011E53A7A6CD76FC" box="[891,977,702,728]" metricMagnitude="3" metricUnit="m" metricValue="1.434" pageId="63" pageNumber="508" unit="m" value="1434.0">1434 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA26FFD5EE85FD0F4019FD4F" box="[167,284,738,765]" direction="south" orientation="latitude" pageId="63" pageNumber="508" precision="5" value="-11.1703">11.1703 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA26FFD5EF04FD0F40A5FD4F" box="[294,416,738,765]" direction="east" orientation="longitude" pageId="63" pageNumber="508" precision="5" value="31.6057">31.6057 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA26FFD5EF94FD0F4345FD4F" box="[438,576,738,765]" pageId="63" pageNumber="508" value="2014-12-04">
<collectingDate id="D572AF1929377561ED45030E71FFF249" box="[438,576,738,765]" pageId="63" pageNumber="508" value="2014-12-04">04-xii- 2014</collectingDate>
</date>
, leg.
<collectorName id="748E08D084BA8B8E538AAEE36075730E">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5ED2AFD0E4261FD4F" box="[776,868,739,765]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5ED53FD0E43AAFC90" ID-GBIF-Occurrence="1224547724" collectingDate="2014-12-06" collectionCode="RMNH" collectorName="J. Kipping" country="Zambia" elevation="1434 m" latitude="-11.1703" location="Shiwa Ngandu, Kapishya Hot Springs" longLatPrecision="7" longitude="31.6057" pageId="63" pageNumber="508" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern Province">
<specimenCount id="9D0D28FBCA26FFD5ED53FD0E4296FD4E" box="[881,915,738,765]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
<specimenCount id="4B28A805FE4A4515E828FCE16F295666" type="female">1 ♀</specimenCount>
, same locality,
<date id="FFB5C5B2CA26FFD5EF21FCE54089FC90" box="[259,396,776,802]" pageId="63" pageNumber="508" value="2014-12-06">
<collectingDate id="DD824540B0707F63CED1BA73E083B7D8" box="[259,396,776,802]" pageId="63" pageNumber="508" value="2014-12-06">06-xii- 2014</collectingDate>
</date>
, leg.
<collectorName id="3EA411F79C0027E427E1348013498BE6">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EC71FCE443AAFC90" box="[595,687,777,802]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EC9EFCE540F9FCFA" ID-GBIF-Occurrence="1224547721" collectingDate="2014-12-04" collectionCode="BMNH" collectorName="J. Kipping" country="Zambia" elevation="1434 m" location="Shiwa Ngandu, Kapishya Hot Springs" pageId="63" pageNumber="508" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern Province">
<specimenCount id="9D0D28FBCA26FFD5EC9EFCE543DBFC90" box="[700,734,776,802]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
<specimenCount id="C4A372F1D784EA902840CFF5753FA563" type="female">1 ♀</specimenCount>
same locality,
<date id="FFB5C5B2CA26FFD5ED9FFCE541E1FCFA" pageId="63" pageNumber="508" value="2014-12-04">
<collectingDate id="9D5B5234B25DDD4C2C361862AC03FE1E" pageId="63" pageNumber="508" value="2014-12-04">04-xii- - 2014</collectingDate>
</date>
, leg.
<collectorName id="48A015942097051D6A36F1ADE91DBFE2">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EF83FCC340F9FCFA" box="[417,508,814,840]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="63" pageNumber="508">BMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EC24FCC34023FCDF" ID-GBIF-Occurrence="1224547720" collectingDate="2014-12-04" collectionCode="CJKL" collectorName="J. Kipping" country="Zambia" elevation="1434 m" location="Shiwa Ngandu, Kapishya Hot Springs" pageId="63" pageNumber="508" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern Province">
<specimenCount id="9D0D28FBCA26FFD5EC24FCC3432DFCF5" box="[518,552,813,840]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
<specimenCount id="3CDD86C60471D2352A99D9D89E3DCAC7" type="female">1 ♀</specimenCount>
same locality,
<date id="FFB5C5B2CA26FFD5ECDFFCC04283FCFA" box="[765,902,813,840]" pageId="63" pageNumber="508" value="2014-12-04">
<collectingDate id="3B333A1B2A99EFCAA95367DB06B7B5DC" box="[765,902,813,840]" pageId="63" pageNumber="508" value="2014-12-04">04-xii- 2014</collectingDate>
</date>
, leg.
<collectorName id="17FD40B90E1C04CB0693748F3B2A871D">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EEC0FCBE4023FCDF" box="[226,294,851,877]" collectionName="CJKL" pageId="63" pageNumber="508">CJKL</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EF10FCBE42DBFC0A" ID-GBIF-Occurrence="1224547711" collectingDate="2010-02-25" collectionCode="CJKL" collectorName="J. Kipping" country="Zambia" elevation="1405 m" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF408-15" latitude="-11.2332" location="Sakeji School on Hillwood Farm" longLatPrecision="7" longitude="24.3125" municipality="Ikelenge" pageId="63" pageNumber="508" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Northwestern Province">
<collectingCountry id="F31CA3E2CA26FFD5EF10FCBE40A4FCDF" box="[306,417,851,877]" name="Zambia" pageId="63" pageNumber="508" value="Zambia">ZAMBIA</collectingCountry>
(
<collectingRegion id="49CF2D90CA26FFD5EF93FCBF43CFFCDF" box="[433,714,850,877]" pageId="63" pageNumber="508">Northwestern Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA26FFD5ECFCFCBE4205FCDE" box="[734,768,850,877]" pageId="63" pageNumber="508" type="generic">1 ♂</specimenCount>
(
<specimenCode id="FCD2337AB90E4264EB639AB5C45D9FC8">RMNH.INS.507960</specimenCode>
),
<collectingMunicipality id="68FBE35456DEDD56C72EFB04E25699F8">Ikelenge</collectingMunicipality>
,
<location id="75563C5A4D62CF23CB48715DFC9B14A3" LSID="urn:lsid:plazi:treatment:03A25264CA22FFABEEE4FD11426AFC90:75563C5A4D62CF23CB48715DFC9B14A3" country="Zambia" latitude="-11.2332" longLatPrecision="7" longitude="24.3125" municipality="Ikelenge" name="Sakeji School on Hillwood Farm" stateProvince="Northwestern Province">Sakeji School on Hillwood Farm</location>
, grassy ditches at forest edge 1
<quantity id="4CF34E97CA26FFD5EE97FC734005FC0A" box="[181,256,926,952]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="63" pageNumber="508" unit="m" value="405.0">
<elevation id="20B32853F19D7C12095EA800F0B0AAE2" box="[181,256,926,952]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="63" pageNumber="508" unit="m" value="405.0">405 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA26FFD5EF6FFC7040C7FC0A" box="[333,450,925,952]" direction="south" orientation="latitude" pageId="63" pageNumber="508" precision="5" value="-11.2332">11.2332 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA26FFD5EFE9FC704340FC0A" box="[459,581,925,952]" direction="east" orientation="longitude" pageId="63" pageNumber="508" precision="5" value="24.3125">24.3125 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA26FFD5EC78FC7043D0FC0A" box="[602,725,925,952]" pageId="63" pageNumber="508" value="2010-02-25">
<collectingDate id="A9A47DDA2D2B1B0259E3B5E233C3938E" box="[602,725,925,952]" pageId="63" pageNumber="508" value="2010-02-25">25 -ii- 2010</collectingDate>
</date>
, leg.
<collectorName id="3AC04922A970FC10EF6C3511FC91A636">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5EDBBFC7342DBFC0A" box="[921,990,926,952]" collectionName="CJKL" pageId="63" pageNumber="508">CJKL</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA26FFD5EDCBFC73439AFC6F" ID-GBIF-Occurrence="1224547709" collectingDate="2014-12-08" collectionCode="RMNH" collectorName="J. Kipping" country="Zambia" elevation="1619 m" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF423-15" latitude="-8.8341" location="Lake Chila near Mbala" longLatPrecision="7" longitude="31.3948" municipality="Ikelenge" pageId="63" pageNumber="508" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-female="1" stateProvince="Northwestern Province">
<specimenCount id="6E59FB4255730B19BC00FE31EA79C264" type="female">1 ♀</specimenCount>
(
<specimenCode id="6EA94D175B6C2B070D5FF606DEC9D639">RMNH.INS.508069</specimenCode>
), as holotype,
<collectionCode id="ED1A7BB7CA26FFD5EC61FC2E439AFC6F" box="[579,671,963,989]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="63" pageNumber="508">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="DBD0E6CC59249FC5366AC051A189F764" ID-GBIF-Occurrence="1224547707" box="[683,981,962,989]" collectingDate="2014-12-11" collectionCode="RMNH" collectorName="J. Kipping" country="Zambia" elevation="1619 m" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF424-15" latitude="-8.8341" location="Lake Chila near Mbala" longLatPrecision="7" longitude="31.3948" pageId="63" pageNumber="508" specimenCode="RMNH.INS.554425" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Northwestern Province">
<specimenCount id="9D0D28FBCA26FFD5EC89FC2E43C8FC6E" box="[683,717,962,989]" pageId="63" pageNumber="508" type="generic">2 ♂</specimenCount>
(
<specimenCode id="B772E2C2DB0A46F8053D9A18FD3B0159">RMNH.INS.508070</specimenCode>
),
<specimenCount id="BE0EFCABFD693636D269396808A89131" type="female">1 ♀</specimenCount>
, locality as holotype,
<date id="FFB5C5B2CA26FFD5EFAFFC054313FBB0" box="[397,534,1000,1026]" pageId="63" pageNumber="508" value="2014-12-11">
<collectingDate id="04FAC32937C4D003CFF7063A90EDD38D" box="[397,534,1000,1026]" pageId="63" pageNumber="508" value="2014-12-11">11 -xii- 2014</collectingDate>
</date>
, leg.
<collectorName id="5FCEF9B2D1D0CC39EF4D6DF2A707FC82">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA26FFD5ECF0FC054212FBB0" box="[722,791,1000,1026]" collectionName="CJKL" pageId="63" pageNumber="508">CJKL</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF74B3FACA27FFD4EEBDFA1A4267F9BD" box="[159,866,1527,1552]" pageId="62" pageNumber="507" targetBox="[158,1035,827,1486]" targetPageId="62">
<paragraph id="8BB4E372CA27FFD4EEBDFA1A4267F9BD" box="[159,866,1527,1552]" pageId="62" pageNumber="507">
<emphasis id="B97F3F60CA27FFD4EEBDFA1A4041F9BD" bold="true" box="[159,324,1527,1552]" pageId="62" pageNumber="507">
Type
<figureCitation id="1330FFF7CA27FFD4EEF8FA1A4045F9BD" box="[218,320,1527,1551]" captionStart="Photo 15" captionStartId="43.[782,848,1384,1408]" captionText="Photo 15. Sandy and clear stream on the Batéké Plateau in Gabon. Habitat of Elattoneura aurifex. Photo: A. Günther (27 - i- 2012)" pageId="62" pageNumber="507">Photo 15</figureCitation>
.
</emphasis>
Holotype of
<taxonomicName id="4C0B98F1CA27FFD4EFEDFA1A43ABF9BD" box="[463,686,1527,1551]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="507" phylum="Arthropoda" rank="species" species="banditum" status="sp. nov.">
<emphasis id="B97F3F60CA27FFD4EFEDFA1A43ABF9BD" box="[463,686,1527,1551]" italics="true" pageId="62" pageNumber="507">Ceriagrion banditum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA27FFD4EC96FA1A420FF9BD" box="[692,778,1527,1551]" pageId="62" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
, RMNH.
</paragraph>
</caption>
<subSubSection id="C311B0F9CA26FFD5EEBDFBDE4007FB2A" pageId="63" pageNumber="508" type="description">
<paragraph id="8BB4E372CA26FFD5EEBDFBDE400CFBFF" box="[159,265,1075,1101]" pageId="63" pageNumber="508">
<heading id="D0FC541ECA26FFD5EEBDFBDE400CFBFF" bold="true" box="[159,265,1075,1101]" fontSize="11" level="3" pageId="63" pageNumber="508" reason="6">
<emphasis id="B97F3F60CA26FFD5EEBDFBDE400CFBFF" bold="true" box="[159,265,1075,1101]" pageId="63" pageNumber="508">Genetics</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA26FFD5EEBDFBB54007FB2A" pageId="63" pageNumber="508">
Three unique haplotypes (n= 6) nearest to
<taxonomicName id="4C0B98F1CA26FFD5ECAFFBB543F0FBC0" box="[653,757,1112,1138]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="508" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B97F3F60CA26FFD5ECAFFBB543F0FBC0" box="[653,757,1112,1138]" italics="true" pageId="63" pageNumber="508">C. bakeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA26FFD5ED0FFBB542AAFBC1" box="[813,943,1112,1139]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="508" phylum="Arthropoda" rank="species" species="junceum" status="sp. nov.">
<emphasis id="B97F3F60CA26FFD5ED0FFBB542AAFBC1" box="[813,943,1112,1139]" italics="true" pageId="63" pageNumber="508">C. junceum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA26FFD5ED97FBB44509FBC0" box="[949,1036,1113,1138]" pageId="63" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1330FFF7CA26FFD5EE85FB9041F6FB2A" box="[167,243,1149,1176]" captionStart="Tree 2. 50" captionStartId="60.[159,207,1440,1464]" captionTargetBox="[159,1033,406,1398]" captionTargetId="figure@60.[156,1038,406,1400]" captionTargetPageId="60" captionText="Tree 2. 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis for the suave - complex of Ceriagrion, related species and two outgroup taxa. For relevant branches posterior probabilities from Bayesian inference analysis are shown first, as is bootstrap support (both as percentages)." pageId="63" pageNumber="508">Tree 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA26FFABEEBDFB244250FED0" lastPageId="65" lastPageNumber="510" pageId="63" pageNumber="508" type="diagnosis">
<paragraph id="8BB4E372CA26FFD5EEBDFB244317FB51" box="[159,530,1224,1251]" pageId="63" pageNumber="508">
<heading id="D0FC541ECA26FFD5EEBDFB244317FB51" bold="true" box="[159,530,1224,1251]" fontSize="11" level="3" pageId="63" pageNumber="508" reason="6">
<emphasis id="B97F3F60CA26FFD5EEBDFB244317FB51" bold="true" box="[159,530,1224,1251]" pageId="63" pageNumber="508">Male morphological diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA26FFABEEBDFB004250FED0" lastPageId="65" lastPageNumber="510" pageId="63" pageNumber="508">
Distinct within the
<taxonomicName id="4C0B98F1CA26FFD5EFAEFB0340C9FABA" box="[396,460,1262,1288]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="508" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA26FFD5EFAEFB0340C9FABA" box="[396,460,1262,1288]" italics="true" pageId="63" pageNumber="508">suave</emphasis>
</taxonomicName>
-complex (see above) by its (1) greater size, Hw
<quantity id="4CF34E97CA26FFD5EEBDFAFE4040FA9F" box="[159,325,1299,1325]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.25" metricValueMin="2.15" pageId="63" pageNumber="508" unit="mm" value="22.0" valueMax="22.5" valueMin="21.5">21.522.5 mm</quantity>
(n = 13) rather than 17.021.0 mm; (2) sleeker build with the wing tips reaching at most a third down the length of S 6, rather than halfway or beyond, and Hw 5051 % of full length (n= 8), versus 5157 % in
<taxonomicName id="4C0B98F1CA26FFD5EEBDFA6E41FBFA2F" box="[159,254,1411,1437]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="508" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA26FFD5EEBDFA6E41FBFA2F" box="[159,254,1411,1437]" italics="true" pageId="63" pageNumber="508">C.suave</emphasis>
</taxonomicName>
(n= 9); (3) two narrow horizontal dark bands on the greenish eyes, which are typically lost with maturity in other species; (4) rather intermediate colour of head and thorax, i.e., neither quite uniformly orange like
<taxonomicName id="4C0B98F1CA26FFD5EEBDFA1E4005F9BF" box="[159,256,1523,1549]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="508" phylum="Arthropoda" rank="species" species="suave">
<emphasis id="B97F3F60CA26FFD5EEBDFA1E4005F9BF" box="[159,256,1523,1549]" italics="true" pageId="63" pageNumber="508">C.suave</emphasis>
</taxonomicName>
, nor more two-toned brown and cream with dark dots on the sutures as
<taxonomicName id="4C0B98F1CA59FFAAEEDDFF104300FEAA" authority="Pinhey, 1963" authorityName="Pinhey" authorityYear="1963" box="[255,517,253,280]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="species" species="sakejii">
<emphasis id="B97F3F60CA59FFAAEEDDFF10406DFEAA" box="[255,360,253,280]" italics="true" pageId="64" pageNumber="509">C.sakejii</emphasis>
Pinhey, 1963
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA59FFAAEC62FF1343C1FEAA" box="[576,708,254,280]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="species" species="junceum">
<emphasis id="B97F3F60CA59FFAAEC62FF1343C1FEAA" box="[576,708,254,280]" italics="true" pageId="64" pageNumber="509">C. junceum</emphasis>
</taxonomicName>
; (5) often distinctly reddish rather than pale brown Pt; (6) penis with finger-like lateral lobes, rather like
<taxonomicName id="4C0B98F1CA59FFAAEEBDFEA5400DFED0" box="[159,264,328,354]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B97F3F60CA59FFAAEEBDFEA5400DFED0" box="[159,264,328,354]" italics="true" pageId="64" pageNumber="509">C. bakeri</emphasis>
</taxonomicName>
(
<figureCitation id="1330FFF7CA59FFAAEF3AFEA5406FFED0" box="[280,362,328,354]" captionStart="Figure 10" captionStartId="61.[159,227,1468,1493]" captionTargetBox="[159,1035,733,1427]" captionTargetId="figure@61.[156,1038,730,1430]" captionTargetPageId="61" captionText="Figure 10. Male appendages in dorsal (left) and lateral view (right) of suave - complex of Ceriagrion, including the holotype of C. moorei currently treated as a synonym of C. suave, as well as penis in lateral view (far left) of two related species." pageId="64" pageNumber="509">Fig. 10</figureCitation>
); (7) apical excision on S 10 being about a third as deep as the segment and bordered with tiny black denticles that appear as dark ridges; (8) cerci that in dorsal view appear narrowed and twisted distally, with the apical black tooth turned in- and base-wards, from above thus being well visible and lying almost halfway the cerci; and (9) tips of the paraprocts that typically fall clearly short of those of the cerci (
<figureCitation id="1330FFF7CA58FFABECD6FEA54242FED0" box="[756,839,328,354]" captionStart="Figure 10" captionStartId="61.[159,227,1468,1493]" captionTargetBox="[159,1035,733,1427]" captionTargetId="figure@61.[156,1038,730,1430]" captionTargetPageId="61" captionText="Figure 10. Male appendages in dorsal (left) and lateral view (right) of suave - complex of Ceriagrion, including the holotype of C. moorei currently treated as a synonym of C. suave, as well as penis in lateral view (far left) of two related species." pageId="65" pageNumber="510">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF74B3FACA59FFAAEEBDFBEB4041FB8F" pageId="64" pageNumber="509" targetBox="[158,1035,461,989]" targetPageId="64">
<paragraph id="8BB4E372CA59FFAAEEBDFBEB4041FB8F" pageId="64" pageNumber="509">
<emphasis id="B97F3F60CA59FFAAEEBDFBEB400CFBAD" bold="true" box="[159,265,1030,1055]" pageId="64" pageNumber="509">Photo 21.</emphasis>
<taxonomicName id="4C0B98F1CA59FFAAEF2CFBEB40E9FBAD" box="[270,492,1030,1055]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="species" species="banditum">
<emphasis id="B97F3F60CA59FFAAEF2CFBEB40E9FBAD" box="[270,492,1030,1055]" italics="true" pageId="64" pageNumber="509">Ceriagrion banditum</emphasis>
</taxonomicName>
, holotype male; Lak e Chila, Mbala, Zambia. Photo: JK (08-xii-2014)
</paragraph>
</caption>
<caption id="DF74B3FACA59FFAAEEBDFAE74000F9BC" pageId="64" pageNumber="509">
<paragraph id="8BB4E372CA59FFAAEEBDFAE74000F9BC" pageId="64" pageNumber="509">
<emphasis id="B97F3F60CA59FFAAEEBDFAE7400CFA90" bold="true" box="[159,265,1290,1314]" pageId="64" pageNumber="509">Photo 22.</emphasis>
Shallow temporary pools close to the Mansha River at Shiwa Ngandu, Zambia. Habitat of
<taxonomicName id="4C0B98F1CA59FFAAEF0FFA924002FA07" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="species" species="banditum">
<emphasis id="B97F3F60CA59FFAAEF0FFA924002FA07" italics="true" pageId="64" pageNumber="509">Ceriagrion banditum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B98F1CA59FFAAEF3FFA714098FA07" box="[285,413,1436,1461]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="species" species="junceum">
<emphasis id="B97F3F60CA59FFAAEF3FFA714098FA07" box="[285,413,1436,1461]" italics="true" pageId="64" pageNumber="509">C. junceum</emphasis>
</taxonomicName>
and some other
<taxonomicName id="4C0B98F1CA59FFAAEF49FA5741E0FA5D" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA59FFAAEF49FA5741E0FA5D" italics="true" pageId="64" pageNumber="509">Ceriagrion</emphasis>
</taxonomicName>
. Photo: JK (05- -xii-2014)
</paragraph>
</caption>
<subSubSection id="C311B0F9CA58FFABEEBDFE79426FFE4A" pageId="65" pageNumber="510" type="etymology">
<paragraph id="8BB4E372CA58FFABEEBDFE794021FE1F" box="[159,292,404,429]" pageId="65" pageNumber="510">
<heading id="D0FC541ECA58FFABEEBDFE794021FE1F" bold="true" box="[159,292,404,429]" fontSize="11" level="3" pageId="65" pageNumber="510" reason="6">
<emphasis id="B97F3F60CA58FFABEEBDFE794021FE1F" bold="true" box="[159,292,404,429]" pageId="65" pageNumber="510">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA58FFABEEBDFE55426FFE4A" pageId="65" pageNumber="510">Medieval Latin “banished”, the origin of the word bandit, which refers to the banded eyes recalling the mask of a bandit (neuter adjective).</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA58FFABEEBDFDC4426AFC90" pageId="65" pageNumber="510" type="distribution">
<paragraph id="8BB4E372CA58FFABEEBDFDC44081FDF1" box="[159,388,553,579]" pageId="65" pageNumber="510">
<heading id="D0FC541ECA58FFABEEBDFDC44081FDF1" bold="true" box="[159,388,553,579]" fontSize="11" level="3" pageId="65" pageNumber="510" reason="6">
<emphasis id="B97F3F60CA58FFABEEBDFDC44081FDF1" bold="true" box="[159,388,553,579]" pageId="65" pageNumber="510">Range and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA58FFABEEBDFDA3426AFC90" pageId="65" pageNumber="510">
Known from northern Zambia to northern Mozambique between 1 000 and 1
<quantity id="4CF34E97CA58FFABEE8AFD9E41FEFD3F" box="[168,251,627,653]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="65" pageNumber="510" unit="m" value="900.0">900 m</quantity>
a.s.l. While poorly preserved, males in
<collectionCode id="ED1A7BB7CA58FFABECE3FD9E4210FD3F" box="[705,789,627,653]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" pageId="65" pageNumber="510">MRAC</collectionCode>
from Lubumbashi in southern Congo-Kinshasa also appear to belong to this species (
<figureCitation id="1330FFF7CA58FFABEDA0FD7542D5FD00" box="[898,976,664,690]" captionStart="Map 4" captionStartId="66.[159,210,1496,1520]" captionTargetBox="[159,1035,543,1454]" captionTargetId="figure@66.[159,1038,525,1460]" captionTargetPageId="66" captionText="Map 4. Distribution of the suave - complex of Ceriagrion (C. banditum sp. nov., C. junceum sp. nov., C. mourae, C. sakejii and C. suave) and C. bakeri." pageId="65" pageNumber="510">Map 4</figureCitation>
). Exact habitat difference with other species unclear, but occurs at shallow grassy pools in dambos, open depressions that flood seasonally (see
<taxonomicName id="4C0B98F1CA58FFABED54FD0E42FEFD4F" box="[886,1019,739,765]" class="Insecta" family="Coenagrionidae" genus="Ceriagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="65" pageNumber="510" phylum="Arthropoda" rank="species" species="junceum">
<emphasis id="B97F3F60CA58FFABED54FD0E42FEFD4F" box="[886,1019,739,765]" italics="true" pageId="65" pageNumber="510">C. junceum</emphasis>
</taxonomicName>
). Also found away from water in half-open miombo woodland.
</paragraph>
</subSubSection>
</treatment>
</document>