513 lines
45 KiB
XML
513 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.869.33997" ID-GBIF-Dataset="5b500434-09e3-4370-82a1-1eb789d9d58a" ID-PMC="PMC6690892" ID-Pensoft-Pub="1313-2970-869-19" ID-Pensoft-UUID="C44C602AB0235FB2922DC2A4915C3288" ID-PubMed="31413659" ID-ZooBank="B7C88A64C06E4B67A352F2F9C8FB0D1C" ModsDocID="1313-2970-869-19" checkinTime="1565056339550" checkinUser="pensoft" docAuthor="Ranarilalatiana, Tolotra, Raveloson Ravaomanarivo, Lala Harivelo & Bergsten, Johannes" docDate="2019" docId="0149666EE3E15D2FA4E180FD888742E5" docLanguage="en" docName="ZooKeys 869: 19-90" docOrigin="ZooKeys 869" docSource="http://dx.doi.org/10.3897/zookeys.869.33997" docTitle="Copelatus pulchellus Klug 1834" docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="5E52FF84FFE5FF9193131A454057613C" masterDocTitle="Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species" masterLastPageNumber="90" masterPageNumber="19" pageNumber="19" updateTime="1668167671827" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ranarilalatiana, Tolotra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Raveloson Ravaomanarivo, Lala Harivelo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bergsten, Johannes</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>869</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>90</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.869.33997</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.869.33997</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-869-19</mods:identifier>
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<mods:identifier type="ZooBank">B7C88A64C06E4B67A352F2F9C8FB0D1C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C44C602AB0235FB2922DC2A4915C3288</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="158995562" LSID="urn:lsid:plazi:treatment:0149666EE3E15D2FA4E180FD888742E5" httpUri="http://treatment.plazi.org/id/0149666EE3E15D2FA4E180FD888742E5" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="0149666e-e3e1-5d2f-a4e1-80fd888742e5" authority="Klug, 1834" authorityName="Klug" authorityYear="1834" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">Copelatus pulchellus (Klug, 1834)</taxonomicName>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figs 5C</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">8D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="reference_group">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">Agabus pulchellus</taxonomicName>
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<bibRefCitation author="Klug, JCF" journalOrPublisher="Officina Academica, Berolini" pageId="0" pageNumber="19" refId="B42" refString="Klug, JCF, 1834. Symbolae physicae, seu icones et descriptiones Insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. -Insecta. Decas quarta. . Officina Academica, Berolini" title="Symbolae physicae, seu icones et descriptiones Insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. - Insecta. Decas quarta." year="1834">Klug 1834</bibRefCitation>
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: t. XXXIII: 7
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus africanus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="africanus">Copelatus africanus</taxonomicName>
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Sharp, 1882: 583; TL: Botswana, Lake Ngami;
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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?
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">Copelatus basalis</taxonomicName>
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Boheman, 1848: 244; TL: South Africa (Caffraria interiore);
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus discoideus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="discoideus">Copelatus discoideus</taxonomicName>
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Sharp, 1882: 582; TL: Mesopotamia;
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus obtusus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusus">Copelatus obtusus</taxonomicName>
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Boheman, 1848: 242; TL: South Africa (Caffraria orientali);
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus strigulosus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="strigulosus">Copelatus strigulosus</taxonomicName>
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Sharp, 1882: 582; TL: Mesopotamia;
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mimetes" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mimetes">Copelatus mimetes</taxonomicName>
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Guignot, 1957: 73; TL: Madagascar, Sakaraha, Lambomakandro; syn. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type locality">
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<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="19">Egypt, Sinai.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type material">
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<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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-HT♂ (GP) (
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus mimetes</emphasis>
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) (MNHN, "coll. Guignot"): // Data in NHRS | JLKB | 000030032 // Sakaraha | Lambomakandro | III-56 A.R. // Type [red label] // F. Guignot Det., 1956 |
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="19">mimetes</emphasis>
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n. sp.| Type ♂ // INSTITUT SCIENTIFIQUE MADAGASCAR // -LT♀ (lectotype here designated) (
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
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) (NHRS): // Caffra | ria. // J. Wahlb // Type. // HoloTypus [red label,
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<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
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printed,
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<normalizedToken originalValue="“Holo”">"Holo"</normalizedToken>
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handwritten with small letters in front, possibly by J. Balfour-Browne who studied the type] // obtusus Boh. //
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<emphasis italics="true" pageId="0" pageNumber="19">Cop. pulchellus</emphasis>
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var: |
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<emphasis italics="true" pageId="0" pageNumber="19">obtusus</emphasis>
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Boh. | J. Balfour-Browne det. // 5657 | E91 // NHRS-JLKB | 000065335 // Lectotype |
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
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| Boheman, 1842 | Des. Ranarilalatiana | & Bergsten, 2019// -? ST♂ (
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
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) (NHRS): // Caffra | ria. // J Wahlb // Type. // Typus [red label, printed] // 158 | 61 // Copelatus | basalis Bhn. //
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="19">pulchellus</emphasis>
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| Klug. | Det. 19.iv.1961 | J. Omer-Cooper. // 5597 | E91 // NHRS-JLKB | 000065337 // -? ST♀ (
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
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) (NHRS): // Caffra | ria. // J. Wahlb // Paratypus [red label, printed] // 160 | 61 // 5596 | E91 // NHRS-JLKB | 000065338 // -? ST♀ (
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
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) (NHRS): // Caffra | ria. // J. Wahlb // Paratypus [red label, printed] // 159 | 61 // 5595 | E91 // NHRS-JLKB | 000065339 //
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="materials_examined">
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<paragraph pageId="0" pageNumber="19">Additional material studied.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Antsiranana. Diana: Antsiranana</emphasis>
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: -1♂(GP) (MNHN): // Data in NHRS | JLKB | 000030259 // Madagascar | Diego-Suarez | Ch. Alluaud 1893 //
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<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Boeny: Mitsinjo</emphasis>
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: -1♀ (NHRS): // NHRS-JLKB | 000065733 // Madagascar: Mahajanga: Boeny: | Mahavavy Kinkony RS: S 16.13337 | E 045.95778, 19 m.a.o. 04.XII.2009 | Water Net, Field# MAD09-25 | leg. J. Bergsten, N.
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<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
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, | T. Ranarilalatiana, H.J. Randriamihaja //
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<emphasis bold="true" pageId="0" pageNumber="19">Mahajanga. Melaky: Antsalova</emphasis>
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: -1♂(GP) (NHRS): // NHRS-JLKB | 000010695 // Madagascar: Mahajanga: Melaky: | Tsingy de Bemaraha NP: S19.03572 | E044.77507, 66 m.a.o. 15.XII.2009 | Water Net, Field# MAD09-58 | Leg. J. Bergsten, N.
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<normalizedToken originalValue="Jönsson">Joensson</normalizedToken>
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, T. | Ranarilalatiana, H.J. Randriamihaja //
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Similar to
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<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
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<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
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</taxonomicName>
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(Laporte, 1835) and
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<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
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<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
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</taxonomicName>
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in overall habitus, but body shape more like the elongated shape of
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<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
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<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
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</taxonomicName>
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.
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
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</taxonomicName>
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on Madagascar is ferrugineous black in overall colouration with or without a rather narrow or vague testaceous band basally on elytra (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8D</figureCitation>
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). Instead testaceous regions on elytra are limited to posterolateral and apical parts. The first elytral stria almost full length. Penis in lateral view rather abruptly curved past middle so that basal and apical parts are approximately at right angles (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5C</figureCitation>
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). Penis overall simpler compared to that of
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<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
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<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
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<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
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</taxonomicName>
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and apex slightly twisted.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">(based on Malagasy specimens):</paragraph>
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<paragraph pageId="0" pageNumber="19">Body length 5.5-6.1 mm. Body shape oblong oval, rather convex and attenuate posteriorly, dark brown to blackish ferrugineous. Head infuscated brown ferrugineous, somewhat lighter posteriorly, covered with dense microreticulation and sparser punctation.</paragraph>
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<paragraph pageId="0" pageNumber="19">Pronotum dark brown to ferrugineous black with testaceous anterolateral corners. Disc covered with fine microsculpture forming regular cells and regularily spread small punctures of about same size as cells. Punctuation becomes coarser in posterolateral corners with a weak tendency to corrugate.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Elytra predominantely dark brown to ferrugineous black on disc and along striae with or without a rather narrow and vague basal testaceous band (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus, dorsal view. A Male: Copelatus marginipennis B Female: Copelatus mahajanga C Female: Copelatus sp. 1 (Bemaraha) D Male: Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19">Fig. 8D</figureCitation>
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). Outer intervals and apical region testaceous to varying degrees, often a testaceous interval 4 stands out. Elytral surface covered with same type of microsculpture and punctures as on pronotum. Six clearly impressed elytral striae present on disc and one submarginal stria: second to sixth stria starting at base, first stria somewhat abbreviated anteriorly (by approximately 1/8th the length of elytra); submarginal stria starting at approx. middle.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Ventral side ferrugineous dark brown, with testaceous spots laterally on abdominal ventrites. Metacoxa and ventrites with strioles. Prosternal process more elongate lanceolate and with blunter apex compared with
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<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
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<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
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<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
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</taxonomicName>
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. Lateral parts of metaventrite medium broad. Metacoxal lines short and rather strongly diverging anteriorly. Antennae, palps, pro- and mesolegs testaceous, metalegs somewhat darker testaceous.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Female: elytral striolation limited to the medial parts of the outer three elytral intervals in the single female studied from Madagascar. From other parts of the distribution a female form is known that has the entire elytra striolated (
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<bibRefCitation pageId="0" pageNumber="19" refId="B68">Sharp 1882</bibRefCitation>
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;
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<bibRefCitation pageId="0" pageNumber="19" refId="B37">
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
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and Reiter 2014
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</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Male: protibia bisinuate and angled at base, distally expanded. Penis thin, strongly angled at middle in lateral view, and apex somewhat twisted to the left in ventral view (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Fig. 5C</figureCitation>
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). Parameres more narrowly triangular apically compared to those in
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<taxonomicName lsidName="C. marginipennis" pageId="0" pageNumber="19" rank="species" species="marginipennis">
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<emphasis italics="true" pageId="0" pageNumber="19">C. marginipennis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="C. mahajanga" pageId="0" pageNumber="19" rank="species" species="mahajanga">
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<emphasis italics="true" pageId="0" pageNumber="19">C. mahajanga</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.869.33997.figure8" httpUri="https://binary.pensoft.net/fig/322933" pageId="0" pageNumber="19" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="19">
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Figure 8. Habitus, dorsal view.
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<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
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Male:
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginipennis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="marginipennis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus marginipennis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
|
||
Female:
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus mahajanga" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="mahajanga">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus mahajanga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
|
||
Female:
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||
</taxonomicName>
|
||
sp. 1 (Bemaraha)
|
||
<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
|
||
Male:
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
As the species
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
is currently interpreted, this is a very widely distributed Afrotropical and Middle Eastern species.
|
||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||
recorded the species from Senegal in the west, Tanzania in the east, South Africa in the south, and Mesopotamia (Iraq and adjacent regions) in the Middle East.
|
||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||
and Reiter (2014)
|
||
</bibRefCitation>
|
||
recorded the species from Yemen and stated that the likely distribution in the Middle East included the entire Arabian Peninsula.
|
||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot (1961)</bibRefCitation>
|
||
also gives India and Ceylon (= Sri Lanka) but this was likely based on the misidentification of closely related oriental species (see
|
||
<bibRefCitation author="Ghosh, SK" journalOrPublisher="Skoervnoepparn Supplement" pageId="0" pageNumber="19" pagination="1 - 77" publicationUrl="http://www.norrent.se/images/stories/upload/SN_Supplement_03_hq.pdf" refId="B20" refString="Ghosh, SK, Nilsson, AN, 2012. Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae). . Skoervnoepparn Supplement 3: 1 - 77" title="Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae)." url="http://www.norrent.se/images/stories/upload/SN_Supplement_03_hq.pdf" volume="3" year="2012">Ghosh and Nilsson 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4459.2.2" author="Sheth, SD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="19" pagination="235 - 260" refId="B70" refString="Sheth, SD, Ghate, HV, Hajek, J, 2018. Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). . Zootaxa 4459: 235 - 260" title="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)." url="https://doi.org/10.11646/zootaxa.4459.2.2" volume="4459" year="2018">Sheth et al. 2018</bibRefCitation>
|
||
).
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
was not previously recorded from Madagascar following the revision by
|
||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||
but as we synonymise
|
||
<taxonomicName lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
, Madagascar now forms part of the distribution. From Madagascar we have seen specimens from Antsiranana, Mahavavy Kinkony Reserve, Tsingy de Bemaraha National Park (Bekopaka), and from Lambomakandro, Sakaraha (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11C</figureCitation>
|
||
). It can likely show up anywhere in lowland Madagascar, but especially in the western lowlands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="19" type="habitat">
|
||
<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
On Madagascar we have collected the species associated with a small forest stream with sidepools in a karstic limestone area (
|
||
<normalizedToken originalValue="“tsingy”">"tsingy"</normalizedToken>
|
||
) and in a muddy stagnant pool in a dried-out river bed. Both localities are in dry deciduous forests of lowland western Madagascar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="19" type="comments">
|
||
<paragraph pageId="0" pageNumber="19">Comment.</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
forms part of a diverse species group with many externally very similar species.
|
||
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="0" pageNumber="19" pagination="359 - 399" refId="B4" refString="Balfour-Browne, J, 1950. On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia). . Occasional Papers of the National Museum of Southern Rhodesia 2: 359 - 399" title="On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyrinidae and Palpicornia)." volume="2" year="1950">Balfour-Browne (1950)</bibRefCitation>
|
||
admitted that his previous treatment of the species (
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" author="Balfour-Browne, J" journalOrPublisher="The Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="19" pagination="57 - 88" refId="B3" refString="Balfour-Browne, J, 1939. On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with description of new species. . The Transactions of the Royal Entomological Society of London 88: 57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with description of new species." url="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
|
||
) was entirely wrong as he had then not studied the male genitalia. When he did so in 1950 it resulted in the description of several new species previously lumped under
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
is now interpreted as a widespread Afrotropical and Middle Eastern species with the male penis similar to that illustrated in
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus marginipennis B Copelatus mahajanga C Copelatus pulchellus." figureDoi="10.3897/zookeys.869.33997.figure5" httpUri="https://binary.pensoft.net/fig/322930" pageId="0" pageNumber="19">Figure 5C</figureCitation>
|
||
. Dorsal colouration is interpreted as very variable, even consisting of several distinct colour forms such as the darker forms described as
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
Boheman and later as
|
||
<taxonomicName lsidName="C. africanus" pageId="0" pageNumber="19" rank="species" species="africanus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. africanus</emphasis>
|
||
</taxonomicName>
|
||
Sharp (compare figure 24 with photographs in
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.595.8614" author="Perissinotto, R" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="85 - 135" refId="B60" refString="Perissinotto, R, Bird, MS, Bilton, DT, 2016. Predaceous water beetles (Coleoptera, Hydradephaga) of the Lake St Lucia system, South Africa: biodiversity, community ecology and conservation implications. . ZooKeys 595: 85 - 135" title="Predaceous water beetles (Coleoptera, Hydradephaga) of the Lake St Lucia system, South Africa: biodiversity, community ecology and conservation implications." url="https://doi.org/10.3897/zookeys.595.8614" volume="595" year="2016">Perissinotto et al. (2016</bibRefCitation>
|
||
: fig. 24) and
|
||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||
and Reiter (2014
|
||
</bibRefCitation>
|
||
: fig. 30)). Females are also interpreted as variable in the striolation pattern on the elytra. Sharp described a female form from Mesopotamia that was entirely striolated over the elytra under the name
|
||
<taxonomicName lsidName="C. strigulosus" pageId="0" pageNumber="19" rank="species" species="strigulosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. strigulosus</emphasis>
|
||
</taxonomicName>
|
||
, which is interpreted today as intraspecific variation of
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
(see photograph in
|
||
<bibRefCitation pageId="0" pageNumber="19" refId="B37">
|
||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||
and Reiter (2014
|
||
</bibRefCitation>
|
||
: fig. 30)). We have seen three males and one female from Madagascar. All males are of the darker colour form, lacking a basal transverse testaceous band on elytra, similar to
|
||
<normalizedToken originalValue="Boheman’s">Boheman's</normalizedToken>
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
. The female has a weak basal testaceous band but is otherwise also most similar to the dark colour form. Striolation is limited to the middle of the outer three intervals. We have no doubt that the three non-type specimens examined are conspecific with
|
||
<taxonomicName lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||
</taxonomicName>
|
||
Guignot. But after our comparison of the male genitalia of the type with
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
specimens from mainland Africa, we concluded that
|
||
<taxonomicName lsidName="C. mimetes" pageId="0" pageNumber="19" rank="species" species="mimetes">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. mimetes</emphasis>
|
||
</taxonomicName>
|
||
cannot be upheld as a separate species given how
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
is interpreted. We noted very minor differences in the penis apex which are not greater than differences between east, south, and west African specimens of
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
. The penis figured by
|
||
<bibRefCitation author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="0" pageNumber="19" pagination="71 - 74" refId="B33" refString="Guignot, F, 1957. Un haliplide et deux dytiscides nouveaux de Madagascar. . Le Naturaliste Malgache 9: 71 - 74" title="Un haliplide et deux dytiscides nouveaux de Madagascar." volume="9" year="1957">Guignot (1957</bibRefCitation>
|
||
: fig. 2, reproduced by
|
||
<bibRefCitation DOI="https://doi.org/10.1080/03946975.1997.10539325" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="19" pagination="11 - 48" refId="B56" refString="Nilsson, AN, Bilardo, A, Rocchi, S, 1997. A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings. . Tropical Zoology 10: 11 - 48" title="A check list of Copelatus Erichson 1832 species (ColeopteraDytiscidae) from Afrotropical and Madagascan regions with a review of published penis drawings." url="https://doi.org/10.1080/03946975.1997.10539325" volume="10" year="1997">Nilsson et al. 1997</bibRefCitation>
|
||
: fig. 8) is very inaccurate, depicting a shape quite different from
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
but that does not correspond with the holotype.
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
may still be a species complex for future studies to solve but currently it is a widespread and variable species distributed over Africa, Madagascar, and the Middle East.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||
</taxonomicName>
|
||
Boheman, 1848 was synonymised with
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
by Omer-Cooper (1965). We have studied the same type material as Omer-Cooper and agree with this conclusion. However, we are not convinced that the material housed at NHRS as these types are the correctly identified types of the name
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.9375" author="Boheman, CH" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="19" refId="B11" refString="Boheman, CH, 1848. . https://doi.org/10.5962/bhl.title.9375" url="https://doi.org/10.5962/bhl.title.9375" year="1848">Boheman (1848)</bibRefCitation>
|
||
describes in Latin four species from Johan August Wahlbergs collectings in Caffraria (South Africa) of which three are new, numbered 259 -
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
Klug; 260 -
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus obtusus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus obtusus</emphasis>
|
||
</taxonomicName>
|
||
; 261 -
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus striatellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="striatellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus striatellus</emphasis>
|
||
</taxonomicName>
|
||
; and 262 -
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus basalis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus basalis</emphasis>
|
||
</taxonomicName>
|
||
. For
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
the disc of elytra is described as having six striae. For the longer description of
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
this is further elaborated to detail that there are six discal stria but entire elytra has seven striae
|
||
<normalizedToken originalValue="“septem-striata”">"septem-striata"</normalizedToken>
|
||
(six discal and one submarginal). This pattern and number of striae is consistent with the
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus pulchellus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus pulchellus</emphasis>
|
||
</taxonomicName>
|
||
species group. The following species is correctly identified as
|
||
<taxonomicName lsidName="C. striatellus" pageId="0" pageNumber="19" rank="species" species="striatellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. striatellus</emphasis>
|
||
</taxonomicName>
|
||
with nine discal striae, and is clearly stated as such: "disco striis 9 tenuibus", of which the innermost is much abbreviated. This description matches very well with the types preserved at NHRS. Finally,
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
is described as the last
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
|
||
</taxonomicName>
|
||
species in
|
||
<normalizedToken originalValue="Boheman’s">Boheman's</normalizedToken>
|
||
work and elytra is described as
|
||
<normalizedToken originalValue="“12-striata”">"12-striata"</normalizedToken>
|
||
. The supposed types at NHRS for both
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
have six discal and one submarginal striae. The type for
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
matches the original description of
|
||
<normalizedToken originalValue="“septem-striata”">"septem-striata"</normalizedToken>
|
||
but the three types of
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
with the same number of striae does not match the original description of
|
||
<normalizedToken originalValue="“12-striata”">"12-striata"</normalizedToken>
|
||
. It would be very inconsistent of Boheman to describe the total number of striae on one elytron in one case
|
||
<normalizedToken originalValue="“elytra…septem-striata”">"elytra...septem-striata"</normalizedToken>
|
||
, and in the other only the discal striae but summing up the number from both elytral halves
|
||
<normalizedToken originalValue="“elytra…12-striata”">"elytra...12-striata"</normalizedToken>
|
||
. It would also be very illogical to place
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
after the 9-striated
|
||
<taxonomicName lsidName="C. striatellus" pageId="0" pageNumber="19" rank="species" species="striatellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. striatellus</emphasis>
|
||
</taxonomicName>
|
||
if it has the same number of striae as
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
, which come first. Similarily,
|
||
<bibRefCitation pageId="0" pageNumber="19" refId="B36">Guignot (1961)</bibRefCitation>
|
||
was confused about
|
||
<normalizedToken originalValue="Boheman’s">Boheman's</normalizedToken>
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
and listed the name both under the 12-striated
|
||
<taxonomicName lsidName="C. mocquerysi" pageId="0" pageNumber="19" rank="species" species="mocquerysi">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. mocquerysi</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||
with a question mark, and under
|
||
<taxonomicName lsidName="C. pulchellus" pageId="0" pageNumber="19" rank="species" species="pulchellus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
. We consider the status of
|
||
<taxonomicName lsidName="C. basalis" pageId="0" pageNumber="19" rank="species" species="basalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
Boheman as uncertain but we have not found any alternative potential type material at NHRS. We designate the single undoubted syntype of
|
||
<taxonomicName lsidName="C. obtusus" pageId="0" pageNumber="19" rank="species" species="obtusus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
Boheman, 1848 in the NHRS collection as lectotype to preserve the stability of the name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |