297 lines
28 KiB
XML
297 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.699.12031" ID-GBIF-Dataset="dbf6bb9f-7ea4-4fa3-83ee-f58263427e39" ID-PMC="PMC5673864" ID-Pensoft-Pub="1313-2970-699-1" ID-PubMed="29200927" ID-ZBK="0F92401F3F7C4896AD9D72BC84348C7D" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-699-1" ModsDocOrigin="ZooKeys 699" ModsDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" checkinTime="1505836577638" checkinUser="pensoft" docAuthor="Cherman, Mariana Alejandra, Mise, Kleber Makoto, Moron †, Miguel Angel, Vaz-de-Mello, Fernando Z. & Almeida, Lucia Massutti de" docDate="2017" docId="89AE40C045EE85501CE2A8B90AC051C0" docLanguage="en" docName="ZooKeys 699: 1-120" docOrigin="ZooKeys 699" docSource="http://dx.doi.org/10.3897/zookeys.699.12031" docTitle="Liogenys suturalis Blanchard 1851" docType="treatment" docVersion="4" lastPageNumber="64" masterDocId="FFD9636FB945C41DFF8AB133FFFA9D76" masterDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="61" updateTime="1668164813467" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cherman, Mariana Alejandra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mise, Kleber Makoto</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moron †, Miguel Angel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vaz-de-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Almeida, Lucia Massutti de</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>699</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>120</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.699.12031</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.699.12031</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-699-1</mods:identifier>
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<mods:identifier type="ZBK">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
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<mods:identifier type="ZooBank">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134311177" LSID="urn:lsid:plazi:treatment:89AE40C045EE85501CE2A8B90AC051C0" httpUri="http://treatment.plazi.org/id/89AE40C045EE85501CE2A8B90AC051C0" lastPageId="63" lastPageNumber="64" pageId="60" pageNumber="61">
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<subSubSection pageId="60" pageNumber="61" type="nomenclature">
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys suturalis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="suturalis">Liogenys suturalis (Blanchard, 1851)</taxonomicName>
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Figs 74, 92
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="reference_group">
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Hilarianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hilarianus suturalis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="suturalis">Hilarianus suturalis</taxonomicName>
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Blanchard, 1851: 169 (orig. desc.);
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<bibRefCitation author="Lacordaire, JT" journalOrPublisher="Librarie Encyclopedique de Roret, Paris" pageId="106" pageNumber="107" title="Histoire naturelle des Insectes. Genera des Coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet odre d'insectes. Tome Troiseme. Contenant les familles des Pectinicornes et Lamellicornes." year="1856">Lacordaire 1856</bibRefCitation>
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: 270 (sys.);
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<bibRefCitation pageId="60" pageNumber="61">Harold 1869a</bibRefCitation>
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: 1141 (check.);
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<bibRefCitation pageId="60" pageNumber="61">Dalla Torre 1913</bibRefCitation>
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: 319 (check.);
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<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 228 (check.).
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</paragraph>
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys suturalis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="suturalis">Liogenys suturalis</taxonomicName>
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:
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<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
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: 40, 55 (key, n. comb.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" pagination="1 - 458" title="A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)." url="https://doi.org/10.11646/zootaxa.211.1.1" volume="211" year="2003">Evans 2003</bibRefCitation>
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: 214 (check.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://digitalcommons.unl.edu/entomologypapers/2" year="2005">Evans and Smith 2005</bibRefCitation>
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: 178 (check.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://museum.unl.edu/research/entomology/SSSA/nwmelos.htm" year="2009">Evans and Smith 2009</bibRefCitation>
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: 182 (check.)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="type material">
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<paragraph pageId="60" pageNumber="61">Type material.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Hilarianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hilarianus suturalis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="suturalis">Hilarianus suturalis</taxonomicName>
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female syntype (MNHN): [light green printed]
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<normalizedToken originalValue="“MUSÉUM">"MUSEUM</normalizedToken>
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<normalizedToken originalValue="PÁRIS">PARIS</normalizedToken>
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/ Bresil/deCastelnau", [red printed]
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<normalizedToken originalValue="“SYNTYPE”">"SYNTYPE"</normalizedToken>
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, [green handwritten] "
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<taxonomicName lsidName="H. suturalis" pageId="60" pageNumber="61" rank="species" species="suturalis">H. suturalis</taxonomicName>
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/ Cat Mus/ Minas-Geraes (
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<normalizedToken originalValue="Brésil">Bresil</normalizedToken>
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)/ M. de Castelnau". This type is here designated the lectotype [white, outlined in red, printed] "LECTOTYPE/
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Hilarianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hilarianus suturalis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="suturalis">Hilarianus suturalis</taxonomicName>
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/Blanchard, 1851/des. M. A. Cherman 2014". Female syntype (MNHN): [white handwritten]
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<normalizedToken originalValue="“1859/34”">"1859/34"</normalizedToken>
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, [white handwritten]
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<normalizedToken originalValue="“2860”">"2860"</normalizedToken>
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, [light green printed]
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<normalizedToken originalValue="“MUSÉUM">"MUSEUM</normalizedToken>
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<normalizedToken originalValue="PÁRIS">PARIS</normalizedToken>
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/ [handwritten] Santa-Cruz/
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<normalizedToken originalValue="d’Orbigny”">d'Orbigny"</normalizedToken>
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, [red printed]
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<normalizedToken originalValue="“SYNTYPE”">"SYNTYPE"</normalizedToken>
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, [green handwritten] "
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<taxonomicName lsidName="H. suturalis" pageId="60" pageNumber="61" rank="species" species="suturalis">H. suturalis</taxonomicName>
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/ Cat Mus/ Santa-Cruz (Bolivie)/ M.
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<normalizedToken originalValue="D’Orbigny”">D'Orbigny"</normalizedToken>
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. This type is here designated the paralectotype [white, outlined in red, printed] "PARALECTOTYPE /
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Hilarianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hilarianus suturalis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="suturalis">Hilarianus suturalis</taxonomicName>
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/Blanchard, 1851/des. M. A. Cherman 2014".
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61" type="type material">
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<paragraph pageId="60" pageNumber="61">Non-type material.</paragraph>
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<paragraph lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61">
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BRAZIL. GO:
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<normalizedToken originalValue="Edéia">Edeia</normalizedToken>
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, 14/X/2003, R. B. da Costa col., 1 ex. (CEMT); Goiania, X/2012, Tissiane Col., 2 ex. (CEMT); DF:
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<normalizedToken originalValue="Brasília">Brasilia</normalizedToken>
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, "Lago Norte",
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.761842">15°45'42.63"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-47.83494">47°50'5.8"W</geoCoordinate>
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, XII/2009, 1023 m, Nunes col., 6 ex. (CEMT); Planaltina,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.6044445">15°36'16"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-47.737778">47°44'16"W</geoCoordinate>
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, 3/XI/2006, C. Oliveira col., 5 ex. (CEMT); MT:
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<normalizedToken originalValue="Rosário">Rosario</normalizedToken>
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Oeste, XI/1976, A. Maller col., 2 ex. (MNRJ);
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<normalizedToken originalValue="Córrego">Corrego</normalizedToken>
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Brigadeiro [
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<normalizedToken originalValue="Figueirópolis">Figueiropolis</normalizedToken>
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<normalizedToken originalValue="D’Oeste">D'Oeste</normalizedToken>
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],
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-15.5">15°30'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-58.716667">58°43'W</geoCoordinate>
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, 29/IX/1984, Binda col., 7 ex. (INPA); 29/IX/1984, Marcolino col., 1 ex. (CEMT); BR 270 km 140 [near
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<normalizedToken originalValue="Rondonópolis">Rondonopolis</normalizedToken>
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], 27/IX/1984, Binda col., 1 ex. (INPA); Chapada dos
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<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
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, 23/X/2011, Tassinari col., 1 ex. (CEMT);
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<normalizedToken originalValue="Cuiabá">Cuiaba</normalizedToken>
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, without date and collector, 1 ex.; 24/VIII/1993, Pinto Col., 1 ex.; 20/VI/2010, Rabello col., 1 ex.; 13/XI/2012, Jacobina col., 1 ex.; 3/X/1988, Silva col.,
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<pageBreakToken pageId="61" pageNumber="62" start="start">1</pageBreakToken>
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ex.; 9/IX/1990, Netto col., 1 ex.; 21/V/1986, Costa col., 1 ex. (CEMT);
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<normalizedToken originalValue="Poconé">Pocone</normalizedToken>
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, 15/IV/2009, Abreu col., 1 ex.; 9/XI/2002, without collector, 1 ex.; 21/XI/2003, without collector, 1 ex. (CEMT); Varzea Grande, 13/XI/1993, Paula col., 1 ex. (CEMT); Campo Verde, "Rio Engano", 14/X/2012, Pena and Pena cols., 1 ex. (CEMT); Barra do Bugre, 7/XI/1993, Serrano col., 1 ex. (CEMT); MS: Campo Grande, X/1947, A. Maller col., 2 ex. (AMNH); Porto Murtinho, XI/1929, W. Mehr col., 14 ex. (MNRJ); Dourados,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.228962">22°13'44.26"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-54.830048">54°49'48,18"W</geoCoordinate>
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, 15/VIII/2010, 429 m, Simioni col. 1 ex.; "Faz. Coqueiro
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<normalizedToken originalValue="Azulão”">Azulao"</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-22.241112">22°14'28"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-54.91889">54°55'08"W</geoCoordinate>
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, 12/IV/2012, D. G. Ribeiro col., 1 ex.;
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<normalizedToken originalValue="“UFGD”">"UFGD"</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.194082">22°11'38,7"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-54.921112">54°55'16"W</geoCoordinate>
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, 9/V/2012, D. G. Ribeiro col., 1 ex. (MuBio); 26/XI/2012, F. Vaz-de-Mello col., 3 ex. (CEMT); Nova Andradina, "Usina Santa Helena",
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-21.992222">21°59'32"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-53.441113">53°26'28"W</geoCoordinate>
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, 18/X/2012, 360 m, G. Coutinho col., 4 ex. (CEMT); Rio Brilhante, 21-28/X/1970, V. O. Becker col., 3 ex. (DZUP); Bonito,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-21.12889">21°07'44"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-56.479446">56°28'46"W</geoCoordinate>
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, 19/XII/2010, F. Vaz-de-Mello col., 1 ex. (CEMT); Rio Verde [Rio Verde do Mato Grosso], XI/1959, 400 m, without collector, 2 ex.; X/1966, A. Maller col., 1 ex. (DZUP); BA: Encruzilhada,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.528417">15°31'42,3"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-40.964474">40°57'52,1"W</geoCoordinate>
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, 16/XII/2012, 780 m, Rafael and Grossi col., 1 ex.;
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.576388">15°34'35"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-40.947502">40°56'51"W</geoCoordinate>
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, 16/XII/2012, 850 m, Rafael and Grossi col., 1 ex. (INPA); XI/1975, without collector, 4 ex. (MNRJ); "Estr. SE", 16/XII/2012, P. Grossi col., 4 ex. (EPGC); MG: Aguas vermelhas, XII/1997, Vaz-de-Mello and Bello cols., 2 ex. (CEMT); 12/XII/2012, P Grossi col., 14 ex. (EPGC); (1)
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Roque de Minas, "Serra da Canastra, Sede
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<normalizedToken originalValue="Jaguaré”">Jaguare"</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-20.254723">20°15'17"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-46.419724">46°25'11"W</geoCoordinate>
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, 2/XI/2007, Souza col., 1 ex. (CEMT); Lavras, 9/X/2001, Vieira col., 1 ex. (CEMT); Uberaba, X/1961, D. Elias col., 15 ex. (DZUP);
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<normalizedToken originalValue="Uberlândia">Uberlandia</normalizedToken>
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, 21/X/1974, H.N.
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<normalizedToken originalValue="Espínola">Espinola</normalizedToken>
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col., 2 ex. (DZUP);
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<normalizedToken originalValue="Ibiá">Ibia</normalizedToken>
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, 20/X/1965, C.T. and C. Elias col., 1 ex. (DZUP); SP: Pontal, 23/X/1984, Osman Dermeka col., 4 ex. (DZUP); Descalvado, "Faz. Itaunas", 27/IX/2007, N. W. Perioto col., 2 ex.; 6/X/2006, N. W. Perioto col., 2 ex.; 17/XII/2007, N. W. Perioto col., 1 ex.; 5/XI/2007, N. W. Perioto col., 1 ex. (CEMT); Neves Paulista, 9/VII/2009, Fanti col., 1 ex. (CEMT); Palmital, "Faz.
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<normalizedToken originalValue="São">Sao</normalizedToken>
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<normalizedToken originalValue="Sebastião”">Sebastiao"</normalizedToken>
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, 12/X2010, Callil col., 18 ex. (CEMT); Batatais, 31/X/1969, L. C. Silva col., 1 ex.(DZUP);
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<normalizedToken originalValue="Ribeirão">Ribeirao</normalizedToken>
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Preto, 31/X/1973, Pe. Moure col., 1 ex. (DZUP); PR: Porecatu, 20/X/1970, Becker-Hastchback col., 1 ex. (DZUP). BOLIVIA. SC: Comarapa, "Aserradero La forestal", 9/XI/1987, D. Salvatierra col., 1 ex. (AMNH); Parapeti, I/1960, A. Martinez col., 1 ex. (CMNC); Camiri, 12/XI/1946, Maldonado col., 1 ex. (1) (ZMNC); CB: Cochabamba, without date, Zischka col., 1 ex. (CMNC). PARAGUAY. Without date, Zikan col., 1 ex. (CEIOC); without locality, date and collector, 10 ex.; "camino vogt", without date, Heys col., 1 ex. (SDEI); CN: San Carlos del Apa,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-22.234446">22°14'04"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-57.296665">57°17'48"W</geoCoordinate>
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, 30/X/2002, B. Garcete col., 2 ex. (CEMT); AS:
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<normalizedToken originalValue="Asunción">Asuncion</normalizedToken>
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, XI/1944, Mis. Cient. Brasil col., 3 ex. (CEIOC); IT: Coronel Bogado,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-27.17">27°10'12"S</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-56.25">56°15'0"W</geoCoordinate>
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, I/1944, A. Martinez col., 1 ex. (CEIOC); GU: Villarica, XII/1928, Schade col., 3 ex. (CEIOC). ARGENTINA. SA: El Naranjo, II/1944, A. Martinez col., 2 ex.; Parque Nacional El Rey, 15/XII/1987, 890 mm Peck col., 2 ex.; Tartagal, XI/1944, Duret and Martinez cols., 4 ex. (CMNC); Cafayate,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-26.064444">26°03'52"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-65.938614">65°56'19"W</geoCoordinate>
|
||
, 24/XI/2006, Ocampo, Ruiz, San Blas, Salazar cols., 3 ex. (IADIZA); JU: XI/1947, without collector, 2 ex. (DZUP); without date, Bruch col., 2 ex. (ZMHB); Marta [?], XI/1946, Duret col., 1 ex. (CMNC);
|
||
<pageBreakToken pageId="62" pageNumber="63" start="start">FO</pageBreakToken>
|
||
: Laguna Yema,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-24.331667">24°19'54"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-61.29381">61°17'37,73"W</geoCoordinate>
|
||
, 11/XII/2008, 161 m, Ocampo, San Blas, Campon cols., 2 ex. (IADIZA); CH: XI/1945, A. Martinez col., 1 ex. (CMNC); CO: El Diquecito, I/1976, A. Martinez col., 2 ex.; San Javier, II/1946, Monros col., 1 ex.; Calamuchita, "La Granadilla", I/1974, A. Martinez col., 1 ex. (CMNC); Lozada, XI/2006, P. Fichetti col., 3 ex. (IADIZA); "Cordova, Pampas", without date and collector, 1 ex. (MNHN); SL:
|
||
<normalizedToken originalValue="Luján">Lujan</normalizedToken>
|
||
, I/2010, M. A. Cherman col., 3 ex. (DZUP).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="62" pageNumber="63" type="diagnosis">
|
||
<paragraph pageId="62" pageNumber="63">Diagnosis.</paragraph>
|
||
<paragraph pageId="62" pageNumber="63">Body brownish, elongate; elytra dull yellow, pruinose, opaque; pronotum reddish brown, shiny; clypeal emargination shallow, rounded and wide; outer sides of anterior teeth follow the lateral margin of clypeus, sub-parallel in females; lateral margin straight, slightly convex in females; pronotal posterior corners sharp, obtuse-angled; meso- and metepisternum, sides of metasternum and metacoxae scaly; mesotibia sub-quadrate in cross section; pygidium flat, pygidial disc bristled throughout, coarsely punctured; in males total length of parameres near five times the length of their apex; inner margins straight, slightly divergent; apex harpoon-like with lateral angle curved projecting almost perpendicular to parameres (Fig. 74F).</paragraph>
|
||
<caption pageId="62" pageNumber="63">
|
||
<paragraph pageId="62" pageNumber="63">
|
||
Figure 74.
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys suturalis" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="suturalis">Liogenys suturalis</taxonomicName>
|
||
(Blanchard). A Dorsal view B Lateral view C Frontal view D Clypeus and pronotum E Pygidium F Parameres, dorsal view G Parameres, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="redescription">
|
||
<paragraph pageId="62" pageNumber="63">Redescription.</paragraph>
|
||
<paragraph lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
Length: 13.5-15.0 mm; width: 6.7-7.6 mm. Reddish brown and elytra dull yellow. Head: distance between eyes nearly twice the width of one eye, more than twice in females; frons equal in length to clypeus; clypeal emargination rounded, shallow and wide; outer sides of anterior teeth follow the lateral margin of clypeus in males, sub-parallel in females; clypeal lateral margin straight, slightly convex in females; outer margin of anterior tooth shorter than one eye; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width twice width of apex; fovea deep, extending to the transverse midline of the palpomere; labium transversely carinated, as wide as it is long; antenna 10-articulated, lamellae lighter in color and longer than flagellum. Thorax: anterior margin of pronotum straight; maximum length of pronotum exceeding the length of tarsomeres I, II and III together; disc glabrous, punctures fine and dense, coarser in females; pronotal posterior corners sharp, obtuse-angled; proepisternum with long bristles; meso- and metepisternum, and sides of metasternum scaly, metasternum with few long bristles on the anterior margin; distance between meso- and metacoxae up to twice longer than the metacoxa; scutellum ogival, coarsely punctured. Elytra: glabrous, dull yellow, darker at the base, pruinose; elytra more than three times longer than the pronotum; elytral suture slightly darker than elytron and distinctly elevated; pair of inner ridges more noticeable than the three outer pairs. Legs: procoxa scaly on infra-carinal and outer surface; punctures visible at 12
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
magnification; three protibial teeth, middle and apical equal in size, the three teeth equally spaced; protibial inner apical spur present; mesofemural disc setose, with a row of long bristles on anterior and posterior margins; mesotibia sub-quadrate in cross section; disc coarsely sculptured; mesotibial apical transverse carina may vary from partial to complete, in females always complete; basal apophysis of metacoxa produced beyond the outer margin of trochanter; metatibia with posterior discontinuous longitudinal carina; metatibial apical spurs equal in length, length equal to the diameter of the tibial apex; inner margin of male metatibia carinated towards apex, apical inner surface setose; metatibial disc coarsely
|
||
<pageBreakToken pageId="63" pageNumber="64" start="start">sculptured</pageBreakToken>
|
||
; two metatibial transverse carinae present posteriorly; basal metatarsomere and tarsomere II equal in size; in males protarsomere II long; pro- and mesotarsomeres I to IV enlarged, protarsomeres slightly wider than the mesotarsomeres, twice as wide as metatarsi; claw bifid, symmetrical, superior tooth longer and as wide as the inferior; distance between teeth as long as the inferior tooth. Abdomen: band of scales visible at the lowest magnification beneath the outer margin of elytra; disc of ventrites bearing sparse short bristles; propygidium visible, with scarce scales; pygidium generally flat, sub-quadrate, as wide as it is long; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc bristled throughout, coarsely punctured; pygidial apex sub-quadrate, more rounded in females. Parameres: basal region slightly narrower than the width of the parameres together at its transverse midline, parameral split at 2/3; total length of parameres near five times the length of their apex; inner margins straight, slightly divergent; apex harpoon-like with lateral angle curved projecting almost perpendicular to parameres (Fig. 74F). In lateral view parameres straight not coplanar with basal region (Fig. 74G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="63" pageNumber="64" type="type-locality">
|
||
<paragraph pageId="63" pageNumber="64">Type-locality.</paragraph>
|
||
<paragraph pageId="63" pageNumber="64">BRAZIL. Minas Gerais.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="63" pageNumber="64" type="distribution">
|
||
<paragraph pageId="63" pageNumber="64">Geographical distribution.</paragraph>
|
||
<paragraph pageId="63" pageNumber="64">BRAZIL (GO, DF, MT, MS, BA, MG, SP, PR); BOLIVIA (CB, SC); PARAGUAY (CN, AS, IT, GU); ARGENTINA (SA, JU, FO, CH, CO, SL).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="63" pageNumber="64" type="remarks">
|
||
<paragraph pageId="63" pageNumber="64">Remarks.</paragraph>
|
||
<paragraph pageId="63" pageNumber="64">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys suturalis" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="suturalis">Liogenys suturalis</taxonomicName>
|
||
together with
|
||
<taxonomicName lsidName="L. opacipennis" pageId="63" pageNumber="64" rank="species" species="opacipennis">L. opacipennis</taxonomicName>
|
||
Frey, 1969 and
|
||
<taxonomicName lsidName="L. rectangula" pageId="63" pageNumber="64" rank="species" species="rectangula">L. rectangula</taxonomicName>
|
||
Frey, 1969, are the only
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
species with body brownish and elytra opaque dull yellow.
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys suturalis" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="suturalis">Liogenys suturalis</taxonomicName>
|
||
is a closely related lineage to the clade
|
||
<taxonomicName lsidName="L. elegans" pageId="63" pageNumber="64" rank="species" species="elegans">L. elegans</taxonomicName>
|
||
+ (
|
||
<taxonomicName lsidName="L. tibialis" pageId="63" pageNumber="64" rank="species" species="tibialis">L. tibialis</taxonomicName>
|
||
+
|
||
<taxonomicName lsidName="L. punctaticollis" pageId="63" pageNumber="64" rank="species" species="punctaticollis">L. punctaticollis</taxonomicName>
|
||
+
|
||
<taxonomicName lsidName="L. spiniventris" pageId="63" pageNumber="64" rank="species" species="spiniventris">L. spiniventris</taxonomicName>
|
||
+
|
||
<taxonomicName lsidName="L. testaceipennis" pageId="63" pageNumber="64" rank="species" species="testaceipennis">L. testaceipennis</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Cherman, MA" journalOrPublisher="Systematic Entomology" pageId="104" pageNumber="105" pagination="744 - 770" title="Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville." url="https://doi.org/10.1111/syen.12188" volume="41" year="2016">Cherman et al. 2016</bibRefCitation>
|
||
), and differs from them mainly in the elytra pruinose and the abundant scales covering the pro-, meso- and metasternum laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |