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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.664.11881" ID-GBIF-Dataset="5affa7d2-a73b-4025-bd9e-a318295516c7" ID-PMC="PMC5523158" ID-Pensoft-Pub="1313-2970-664-1" ID-PubMed="28769623" ID-ZBK="C86AA26D022948D8A36E5BBBE871F7EA" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-664-1" ModsDocOrigin="ZooKeys 664" ModsDocTitle="A revision of the Oriental species of Bolitogyrus Chevrolat (Coleoptera, Staphylinidae, Staphylininae)" checkinTime="1490795552514" checkinUser="pensoft" docAuthor="Brunke, Adam J." docDate="2017" docId="3717A13BB9A09890137DB69D9F3060C5" docLanguage="en" docName="ZooKeys 664: 1-97" docOrigin="ZooKeys 664" docSource="http://dx.doi.org/10.3897/zookeys.664.11881" docTitle="Bolitogyrus nanus Brunke, sp. n." docType="treatment" docUuid="BF37C021-CD3F-40E5-8AC1-88F6F4AB6BC3" docUuidSource="ZooBank" docVersion="5" lastPageNumber="59" masterDocId="1C11EF08FFA0FF81FFDBB829D66FE206" masterDocTitle="A revision of the Oriental species of Bolitogyrus Chevrolat (Coleoptera, Staphylinidae, Staphylininae)" masterLastPageNumber="97" masterPageNumber="1" pageNumber="58" updateTime="1668164339952" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Oriental species of Bolitogyrus Chevrolat (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brunke, Adam J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>664</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>97</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.664.11881</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.664.11881</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-664-1</mods:identifier>
<mods:identifier type="ZBK">C86AA26D022948D8A36E5BBBE871F7EA</mods:identifier>
<mods:identifier type="ZooBank">C86AA26D022948D8A36E5BBBE871F7EA</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="128090122" LSID="urn:lsid:zoobank.org:act:BF37C021-CD3F-40E5-8AC1-88F6F4AB6BC3" httpUri="http://treatment.plazi.org/id/3717A13BB9A09890137DB69D9F3060C5" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<subSubSection pageId="57" pageNumber="58" type="nomenclature">
<paragraph pageId="57" pageNumber="58">
<taxonomicName LSID="http://zoobank.org/BF37C021-CD3F-40E5-8AC1-88F6F4AB6BC3" authority="Brunke" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus nanus" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nanus">Bolitogyrus nanus Brunke</taxonomicName>
<taxonomicNameLabel pageId="57" pageNumber="58">sp. n.</taxonomicNameLabel>
Fig. 17
<normalizedToken originalValue="KL">K-L</normalizedToken>
, 20C (map)
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="type locality">
<paragraph pageId="58" pageNumber="59">
<pageBreakToken pageId="58" pageNumber="59" start="start">Type</pageBreakToken>
locality.
</paragraph>
<paragraph pageId="58" pageNumber="59">Mawsynram-Balat Road, Khasi Hills, Meghalaya, India.</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="type material">
<paragraph pageId="58" pageNumber="59">Type material.</paragraph>
<paragraph pageId="58" pageNumber="59">
Holotype (♂, MHNG): INDIA, Meghalaya, Khasi Hills, 1000m, Mawsynram-Balat,
<normalizedToken originalValue="Besuchet-Löbl">Besuchet-Loebl</normalizedToken>
, 27.X.78, [1978] [printed] /
<taxonomicName class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">Cyrtothorax</taxonomicName>
spec. det. M. Uhlig 1982 [written] /
<taxonomicName class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax vulneratus" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="vulneratus">Cyrtothorax vulneratus</taxonomicName>
Smetana det. 1986 [written] / HOLOTYPE
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus nanus" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nanus">Bolitogyrus nanus</taxonomicName>
Brunke, des. A. Brunke 2017 [red label] / AJB0000431 [identifier label].
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="diagnosis">
<paragraph pageId="58" pageNumber="59">Diagnosis.</paragraph>
<paragraph pageId="58" pageNumber="59">
This species can be distinguished by the following character states: head entirely dark; elytral disc dark with pale markings extending halfway to epipleural margin (Fig. 5G); pronotal margin at its widest point no more than three lateral puncture widths wide but still distinctly expanded at hind angles; forebody less than four millimeters; apex of median lobe with double-toothed carina or
<normalizedToken originalValue="lip">'lip'</normalizedToken>
(Fig. 17L); paramere with only simple rows of peg setae and without dense field (Fig. 17K).
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="description">
<paragraph pageId="58" pageNumber="59">Description.</paragraph>
<paragraph pageId="58" pageNumber="59">Measurements ♂ (n = 1): HW/HL 1.40; PW/PL 1.29; EW/ EL 1.23; ESut/PL 0.84; PW/HW 1.01; forebody length 3.5 mm.</paragraph>
<paragraph pageId="58" pageNumber="59">
Extremely similar to
<taxonomicName lsidName="B. khasiensis" pageId="58" pageNumber="59" rank="species" species="khasiensis">B. khasiensis</taxonomicName>
and differing only in the following: medial and lateral elytral markings more broadly connected, pale area of epipleuron not restricted to humeral spot, pale in up to entire basal half; pronotum slightly more transverse; elytral suture relatively longer than pronotum at middle; forebody distinctly shorter and thinner; median lobe in parameral view with apex distinctly acuminate and acute, apex with folded, double-tooted carina (Fig. 17L); paramere with sparse and simple rows of peg setae placed close to margin, removed from margin only at base of rows (Fig. 17K). Female unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="distribution">
<paragraph pageId="58" pageNumber="59">Distribution.</paragraph>
<paragraph pageId="58" pageNumber="59">Figure 20C. Known only from the Khasi Hills of Meghalaya.</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="bionomics">
<paragraph pageId="58" pageNumber="59">Bionomics.</paragraph>
<paragraph pageId="58" pageNumber="59">The holotype was collected in October at 1000 m.</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="etymology">
<paragraph pageId="58" pageNumber="59">Etymology.</paragraph>
<paragraph pageId="58" pageNumber="59">The species epithet refers to the fact that this species is the smallest known in the Oriental fauna at 3.5 millimeters in forebody length.</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="comments">
<paragraph pageId="58" pageNumber="59">Comments.</paragraph>
<paragraph pageId="58" pageNumber="59">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus nanus" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nanus">Bolitogyrus nanus</taxonomicName>
is extremely similar to allopatric
<taxonomicName lsidName="B. himalayicus" pageId="58" pageNumber="59" rank="species" species="himalayicus">B. himalayicus</taxonomicName>
from West Bengal but is slightly smaller and the apex of the median lobe bears a double-toothed carina and the paramere lacks the dense apical field of peg setae.
</paragraph>
</subSubSection>
</treatment>
</document>