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<document id="82AB84D5FE446A73089698FBC8D1CB8C" ID-DOI="10.5281/zenodo.156636" ID-GBIF-Dataset="c9e80f8e-36cc-4c6b-bfb8-bc53428ee8f4" ID-ISSN="1175­5326" ID-Zenodo-Dep="156636" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459622659579" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2003" docId="36558784E16AAC2BFE80FE17FBA416CD" docLanguage="en" docName="zt00263.pdf" docOrigin="Zootaxa 263" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Haplohyphes yanahuicsa Molineri, 2003, sp. nov." docType="treatment" docVersion="10" lastPageNumber="8" masterDocId="CA6CFFFCE16FAC2CFF88FFF3FFAE132B" masterDocTitle="Haplohyphes (Ephemeroptera: Leptohyphidae), new species and stage descriptions with a key to separate the species of the genus" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1698203766015" updateUser="plazi">
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<mods:title id="F4681FAF1557BF7B9B4C9126328CB359">Haplohyphes (Ephemeroptera: Leptohyphidae), new species and stage descriptions with a key to separate the species of the genus</mods:title>
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<mods:namePart id="AA8132563512E0DFAC661FF656AE2018">Molineri, Carlos</mods:namePart>
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<mods:date id="56547521170CA38D30A1C71A8E284F8E">2003</mods:date>
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<subSubSection id="F6E66519E16AAC29FE80FE17FD061752" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="BE433692E16AAC29FE80FE17FC3A12D6" blockId="5.[264,916,483,510]" box="[264,916,483,510]" pageId="5" pageNumber="6">
<heading id="E50B81FEE16AAC29FE80FE17FC3A12D6" bold="true" box="[264,916,483,510]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="8C88EA80E16AAC29FE80FE17FC3A12D6" bold="true" box="[264,916,483,510]" pageId="5" pageNumber="6">
<taxonomicName id="79FC4D11E16AAC29FE80FE17FD9412D5" box="[264,570,484,510]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yanahuicsa" status="sp. nov.">
<emphasis id="8C88EA80E16AAC29FE80FE17FD9412D5" bold="true" box="[264,570,484,510]" italics="true" pageId="5" pageNumber="6">Haplohyphes yanahuicsa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97BB57FBE16AAC29FDC9FE10FD3012D6" box="[577,670,483,509]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="26C72A17E16AAC29FD26FE10FC9612D6" box="[686,824,483,509]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="5" pageNumber="6">Figs. 20­41</figureCitation>
,
<figureCitation id="26C72A17E16AAC29FCCEFE10FC2512D6" box="[838,907,483,509]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="5" pageNumber="6">46­47</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="BE433692E16AAC29FE80FDDCFD061752" blockId="5.[264,1324,559,1225]" pageId="5" pageNumber="6">
Male imago. Length: 4.0­
<quantity id="79049B77E16AAC29FDB2FDDCFD341162" box="[570,666,559,585]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="6" unit="mm" value="4.2">4.2 mm</quantity>
; fore wings,
<quantity id="79049B77E16AAC29FCB0FDDCFC691162" box="[824,967,559,585]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="5" pageNumber="6" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.7­3.9 mm</quantity>
. General coloration yellowish white, last segments of abdomen blackish. Head shaded with brownish gray from a transverse line at the median ocellus to posterior margin; antennae yellowish white. Thorax. Pronotum whitish translucent shaded almost completely with brownish gray, meso­ and metanotum whitish yellow with few very light gray marks; pleurae and sterna whitish yellow translucent shaded with light brownish on pro­ and mesopleurae and with gray in thoracic sterna. Legs: coxae I and II, fore femora and apex of fore tibiae and fore tarsi shaded with brownish gray, remaining segments of all legs whitish. Wings (
<figureCitation id="26C72A17E16AAC29FBCCFCB4FB07104A" box="[1092,1193,839,865]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="5" pageNumber="6">Figs. 20</figureCitation>
a­c): membrane of wings hyaline, fore wings shaded with gray on C and Sc area, basally to bullae; longitudinal vein grayish, cross veins paler; hind wings only with two longitudinal veins. Abdomen (
<figureCitation id="26C72A17E16AAC29FE06FC4CFE4910F2" box="[398,487,959,985]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="5" pageNumber="6">Fig. 46</figureCitation>
) whitish translucent shaded with blackish gray on terga, shaded very slightly on first segments and becoming darker and more extensive toward rear segments (segments VI­IX are much darker than the rest of body). Genitalia (
<figureCitation id="26C72A17E16AAC29FB92FBFCFB0B1702" box="[1050,1189,1039,1065]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="5" pageNumber="6">Figs. 21­24</figureCitation>
): yellowish white shaded with gray on margins of styliger plate and forceps. Caudal filaments hyaline shaded with gray, except at joining.
</paragraph>
</subSubSection>
<subSubSection id="F6E66519E16AAC2BFEB0FB74FBA416CD" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="BE433692E16AAC29FEB0FB74FC1417E2" blockId="5.[264,1324,559,1225]" pageId="5" pageNumber="6">
Female imago. Unknown, but two mature female nymphs ready to moult (
<figureCitation id="26C72A17E16AAC29FB41FB74FA8D178A" box="[1225,1315,1159,1185]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="5" pageNumber="6">Fig. 47</figureCitation>
) show the same characteristic coloration of the male imago.
</paragraph>
<caption id="EA83661AE16AAC29FE80F995FEF81400" pageId="5" pageNumber="6">
<paragraph id="BE433692E16AAC29FE80F995FEF81400" blockId="5.[264,1323,1638,1835]" pageId="5" pageNumber="6">
<emphasis id="8C88EA80E16AAC29FE80F995FE651555" bold="true" box="[264,459,1638,1662]" pageId="5" pageNumber="6">FIGURES 20­41.</emphasis>
<taxonomicName id="79FC4D11E16AAC29FE5BF994FD4E1555" box="[467,736,1639,1662]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yanahuicsa" status="sp. nov.">
<emphasis id="8C88EA80E16AAC29FE5BF994FD4E1555" box="[467,736,1639,1662]" italics="true" pageId="5" pageNumber="6">Haplohyphes yanahuicsa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97BB57FBE16AAC29FD60F995FC931556" box="[744,829,1638,1661]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, male imago: 20a, fore wing; 20b, hind wing; 20c, idem, detail; 21, male genitalia, v.v.; 22, idem, l.v.; 23a­b, penes, details v.v.; 24, detail of forceps, l.v.; nymph: 25, head, d.v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30a, labium, v.v.; 30b, detail of labial palp; 31, labrum, d.v.; 32, hypopharynx, v.v.; 33, left mandible, d.v.; 34, right mandible, d.v.; 35a, maxillae, d.v.; 35b, detail of maxillary palp; 36, gill II, d.v.; 37­41, gills II­ VI, v.v.
</paragraph>
</caption>
<paragraph id="BE433692E168AC2BFEB0FEEFFECD104D" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">
Nymph (
<figureCitation id="26C72A17E168AC2BFE2AFEEFFE52121D" box="[418,508,284,310]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 47</figureCitation>
). Length: body, 5.0­
<quantity id="79049B77E168AC2BFD78FEEFFCE1121D" box="[752,847,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="8" unit="mm" value="5.1">5.1 mm</quantity>
; mesonotum,
<quantity id="79049B77E168AC2BFC7FFEEFFBF8121D" box="[1015,1110,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="mm" value="1.5">1.5 mm</quantity>
; hind femora, 1.0 mm. General coloration whitish yellow with blackish marks. Head yellowish white surrounded by thick yellowish setae; shaded with blackish gray on a transversal band between eyes; occipute shaded more slightly with a reticular pattern; antennae whitish translucent; mouth parts whitish. Mouthparts as in
<figureCitation id="26C72A17E168AC2BFD7BFE4FFCD512FD" box="[755,891,444,470]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Figs 30­35</figureCitation>
. Thorax whitish yellow, pronotum shaded extensively with blackish gray, mesonotum shaded more slightly; wing buds shaded with black on basal half of costal margin. Thoracic sterna shaded with gray, metanotum and pleurae without shading. Legs (
<figureCitation id="26C72A17E168AC2BFD64FDC7FCDD1165" box="[748,883,564,590]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Figs. 26­28</figureCitation>
) whitish shaded with gray on coxae I and II; tarsal claws with double row of submarginal denticles near apex (
<figureCitation id="26C72A17E168AC2BFBF8FDAFFB64115D" box="[1136,1226,604,630]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Fig. 29</figureCitation>
). Abdomen (
<figureCitation id="26C72A17E168AC2BFEC3FD77FE0F11B5" box="[331,417,644,670]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 47</figureCitation>
) yellowish white shaded with light gray on terga I­IV (sometimes also on V) and shaded with blackish gray on terga VI­IX (sometimes also on V); tergum X slightly shaded with gray. Abdominal sterna paler shaded very slightly with gray. Opercullar gill (
<figureCitation id="26C72A17E168AC2BFE99FD0FFE37103D" box="[273,409,764,790]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Figs. 36­37</figureCitation>
) whitish translucent shaded completely with light gray, remaining gills (
<figureCitation id="26C72A17E168AC2BFB78FD0FFEE31015" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Figs. 38­41</figureCitation>
) whitish translucent. Caudal filaments whitish yellow with whorls of setae at each joining.
</paragraph>
<paragraph id="BE433692E168AC2BFEB0FC87FDA2109D" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">Etymology. Combination of letters including the Quechua voices “yana” (=black) and “huicsa” (=abdomen).</paragraph>
<paragraph id="BE433692E168AC2BFEB0FC37FEF7178D" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">
Material.
<typeStatus id="61478830E168AC2BFE24FC37FDB710F5" box="[428,537,964,990]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
male imago from
<collectingCountry id="C6EB7602E168AC2BFD74FC37FCD410F5" box="[764,890,964,990]" name="Bolivia" pageId="7" pageNumber="8">BOLIVIA</collectingCountry>
: Dept. La Paz, stream between Caranavi and Guanai,
<geoCoordinate id="DBC85055E168AC2BFE6FFC1FFD29172D" box="[487,647,1004,1030]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-15.671667">S 15° 40' 18&quot;</geoCoordinate>
­
<geoCoordinate id="DBC85055E168AC2BFD29FC1FFCE2172D" box="[673,844,1004,1030]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="-67.70111">W 67° 42' 04&quot;</geoCoordinate>
,
<quantity id="79049B77E168AC2BFCECFC1FFC1C172D" box="[868,946,1004,1030]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="m" value="500.0">500 m</quantity>
,
<date id="CA421052E168AC2BFC37FC1FFBE7172D" box="[959,1097,1004,1030]" pageId="7" pageNumber="8" value="2000-11-27">27/XI/2000</date>
, E. Domínguez, C. Molineri y C. Nieto Cols.
<typeStatus id="61478830E168AC2BFDCDFBE7FD121705" box="[581,700,1044,1070]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A8FAFD1BE168AC2BFD40FBE7FCB41705" box="[712,794,1044,1070]" pageId="7" pageNumber="8" type="male">3 male</specimenCount>
imagos and
<specimenCount id="A8FAFD1BE168AC2BFC3DFBE7FBB31705" box="[949,1053,1044,1070]" pageId="7" pageNumber="8" type="female">2 female</specimenCount>
mature nymphs, same data as
<typeStatus id="61478830E168AC2BFEE1FBCFFE7D177D" box="[361,467,1084,1110]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
.
<typeStatus id="61478830E168AC2BFE67FBCFFDF1177D" box="[495,607,1084,1110]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
, 1
<typeStatus id="61478830E168AC2BFD02FBCFFD41177D" box="[650,751,1084,1110]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
male imago and 1
<typeStatus id="61478830E168AC2BFC6EFBCFFBE5177D" box="[998,1099,1084,1110]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
nymph housed in UMSA; remaining material in IFML. Nymphs and adults are associated by a pharate subimago.
</paragraph>
<paragraph id="BE433692E168AC2BFEB0FB47FBA416CD" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">
Discussion.
<taxonomicName id="79FC4D11E168AC2BFE4EFB47FD4317E6" box="[454,749,1204,1229]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yanahuicsa" status="sp. nov.">
<emphasis id="8C88EA80E168AC2BFE4EFB47FD4317E6" box="[454,749,1204,1229]" italics="true" pageId="7" pageNumber="8">Haplohyphes yanahuicsa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97BB57FBE168AC2BFD7DFB47FCE117E5" box="[757,847,1204,1230]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from the other species of the genus by the following combination of characters. In the imago: 1) hind wings with two longitudinal veins (
<figureCitation id="26C72A17E168AC2BFDEFFAF7FD6C1635" box="[615,706,1284,1310]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Fig. 20</figureCitation>
c); 2) segment 1 and 2 of forceps similar in length (
<figureCitation id="26C72A17E168AC2BFE99FADFFEC5166D" box="[273,363,1324,1350]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Fig. 24</figureCitation>
); 3) penes divided on apical 0.22 (
<figureCitation id="26C72A17E168AC2BFC9CFADFFCD9166D" box="[788,887,1324,1350]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Figs. 23</figureCitation>
a­b); 4) membranous lobes of penes rounded (
<figureCitation id="26C72A17E168AC2BFEF3FAA7FE7D1645" box="[379,467,1364,1390]" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Fig. 23</figureCitation>
a­b); 5) distal spines of penes shorter than membranous lobes; 6) male fore wing length: 4.0 mm; 7) general coloration pale except apical segments of abdomen blackish (
<figureCitation id="26C72A17E168AC2BFEF2FA57FE7D1695" box="[378,467,1444,1470]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 46</figureCitation>
). In the nymph: 1) markings on head not reaching base of antennae (
<figureCitation id="26C72A17E168AC2BFB73FA57FE8716CD" captionStart="FIGURES 20 ­ 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 ­ 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 a­b, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 ­ 41, gills II­ VI, v. v." pageId="7" pageNumber="8">Fig. 25</figureCitation>
); 2) abdomen shaded with blackish on rear segments (
<figureCitation id="26C72A17E168AC2BFC2DFA3FFC5216CD" box="[933,1020,1484,1510]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>