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<mods:title id="189132A7A5A50E0507C7A7F235219EAA">Taxonomic revision of genus Ablattaria Reitter (Coleoptera, Silphidae) using geometric morphometrics</mods:title>
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<mods:namePart id="2970121FDCD6B129C8746D855EAFCF0C">Ruzicka, Jan</mods:namePart>
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<treatment id="20CF6EA6BB3BEE77746899E06C38D0FA" ID-GBIF-Taxon="152057996" LSID="urn:lsid:plazi:treatment:20CF6EA6BB3BEE77746899E06C38D0FA" httpUri="http://treatment.plazi.org/id/20CF6EA6BB3BEE77746899E06C38D0FA" lastPageId="10" lastPageNumber="89" pageId="7" pageNumber="86">
<subSubSection id="E8D34CFFE975D995F49960F7CBBE12A5" pageId="7" pageNumber="86" type="multiple">
<paragraph id="6E98E3B89AD6AC1DC4242E4F947034BC" pageId="7" pageNumber="86">
<pageBreakToken id="0EA0023ED06DCF0502F63A79D74619FC" pageId="7" pageNumber="86" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Silphidae
</paragraph>
</subSubSection>
<subSubSection id="F46B0D85B2D8CACA0B9037AF9CEEA1E1" pageId="7" pageNumber="86" type="nomenclature">
<paragraph id="616FA55FB2347B42E6305EF27B91B6E3" pageId="7" pageNumber="86">
<taxonomicName id="FD53D4854F28FFD42689006FC6D986FF" ID-CoL="9MKQQ" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata (Fabricius, 1775)</taxonomicName>
Figs 4, 5-8, 9-11, 15, 18, 21, 22, 26, 28-31
</paragraph>
</subSubSection>
<subSubSection id="D4438D552D3B76CA60C26BE74E19B233" pageId="7" pageNumber="86" type="reference_group">
<paragraph id="5BDCEFDB7E3FC6C21508F622ED3E5D90" pageId="7" pageNumber="86">
<taxonomicName id="7D5183B5D30EF96AC810B051472E0FBA" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
<bibRefCitation id="E1A748F979B990B3B1373AB2C529F454" author="Fabricius, JC" journalOrPublisher="Libraria Kortii, Flensburgi et Lipsiae" pageId="20" pageNumber="99" title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus" year="1775">Fabricius 1775</bibRefCitation>
: 74 (with reference to
<bibRefCitation id="8CE955BB53C7AC00BB789CE1D3FA0B21" author="Geoffroy, E-L" journalOrPublisher="Durand, Paris" pageId="21" pageNumber="100" title="Histoire abregee des Insectes qui se trouvent aux environs de Paris: Dans laquelle ces Animaux sont ranges suivant un ordre methodique" year="1762">Geoffroy (1762)</bibRefCitation>
) (type locality: Europe [&quot;Habitat in sylvis Europae&quot;]).
</paragraph>
<paragraph id="042BA0810B231F8405B3C5BEE75E9426" pageId="7" pageNumber="86">
<taxonomicName id="9B03D75217BC2ACA29DAE9B0B95359E1" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha polita" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="polita">Silpha polita</taxonomicName>
<bibRefCitation id="2BB2D89D7B4DBA1EBB025E5858764CA6" author="Sulzer, JH" journalOrPublisher="H. Steiner und Comp., Winterthur" pageId="23" pageNumber="102" title="Abgekuerzte Geschichte der Insecten nach dem Linaeischen System" year="1776">Sulzer 1776</bibRefCitation>
: 28 (with reference to
<bibRefCitation id="B33F54605F684DD54F2A3A913D8C8C54" author="Geoffroy, E-L" journalOrPublisher="Durand, Paris" pageId="21" pageNumber="100" title="Histoire abregee des Insectes qui se trouvent aux environs de Paris: Dans laquelle ces Animaux sont ranges suivant un ordre methodique" year="1762">Geoffroy (1762)</bibRefCitation>
) (type locality:
<normalizedToken id="1BCBCEBFEA8BA26CC7664DFB03E74FDA" originalValue="“Schweiz”">&quot;Schweiz&quot;</normalizedToken>
[Switzerland]; preoccupied, not
<bibRefCitation id="33F86CB401A61F7E8CC39168D542B04A" author="Fuessli, JC" journalOrPublisher="Fuessly und Winterthur, H. Steiner, Zuerich" pageId="20" pageNumber="99" title="Verzeichnis der ihm bekannten Schweitzerischen Inseckten mit einer ausgemahlten Kupfertafel: nebst der Ankuendigung eines neuen Insecten Werks" year="1775">
<normalizedToken id="789B895171292797C4FEBD001F284502" originalValue="Fueßli">Fuessli</normalizedToken>
1775
</bibRefCitation>
: 6 (junior primary homonym)) (as junior synonym of
<taxonomicName id="8FDEBB3D600D953FD0EAA4F85A7C9DE3" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="subSpecies" species="laevigata" subSpecies="laevigata">Ablattaria laevigata laevigata</taxonomicName>
by
<bibRefCitation id="7A952626F5FEECDFA3FFBE63CEA6EAD3" author="Reitter, E" journalOrPublisher="Verhandlungen des Naturforschenden Vereines in Bruenn" pageId="22" pageNumber="101" pagination="3 - 122" title="Bestimmungs-Tabellen der europaeischen Coleopteren. XII. Necrophaga (Platypsyllidae, Leptinidae, Silphidae, Anisotomidae und Clambidae)." volume="23" year="1884">Reitter 1884</bibRefCitation>
: 75, confirmed by
<bibRefCitation id="65A5FFEE2A29C57EEB6C8BB6D95DFD4C" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller 1979</bibRefCitation>
: 5).
</paragraph>
<paragraph id="9B471562508A8028B064C63C1F52BB20" pageId="7" pageNumber="86">
<taxonomicName id="3314E016E604BB78BDB2926FB361DD2B" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Silpha gibba</taxonomicName>
<bibRefCitation id="B761A30FDAF663B1F34886EFE65CD98E" author="Brulle, GA" journalOrPublisher="F. G. Levrault, Paris" pageId="20" pageNumber="99" title="Expedition scientifique de Moree. Section des sciences physiques, Tome III, 2 e Partie, Insectes" year="1832">
<normalizedToken id="30D96643D6567AFD1D673362DA02D43D" originalValue="Brullé">Brulle</normalizedToken>
1832
</bibRefCitation>
: 162 (type locality:
<normalizedToken id="B8D68743EB101F8FF26064CE4B2A0825" originalValue="“Arcadie”">&quot;Arcadie&quot;</normalizedToken>
[= Greece: Peloponnese Peninsula, Arcadia region]) (as junior synonym of
<taxonomicName id="E28D5F2480C0E18047857969071F2435" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
by
<bibRefCitation id="DDF9FE2233D6B4AA5CAA1E21D154BF18" author="Nikolaev, GV" journalOrPublisher="Almaty, Kazak universiteti" pageId="21" pageNumber="100" title="Zhuky-mertvoedy (Coleoptera: Agyrtidae, Silphidae) Kazakhstana, Rossii i ryada sopredel'nykh stran: Opredelitel. (The carrion beetles (Coleoptera: Agyrtidae, Silphidae) of Kazakhstan, Russia and adjacent countries)." year="2002">Nikolaev and Kozminykh 2002</bibRefCitation>
: 70).
</paragraph>
<paragraph id="4A2F8464F613A3B16B360998E735B826" pageId="7" pageNumber="86">
<taxonomicName id="0703972E14F170A5490280A3FA59CEEF" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="subSpecies" species="laevigata" subSpecies="gibba">Ablattaria laevigata gibba</taxonomicName>
:
<bibRefCitation id="AD6E424C23EEC6F2B8EFCE671CDD093D" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller 1979</bibRefCitation>
: 5.
</paragraph>
<paragraph id="FF71BD8F40DE0A21B74EEC09037A76A4" pageId="7" pageNumber="86">
<taxonomicName id="9C812897085ACCB1EA521B722741E765" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. costulata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="costulata">Ablattaria gibba var. costulata</taxonomicName>
<bibRefCitation id="9585A8C0E42FEFBF85A93B2A775B93BE" author="Portevin, G" journalOrPublisher="Lechevalier, Paris" pageId="22" pageNumber="101" title="Les Grands Necrophages du Globe. Silphini - Necrodini - Necrophorini." volumeTitle="Encyclopedie Entomologique" year="1926">Portevin 1926</bibRefCitation>
: 25 (type locality: Istanbul [ca.
<geoCoordinate id="51710E38717FCAD6F11593C5008B7117" direction="north" orientation="latitude" precision="925" value="41.016666">41°01'N</geoCoordinate>
,
<geoCoordinate id="B5D4FCB384A2940A4032BBCC3C9942BD" direction="east" orientation="longitude" precision="925" value="28.95">28°57'E</geoCoordinate>
]) (as junior synonym of
<taxonomicName id="F25BC8DAEBF8E910900D855277128185" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="subSpecies" species="laevigata" subSpecies="gibba">Ablattaria laevigata gibba</taxonomicName>
by
<bibRefCitation id="A40EA7A8AFB685B64D23A9970A0B8A7A" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller 1979</bibRefCitation>
: 5).
</paragraph>
<paragraph id="A0D49C1DD4D92EAF8C0462E1F6704BC8" pageId="7" pageNumber="86">
<taxonomicName id="103DF7AF360A7AAFB88F025263548E92" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. distinguenda" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="distinguenda">Ablattaria gibba var. distinguenda</taxonomicName>
<bibRefCitation id="9A9E97B8A1542EC094E65A214E209BB2" author="Portevin, G" journalOrPublisher="Lechevalier, Paris" pageId="22" pageNumber="101" title="Les Grands Necrophages du Globe. Silphini - Necrodini - Necrophorini." volumeTitle="Encyclopedie Entomologique" year="1926">Portevin 1926</bibRefCitation>
: 25 (type locality not stated) (as junior synonym of
<taxonomicName id="F996AFDBE05E503EC6DA37F820DDE856" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="subSpecies" species="laevigata" subSpecies="gibba">Ablattaria laevigata gibba</taxonomicName>
by
<bibRefCitation id="67450DAD8D97E6945A7CBF8EEBC91223" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller 1979</bibRefCitation>
: 5).
</paragraph>
<paragraph id="0CA9EA73F6450AC3AA5A8E195D50A6CC" pageId="7" pageNumber="86">
<taxonomicName id="B4D20F595BC55B132C6212B0C0574B4C" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. punctata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="punctata">Ablattaria gibba var. punctata</taxonomicName>
<bibRefCitation id="A4581C4D1B5ACDB69FB1EC8857F804F8" author="Portevin, G" journalOrPublisher="Lechevalier, Paris" pageId="22" pageNumber="101" title="Les Grands Necrophages du Globe. Silphini - Necrodini - Necrophorini." volumeTitle="Encyclopedie Entomologique" year="1926">Portevin 1926</bibRefCitation>
: 26 (type locality not stated) (as junior synonym of
<taxonomicName id="670D31A8A595FFCE132B734D2481D607" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="subSpecies" species="laevigata" subSpecies="gibba">Ablattaria laevigata gibba</taxonomicName>
by
<bibRefCitation id="6B6B7355644D5C360995FF786DF2F883" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller 1979</bibRefCitation>
: 5).
</paragraph>
<paragraph id="FA2BA3D76EC247069155F9C9B441A473" pageId="7" pageNumber="86">
<taxonomicName id="AA53FB3503CC6AABFDCEBA39E4085946" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata var. meridionalis" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="laevigata" variety="meridionalis">Ablattaria laevigata var. meridionalis</taxonomicName>
<bibRefCitation id="E174072432CC8C562E4EE17CCDDB7763" author="Ganglbauer, L" journalOrPublisher="C. Gerold's Sohn, Wien" pageId="20" pageNumber="99" title="Die Kaefer von Mitteleuropa. Die Kaefer der oesterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzoesischen und italienischen Alpengebietes. Dritter Band, Familienreihe Staphylinoidea, II. Theil. Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeriidae, Trichopterygidae, Hydroscaphidae, Scaphidiidae, Histeridae. Familienreihe Clavicornia. Sphaeritidae, Ostomidae, Byturidae, Nitidulidae, Cucuidae, Erotylidae, Phalacridae, Thorictidae, Lathridiidae, Mycetophagidae, Colydidae, Endomychidae, Coccinellidae" year="1899">Ganglbauer 1899</bibRefCitation>
: 191 (type locality: &quot;Illirien, Dalmatien,
<normalizedToken id="73FB34A814689445C46747FC63AE86D0" originalValue="Südungarn">Suedungarn</normalizedToken>
und Griechenland&quot; [= Illiria, Dalmatia, southern Hungary and Greece], syn. n.
</paragraph>
<paragraph id="48DC51CC762A20C96C725FA1806A84F8" pageId="7" pageNumber="86">
<taxonomicName id="9B6B0EB9FBE2780B4ADAE95A49F7FFA2" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="subSpecies" species="laevigata" subSpecies="meridionalis">Ablattaria laevigata meridionalis</taxonomicName>
:
<bibRefCitation id="5D20A9B45083F85446BAD4391E5D3E65" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller 1979</bibRefCitation>
: 5.
</paragraph>
</subSubSection>
<subSubSection id="DD4A42045C524E7E06CA18A571CF7254" pageId="7" pageNumber="86" type="type material examined">
<paragraph id="7433139A46BF4F25742EF63C9C2E74AB" pageId="7" pageNumber="86">Type material examined.</paragraph>
<paragraph id="542A2B7F8684DFE24D235AE1514E14A5" pageId="7" pageNumber="86">
Lectotype female of
<taxonomicName id="988F8B2467B3165D915924AB580B952E" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
(here designated) (ZMUC, Collection of Ove Ramel Sehested and Niels
<normalizedToken id="F9088FF8243F21068526FFA025BBE951" originalValue="Tønder">Tonder</normalizedToken>
Lund, the &quot;Copenhagen collection&quot; [= coll. S &amp; TL]) (Figs 28-29), pinned, labelled: &quot;Lectotype /
<taxonomicName id="D7E991AED0E852A4CB16098587EB0ADF" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Silpha</taxonomicName>
/
<taxonomicName id="BE8EE8A2EB3AB9FA7E6B03B89043A825" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ Fabricius, 1775 / Jan
<normalizedToken id="24384AAAADAFB7CFEB980E22176E47B6" originalValue="Růžička">Ruzicka</normalizedToken>
des. 2012 [p, red label] //
<taxonomicName id="890B06F65FE9ED2515F765EEA03B16EF" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="68EA3F4239AB06E4C21E84D21FDBCC81" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ (Fabricius, 1775) / Jan
<normalizedToken id="26ACFEB389284ED5DC8A4466ED9CAFD5" originalValue="Růžička">Ruzicka</normalizedToken>
det. 20 [p] 12 [hw] // zmuc / 00021148 [p, this and subsequent numbers below are associated with photodocumentation]&quot;. Paralectotypes: 1 male (ZMUC, coll. S &amp; TL), pinned, labelled &quot;Paralectotype /
<taxonomicName id="FA4AE78A9E1FB80A68755AD5DBD7ADF7" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Silpha</taxonomicName>
/
<taxonomicName id="8B277A96FFB5E476CA4BBACFCAB7927E" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ Fabricius, 1775 / Jan
<normalizedToken id="46F4BB6C5173317B3260E8807D24C245" originalValue="Růžička">Ruzicka</normalizedToken>
des. 2012 [p, red label] //
<taxonomicName id="CFA66E518C9A4508B831D794AF006A8D" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="835F3F1C5A9F8EA1D283A09316608C24" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ (Fabricius, 1775) / Jan
<normalizedToken id="743C1E76D8358A65AFD902A01182CED9" originalValue="Růžička">Ruzicka</normalizedToken>
det. 20 [p] 12 [hw] // zmuc / 00021149 [p]&quot;; 1 male and 1 female (ZMUC, Fabricius personal collection, the &quot;Kiel collection&quot;), pinned, identical labels as previous except for &quot;laeviga / ta [hw,
<normalizedToken id="F6136C29DDC47FF7FA5560AFD8CFB1A3" originalValue="Fabricius’s">Fabricius's</normalizedToken>
handwriting, label pinned in box left to the first specimen]&quot; and &quot;zmuc00021468 [p]&quot; or &quot;zmuc00021469 [p]&quot;; 1 female of
<taxonomicName id="77A5302ED0259E1C56F8736935D7A3FF" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha tyrolensis" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="tyrolensis">Silpha tyrolensis</taxonomicName>
Laicharting, 1781 (ZMUC, &quot;Kiel collection&quot;) (Figs 30-31), pinned, labelled &quot;Paralectotype /
<taxonomicName id="CF450E93D290DE3CF98C98177673B9B2" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Silpha</taxonomicName>
/
<taxonomicName id="1152C5748C14EACD286139555382AFD2" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ Fabricius, 1775 / Jan
<normalizedToken id="2F0D9AE90F2A48DBB0B2F976FD5358C0" originalValue="Růžička">Ruzicka</normalizedToken>
des. 2012 [p, red label] //
<taxonomicName id="B1C9F79C5E2C91B839547CB0ADCBB430" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Silpha</taxonomicName>
/
<taxonomicName id="3E33DF661E9F20FD093C238A045AF226" lsidName="tyrolensis" pageId="7" pageNumber="86" rank="species" species="tyrolensis">tyrolensis</taxonomicName>
/ Laicharting, 1781 / Jan
<normalizedToken id="4D17B7F1D77E753B9E712AF920523CC5" originalValue="Růžička">Ruzicka</normalizedToken>
det. 20 [p] 12 [hw] // zmuc00021467 [p]&quot;; 1 male (BMNH, coll. Banks), pinned, labelled: &quot;HOLO- / TYPE [p, modern round label with thick red margin] //
<taxonomicName id="7CCB7B4AEF2D31D3FD1F15A6DED097EA" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
[hw] / Fab. Entom. p. [p] 74. n / 10 [hw, double black frame] // Paralectotype /
<taxonomicName id="EED3E537894059DEBD8CB821D9E1F40A" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Silpha</taxonomicName>
/
<taxonomicName id="2B166B5259908BEA2D21941722C01F0C" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ Fabricius, 1775 / Jan
<normalizedToken id="FEE395E289BAF22CDBC10DE069BDB8EA" originalValue="Růžička">Ruzicka</normalizedToken>
des. 2012 [p, red label] //
<taxonomicName id="4EAED6CFA5E5D15FF76E768D243C97FA" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="42318E7E7083D5845433BF2D55908E81" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ (Fabricius, 1775) / Jan
<normalizedToken id="BDA2828C8C1532D2401462013D55D9E3" originalValue="Růžička">Ruzicka</normalizedToken>
det. 20 [p] 12 [hw]&quot;.
</paragraph>
<paragraph id="BB21ADE0484D99584774C771451B708B" pageId="7" pageNumber="86">
Lectotype female of
<taxonomicName id="803ECD494822E350B3C5B0E087FDD0AE" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Silpha gibba</taxonomicName>
(here designated) (MNHN, coll. generale), labelled: &quot;Type / de
<normalizedToken id="22235846C4859E3C7B59CB2482A7A8C1" originalValue="Brullé">Brulle</normalizedToken>
[hw] // TYPE [p, red label] // MUSEUM PARIS /
<normalizedToken id="46274F55DB2BF3D67825C7F291CD4123" originalValue="MORÉE">MOREE</normalizedToken>
/
<normalizedToken id="0B1AD504861216CFE7661DA740C38275" originalValue="BRULLÉ">BRULLE</normalizedToken>
4187-33 [p] // Lectotype /
<taxonomicName id="FE9BB859747E05780518447AB9DD1A5C" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Silpha gibba</taxonomicName>
/
<normalizedToken id="C555F7DB0C1234358BAB663EF744D5E7" originalValue="Brullé">Brulle</normalizedToken>
, 1832 / J.
<normalizedToken id="70E4555F9A2247A58BFFDB7B07819820" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="90DCFEC021A324832B702F6D6E4448DA" originalValue="Růžička">Ruzicka</normalizedToken>
/ des. 2014 [p, red label] //
<taxonomicName id="52E9E6EBF69A4E87A518B6AA5B9DA1B1" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="1D6EF1CB9A37498F3973E96382997FA5" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
(Fabricius, 1775) / J.
<normalizedToken id="CC64346B9B996F39C0D8C23D22871915" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="1E6F555DD5E1B39763CE325AB97CA4D7" originalValue="Růžička">Ruzicka</normalizedToken>
det. 2014 [p]&quot;. Paralectotype: 1 male (MNHN, coll. generale), labelled: &quot;
<taxonomicName id="899EDB82C22707359B0C31D625E9E3D7" lsidName="gibba" pageId="7" pageNumber="86" rank="species" species="gibba">gibba</taxonomicName>
Br. [hw] // TYPE [p, red label] // MUSEUM PARIS /
<normalizedToken id="7C615E6DE987F7188B0C11078DA6203F" originalValue="MORÉE">MOREE</normalizedToken>
/
<normalizedToken id="0434DA322F90DA960C247F806AF54121" originalValue="BRULLÉ">BRULLE</normalizedToken>
4187-33 [p] // Paralectotype
<taxonomicName id="25A83C942C04AA7C961FCE40FE29DB95" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Silpha gibba</taxonomicName>
/
<normalizedToken id="6C5012A03040A959A3DBF79955B2255B" originalValue="Brullé">Brulle</normalizedToken>
, 1832 / J.
<normalizedToken id="7E4FDD33EEE8B34057D6A8DE7CC09695" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="156C68B29AB73E365EE957618A3C6604" originalValue="Růžička">Ruzicka</normalizedToken>
des. 2014 [p, red label] //
<taxonomicName id="C19012467E165B30AD5DD48179A2D73B" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="E333CD39844DCC369237FBE0583DDB9A" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
(Fabricius, 1775) / J.
<normalizedToken id="D7E1DF34DF8FD925EBA8527A9F2856D7" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="3FFDC0E94DC6A1070934EAE7D08CA929" originalValue="Růžička">Ruzicka</normalizedToken>
det. 2014 [p]&quot;.
</paragraph>
<paragraph id="42452D6E0E81F6FF6E2B18B393ACDEA2" pageId="7" pageNumber="86">
Lectotype female of
<taxonomicName id="9A4F44F6E4B0F161F87455DF9FE96C55" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. costulata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="costulata">Ablattaria gibba var. costulata</taxonomicName>
(here designated) (MNHN, coll. Pic), labelled &quot;TURQUIE / Constantinople [= Istanbul] [p] //
<taxonomicName id="64D7ABE851C2DA1E0B78DBE10A4966AF" lsidName="var. costatula" pageId="7" pageNumber="86" rank="variety" variety="costatula">var. costatula</taxonomicName>
[sic] / m. [hw,
<normalizedToken id="3820F51B10E73365B46411D28E52C6A4" originalValue="Portevin’s">Portevin's</normalizedToken>
handwriting] // TYPE [p, red label] //
<normalizedToken id="1FA99CA7FAA51358FE5F66D724DF46E1" originalValue="Muséum">Museum</normalizedToken>
Paris / Coll. M. Pic [p] // Lectotype /
<taxonomicName id="D492BFE19CAD1634B34BB84934EE64ED" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Ablattaria gibba</taxonomicName>
/
<taxonomicName id="AE97CBA99CEDB8C60049E8E16B7A7068" lsidName="var. costulata" pageId="7" pageNumber="86" rank="variety" variety="costulata">var. costulata</taxonomicName>
/ Portevin, 1926 / J.
<normalizedToken id="5BBAF687EF1DCC6780DF7E65044D8972" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="E2B9AF1717FD146E8843422704B1A8BB" originalValue="Růžička">Ruzicka</normalizedToken>
/ des. 2014 [p, red label] //
<taxonomicName id="7D78A7B195D9E0AFE40A9F53CFCB628C" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="227BCFF150B7C843687766F7B731CC1B" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
(Fabricius, 1775) / J.
<normalizedToken id="DA3D7FB4807ED0E524B14917AAB05D06" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="11BF643068CC40960C7DDC37643D6017" originalValue="Růžička">Ruzicka</normalizedToken>
/ det. 2014 [p]&quot;.
</paragraph>
<paragraph id="654B4EFE69A2E999230626424049D532" pageId="7" pageNumber="86">
Lectotype male of
<taxonomicName id="4158A779F5AE27884C083CA0831AB100" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. distinguenda" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="distinguenda">Ablattaria gibba var. distinguenda</taxonomicName>
(here designated) (MNHN, coll. Pic), pinned, labelled: &quot;
<taxonomicName id="6188F654903AEAAC43C1F076D5E002F1" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
/ var. [hw] //
<taxonomicName id="23BBBD02FD3EEBE01F6A30C93D2D75C9" lsidName="var. distinguenda" pageId="7" pageNumber="86" rank="variety" variety="distinguenda">var. distinguenda</taxonomicName>
/ m. [hw,
<normalizedToken id="A36E9129EA544BDF0879AC1475C6C990" originalValue="Portevin’s">Portevin's</normalizedToken>
handwriting] // TYPE [p, red modern label] //
<normalizedToken id="14E0CD07C695218A4C8C76E806CC2947" originalValue="Muséum">Museum</normalizedToken>
Paris / Coll. M. Pic [p] // Lectotype /
<taxonomicName id="F652E969DA21FCB4DBAE4F64F3C70624" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Ablattaria gibba</taxonomicName>
/
<taxonomicName id="26AA7ACD255A7C6522E868639D5CACD3" lsidName="var. distinguenda" pageId="7" pageNumber="86" rank="variety" variety="distinguenda">var. distinguenda</taxonomicName>
/ Portevin, 1926 / J.
<normalizedToken id="1F4FFAF3512C9A299D18162716A66B22" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="235D7C115A9AD1FAF9A302C90E38C3A6" originalValue="Růžička">Ruzicka</normalizedToken>
/ des. 2014 [p, red label] //
<taxonomicName id="895100A60CA692D9513D948DE3FC17FC" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="FD36F8BF97B158F6E4F67181C0894DD1" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
(Fabricius, 1775) / J.
<normalizedToken id="0211053BB6C6807C7A745A1A959AD62A" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="4926EF14BC0346BA78A9B11BD0833111" originalValue="Růžička">Ruzicka</normalizedToken>
det. 2014 [p]&quot;.
</paragraph>
<paragraph id="E343F00A26A291777AC8AD1AD15CF9AF" pageId="7" pageNumber="86">
Lectotype male of
<taxonomicName id="E648D3B3BCEC8035BD9BCF751F3A977C" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. punctata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="punctata">Ablattaria gibba var. punctata</taxonomicName>
(here designated) (MNHN, coll. Marmottan) (Fig. 26), pinned, labelled: &quot;Turquie [hw] // TYPE [p, red modern label] //
<taxonomicName id="C0334EF662A02B354B6CFF2D11D6A74D" lsidName="var. punctata" pageId="7" pageNumber="86" rank="variety" variety="punctata">var. punctata</taxonomicName>
/ m. [hw, probably
<normalizedToken id="86E9DC83E7CA5C12656A613929F5DE54" originalValue="Portevin’s">Portevin's</normalizedToken>
handwriting] //
<normalizedToken id="595FEFEA2808F7A93D522693F66F277A" originalValue="Muséum">Museum</normalizedToken>
Paris / 1914 / Coll. H. Marmottan [p, modern label] // Lectotype /
<taxonomicName id="1C3A0DF08E7BE4BACD807B721A45A2FA" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="gibba">Ablattaria gibba</taxonomicName>
/
<taxonomicName id="A7DBC40C1FD0D965300DF1109B02B08B" lsidName="var. punctata" pageId="7" pageNumber="86" rank="variety" variety="punctata">var. punctata</taxonomicName>
/ Portevin, 1926 / J.
<normalizedToken id="07B5DA441FB36B890604F45CE3B9F0AD" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="13DD90AFDA7458CB77B8E48F36538186" originalValue="Růžička">Ruzicka</normalizedToken>
/ des. 2014 [p, red label] //
<taxonomicName id="94D3CBE050C28C5249A9BF98BB18BADD" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="73CF350304466C63362F7F37A27AA9AD" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
(Fabricius, 1775) / J.
<normalizedToken id="BE61D7B74632EC654570818261A2865F" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="2453EE061DBD9217CAA4A7E283F0E52D" originalValue="Růžička">Ruzicka</normalizedToken>
/ det. 2014 [p]&quot;. Paralectotype: 1 male, labelled: &quot;MUSEUM PARIS [p] / Turquie / Jejeune 1881 [hw] // TYPE [p, red label] // Paralectotype /
<taxonomicName id="BC92CEB22ADA5BDF8AE107C0799315DE" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. punctata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="variety" species="gibba" variety="punctata">Ablattaria gibba var. punctata</taxonomicName>
/ Portevin, 1926 / J.
<normalizedToken id="49CC52E2CEEDAEE4292E30484FB2B2B6" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="A6002C0C21AA2943A59B8DAFFFF6B2B9" originalValue="Růžička">Ruzicka</normalizedToken>
des. 2014 [p, red label] //
<taxonomicName id="245B9EC9102684B12CA8298E8B0BB44B" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
/
<taxonomicName id="CFC598F593544627EA389000602E4C92" lsidName="laevigata" pageId="7" pageNumber="86" rank="species" species="laevigata">laevigata</taxonomicName>
(Fabricius, 1775) / J.
<normalizedToken id="C4042BD9E9069C61A017235061A8226C" originalValue="Qubaiová">Qubaiova</normalizedToken>
&amp; J.
<normalizedToken id="662F6293E82574A4F72665DAC7BD7D70" originalValue="Růžička">Ruzicka</normalizedToken>
det. 2014 [p]&quot;.
</paragraph>
<caption id="DE6F898D4CE99D5A41CB3674416585E6" pageId="7" pageNumber="86">
<paragraph id="13088BDEF5DCA76F87152D8D4DA81E87" pageId="7" pageNumber="86">
Figures 28-31. Lectotype and paralectotype of
<taxonomicName id="E03E8B26BD90F8292C74F997FB839476" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
Fabricius: 28, 29 lectotype (female) 30, 31 paralectotype (female,
<taxonomicName id="125AE08A730A82096213FAEB445B65DF" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha tyrolensis" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="tyrolensis">Silpha tyrolensis</taxonomicName>
; see text) 28, 30 dorsal view 29, 31 lateral view. (Photo K.P. Puliafico).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="646E2CF64BEFC06A72DAEDAADE9F46B2" pageId="8" pageNumber="87" type="additional material examined">
<paragraph id="4280FC6E71B8EF2E01BDB68A394FF4E2" pageId="8" pageNumber="87">
<pageBreakToken id="4C5FAB0D926ACCCB54F477F1B087A3AD" pageId="8" pageNumber="87" start="start">Additional</pageBreakToken>
material examined.
</paragraph>
<paragraph id="EC39347E74C84160187938BD332D0511" pageId="8" pageNumber="87">2206 specimens, see Appendix 1.</paragraph>
</subSubSection>
<subSubSection id="9DEA731B8D7DE33C3045C68D16D6EB45" pageId="8" pageNumber="87" type="diagnostic description">
<paragraph id="05301CDF2603E00A263FF1F71806DBC5" pageId="8" pageNumber="87">Diagnostic description.</paragraph>
<paragraph id="30901E2D477FCD1327598F1828E0D807" pageId="8" pageNumber="87">
Total body length 9-19 mm, body matt. Pronotum semi-elliptical, with evenly distributed, homogenous, distinct punctures on whole dorsal surface (Fig. 15). Elytra regularly rounded (more in larger specimens - compare Figs 9-11); with densely and evenly distributed medium-sized, distinct punctures of similar size (Fig. 18), rarely with two very fine, longitudinal lines that are occasionally more visible (Fig. 21). Rarely, individual larger punctures are intermixed, but never in such regular pattern as in
<taxonomicName id="9DE96F624E894B94C4E2F77A934EE8B6" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria arenaria" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="arenaria">Ablattaria arenaria</taxonomicName>
and
<taxonomicName id="A73DF2D0556170C6DC0FE577D5DF2095" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria cribrata" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="cribrata">Ablattaria cribrata</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C16785CE413AD13895200F023CD99DAF" lastPageId="10" lastPageNumber="89" pageId="8" pageNumber="87" type="remarks">
<paragraph id="6F9CAE3BC53ECF19443CB593A932DB3F" pageId="8" pageNumber="87">Remarks.</paragraph>
<paragraph id="C963C63C5AE9B89EB8F307332486C6E2" pageId="8" pageNumber="87">
Both
<bibRefCitation id="B9B0BBA2B9513190B7D6712AC474F0C6" author="Fabricius, JC" journalOrPublisher="Libraria Kortii, Flensburgi et Lipsiae" pageId="20" pageNumber="99" title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus" year="1775">Fabricius (1775)</bibRefCitation>
and
<bibRefCitation id="3272EFC3897551516A37EE1045D71A99" author="Sulzer, JH" journalOrPublisher="H. Steiner und Comp., Winterthur" pageId="23" pageNumber="102" title="Abgekuerzte Geschichte der Insecten nach dem Linaeischen System" year="1776">Sulzer (1776)</bibRefCitation>
refer in their descriptions of
<taxonomicName id="6DED4E2AC682FE0EB7B550538EEAF925" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
and
<taxonomicName id="379137B8046ED329BC152E4C48D601A4" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha polita" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="polita">Silpha polita</taxonomicName>
to
<bibRefCitation id="9DD9B8A77425087AAFA1D34F9165FE25" author="Geoffroy, E-L" journalOrPublisher="Durand, Paris" pageId="21" pageNumber="100" title="Histoire abregee des Insectes qui se trouvent aux environs de Paris: Dans laquelle ces Animaux sont ranges suivant un ordre methodique" year="1762">Geoffroy (1762</bibRefCitation>
: 122, species #8). However, the book of Geoffroy is not consistently binominal and Opinion 1754 (
<bibRefCitation id="1E10054F8EDA60DDFCC4EF19BCB94FFE" author="Horion, A" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="21" pageNumber="100" pagination="58 - 70" title="Opinion 1754. Histoire abregee des insectes qui se trouvent aux environs de Paris (Geoffroy, 1762): some generic names conserved (Crustacea, Insecta)." volume="51" year="1994">ICZN 1994</bibRefCitation>
) placed it on the Official List of Works in Zoological Nomenclature with only some generic names available. Accordingly, the author of
<taxonomicName id="6DF9306B09907B6D48CD2CC0A53A4EA7" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
is Fabricius and the author of
<taxonomicName id="77B3BD94D218A745C26E592B1732C7FB" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha polita" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="polita">Silpha polita</taxonomicName>
is Sulzer.
</paragraph>
<paragraph id="3E63EE978237CB8DC1CA15C0BEEBD0AC" pageId="9" pageNumber="88">
<pageBreakToken id="83F52E3F01731537189971C7F209B853" pageId="9" pageNumber="88" start="start">In</pageBreakToken>
the syntype series of
<taxonomicName id="ED61FC754E03C480832EBD1030955DB9" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha laevigata" order="Coleoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="laevigata">Silpha laevigata</taxonomicName>
from ZMUC and BMNH, consistent with current understanding of
<taxonomicName id="D94878EED7613AC7116F2E0470570364" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
, we also found intermixed a single specimen of
<taxonomicName id="ADE29044B204D6430A9FB99D3DB2A1B9" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha tyrolensis" order="Coleoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="tyrolensis">Silpha tyrolensis</taxonomicName>
Laicharting, 1781 (in ZMUC, &quot;Kiel collection&quot;; see above for details). This syntype specimen is here considered a paralectotype. We have designated a female from ZMUC, the &quot;Copenhagen collection&quot;, as the lectotype to fix this name as currently used.
</paragraph>
<paragraph id="1C7B840DDB9FDAA72DD2FC0ACD0E35AC" pageId="10" pageNumber="89">
<taxonomicName id="C388EB518298F271359F8CB3FDF2FBC4" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">
<pageBreakToken id="DE182F4F8BB41B3D6B843A8D4E4F7A24" pageId="10" pageNumber="89" start="start">Ablattaria</pageBreakToken>
laevigata
</taxonomicName>
is a widely distributed species with regional variation in size and shape between populations (see Geometric morphometrics section below), and also with some variability in punctation of elytra, sometimes with intermixed larger punctures or an impunctate pair of longitudinal lines present on elytra.
</paragraph>
<paragraph id="7D60361443BF2EEEFF674419729C5BFE" pageId="10" pageNumber="89">
There are no distinct differences in the description of
<taxonomicName id="6AD6FC373B045CC4CF0A6CED79321B7F" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha polita" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="polita">Silpha polita</taxonomicName>
to separate it from
<taxonomicName id="85DDD288B4D40746BB1622290A02C581" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
, and we believe that this taxon is correctly considered as a junior subjective synonym of
<taxonomicName id="44ED7DBD48FF96DE00489F5025965ABB" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
by
<bibRefCitation id="7A7349DC75E57A7994DBD79CDBAFE468" author="Reitter, E" journalOrPublisher="Verhandlungen des Naturforschenden Vereines in Bruenn" pageId="22" pageNumber="101" pagination="3 - 122" title="Bestimmungs-Tabellen der europaeischen Coleopteren. XII. Necrophaga (Platypsyllidae, Leptinidae, Silphidae, Anisotomidae und Clambidae)." volume="23" year="1884">Reitter (1884)</bibRefCitation>
and
<bibRefCitation id="03FFBA180B8E8C56F5857795762A2FAB" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller (1979)</bibRefCitation>
. In our opinion, the variation in body size, proportions and surface sculpturation which led to the description of
<taxonomicName id="565854723DF95F54DAAFD130CCE8607B" class="Insecta" family="Leiodidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha gibba" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="gibba">Silpha gibba</taxonomicName>
and several varieties of
<bibRefCitation id="8227AECB1D0485D4D7ABA310EDC149C4" author="Ganglbauer, L" journalOrPublisher="C. Gerold's Sohn, Wien" pageId="20" pageNumber="99" title="Die Kaefer von Mitteleuropa. Die Kaefer der oesterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzoesischen und italienischen Alpengebietes. Dritter Band, Familienreihe Staphylinoidea, II. Theil. Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeriidae, Trichopterygidae, Hydroscaphidae, Scaphidiidae, Histeridae. Familienreihe Clavicornia. Sphaeritidae, Ostomidae, Byturidae, Nitidulidae, Cucuidae, Erotylidae, Phalacridae, Thorictidae, Lathridiidae, Mycetophagidae, Colydidae, Endomychidae, Coccinellidae" year="1899">Ganglbauer (1899)</bibRefCitation>
and
<bibRefCitation id="E8C15B9DB647A9836EA2A621C75F1D93" author="Portevin, G" journalOrPublisher="Lechevalier, Paris" pageId="22" pageNumber="101" title="Les Grands Necrophages du Globe. Silphini - Necrodini - Necrophorini." volumeTitle="Encyclopedie Entomologique" year="1926">Portevin (1926)</bibRefCitation>
fall within the infrasubspecific variation of
<taxonomicName id="5221C74102BCC71A2EF21C03A9E076C9" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
. We agree with
<bibRefCitation id="D125EE66B97C02FE9710F02E6D0629B6" author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde (Serie A (Biologie ))" pageId="23" pageNumber="102" pagination="1 - 8" title="Revision der Gattung Ablattaria Reitter 1884 (Coleoptera: Silphidae)." volume="321" year="1979">Schawaller (1979)</bibRefCitation>
, who considered
<taxonomicName id="086B324B16A7498A21CAA7191719FABC" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. costulata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="variety" species="gibba" variety="costulata">Ablattaria gibba var. costulata</taxonomicName>
,
<taxonomicName id="1232E306199789CBC7E3B356174A7B19" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. distinguenda" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="variety" species="gibba" variety="distinguenda">Ablattaria gibba var. distinguenda</taxonomicName>
and
<taxonomicName id="339E649517AA17B97CD0BA86BAA10C51" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria gibba var. punctata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="variety" species="gibba" variety="punctata">Ablattaria gibba var. punctata</taxonomicName>
as junior subjective synonyms of
<taxonomicName id="12A25FB2D1EDE8CE9189700A477CF5A1" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
. Further, we consider
<taxonomicName id="F7047E8C4636A3A84E11BBF2DA7F1E75" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata var. meridionalis" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="variety" species="laevigata" variety="meridionalis">Ablattaria laevigata var. meridionalis</taxonomicName>
of
<bibRefCitation id="65ED699F7A468DDCEDB24E172F9FA4FB" author="Ganglbauer, L" journalOrPublisher="C. Gerold's Sohn, Wien" pageId="20" pageNumber="99" title="Die Kaefer von Mitteleuropa. Die Kaefer der oesterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzoesischen und italienischen Alpengebietes. Dritter Band, Familienreihe Staphylinoidea, II. Theil. Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeriidae, Trichopterygidae, Hydroscaphidae, Scaphidiidae, Histeridae. Familienreihe Clavicornia. Sphaeritidae, Ostomidae, Byturidae, Nitidulidae, Cucuidae, Erotylidae, Phalacridae, Thorictidae, Lathridiidae, Mycetophagidae, Colydidae, Endomychidae, Coccinellidae" year="1899">Ganglbauer (1899)</bibRefCitation>
as a junior subjective synonym of
<taxonomicName id="ADE428FF5B52AE5201B28BFF9590CAFB" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9154251718117DC4A5FEF32A43C0D92D" pageId="10" pageNumber="89" type="biology">
<paragraph id="E26ED9218725F2EECD01DE3C22375AAA" pageId="10" pageNumber="89">Biology.</paragraph>
<paragraph id="A6112C75B96ABD20A12106DBB4D152EF" pageId="10" pageNumber="89">
Seasonal activity of adults with a peak in
<normalizedToken id="23CAAAC45B854E345D81DDBE1C87B6C6" originalValue="April–June">April-June</normalizedToken>
(Table 1).
</paragraph>
</subSubSection>
<subSubSection id="9BABB8B5E0695862735A3D3181EB31AA" pageId="10" pageNumber="89" type="distribution">
<paragraph id="CCBDC0C07BD4145572C6667AA08EABEA" pageId="10" pageNumber="89">Distribution.</paragraph>
<paragraph id="7685ED240BE734E8DE0BE8B33431F6B5" pageId="10" pageNumber="89">Most of Europe; from the west (Spain to United Kingdom), through all of central and southern Europe, reaching to the east and north of Turkey; Ukraine, southern Russia, Georgia to Armenia (Fig. 32).</paragraph>
<caption id="0062466267C9DA13A3F8BAB496E51739" pageId="10" pageNumber="89">
<paragraph id="93F38A099CD09BB1A855D58AB704D585" pageId="10" pageNumber="89">
Figures 32-33. 32 Distribution of
<taxonomicName id="2BDA379F7BCA066821EF04316D491EE0" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="genus">Ablattaria</taxonomicName>
spp. in Western Palaearctic region 33 Delimitation and distribution of samples of
<taxonomicName id="E81B14EDC5498EB8BB9B7D27CAE2C500" class="Insecta" family="Staphylinidae" genus="Ablattaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ablattaria laevigata" order="Coleoptera" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="laevigata">Ablattaria laevigata</taxonomicName>
used in geometric morphometrics.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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